Provanna

Provanna; Temporal range: Cretaceous-Recent PreꞒ Ꞓ O S D C P T J K Pg N
	Provanna hirokoae
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Superfamily:	Abyssochrysoidea
Family:	Provannidae
Genus:	Provanna; Dall, 1918
Type species
	Trichotropis (Provanna) lomana Dall, 1918;
Synonyms
	Trichotropis (Provanna) Dall, 1918 (original rank)

Provanna is a genus of sea snails, marine gastropod mollusks in the family Provannidae.^[1]

The species was named after Dall's first wife.

This is a very old genus with species already existing in Cretaceous seep deposits off Japan (93 - 100 million years ago). The Caribbean and the Pacific fauna were isolated about 3 million years ago by the closure of the Central American Isthmus.

Description

The apical whorls of the protoconch are lacking through decollation. The opening is then sealed with a calcareous plug. The sculpture of the ovate fusiform shell shows scattered sigmoidal axial ribs that are crossed by spiral cords. The whorls are broad.

Habitat

They are common in hydrothermal vents, hydrocarbon cold seeps and (more rarely) on deep-water food-falls, depending on the species. Most species feed on filamentous bacteria, while others feed on detritus.

Species

Species within the genus Provanna include:

Provanna abyssalis Okutani & Fujikura, 2002^[2]
Provanna admetoides Warén & Ponder, 1991^[3]
† Provanna alexi Amano & Little, 2014
† Provanna antiqua Squires, 1995
† Provanna azurini Kiel, Aguilar & Kase, 2020
Provanna beebei Linse, Nye, Copley & C. Chen, 2019
Provanna buccinoides Warén & Bouchet, 1993^[4]
Provanna chevalieri Warén & Bouchet, 2009^[5]
Provanna cingulata C. Chen, Watanabe & Ohara, 2018
Provanna clathrata Sasaki, Ogura, Watanabe & Fujikura, 2016
Provanna cooki Linse, Nye, Copley & C. Chen, 2019
Provanna fenestrata C. Chen, Watanabe & Sasaki, 2019
† Provanna fortis Hybertsen & Kiel, 2018
Provanna glabra Okutani, Tsuchida & Fujikura, 1992^[6]
Provanna goniata Warén & Bouchet, 1986^[7]
† Provanna hirokoae Amano & Little, 2014
Provanna ios Warén & Bouchet, 1986^[8]
Provanna kuroshimensis Sasaki, Ogura, Watanabe & Fujikura, 2016
Provanna laevis Warén & Ponder, 1991^[9]
Provanna lomana (Dall, 1918)^[10]
Provanna lucida Sasaki, Ogura, Watanabe & Fujikura, 2016
Provanna macleani Warén & Bouchet, 1989^[11]
† Provanna marshalli Saether, Little & K. A. Campbell, 2010
Provanna muricata Warén & Bouchet, 1986^[12]
† Provanna nakagawaensis Kaim, Jenkins & Hikida, 2009
Provanna nassariaeformis Okutani, 1990^[13]
Provanna pacifica (Dall, 1908)^[14]
† Provanna pelada Kiel, Hybertsen, Hyžný & Klompmaker, 2019
Provanna reticulata Warén & Bouchet, 2009^[15]
Provanna sculpta Warén & Ponder, 1991^[16]
Provanna segonzaci Warén & Ponder, 1991^[17]
Provanna shinkaiae Okutani & Fujikura, 2002^[18]
Provanna stephanos C. Chen, Watanabe & Sasaki, 2019
Provanna subglabra Sasaki, Ogura, Watanabe & Fujikura, 2016
† Provanna tappuensis Kaim, Jenkins & Warén, 2008
† Provanna urahoroensis Amano & Jenkins, 2013
Provanna variabilis Warén & Bouchet, 1986^[19]

Several fossil species have been described, predominantly identified from hydrocarbon seep deposits (Saether et al., 2010), including:

Provanna antiqua Squires & Goedert, 1995 ^[20]
Provanna marshalli Saether, Little & Campbell, 2010
Provanna nakagawaensis Kaim, Jenkins & Hikida, 2009
Provanna tappuensis Kaim, Jenkins & Warén, 2008

There also are several fossil species known, but not formally described and named, from Central America, Japan, and possibly the Philippines (see Saether et al., 2010).

Species brought into synonymy

Provanna marianensis Okutani & Fujikura, 1990: synonym of Desbruyeresia marianensis (Okutani & Fujikura, 1990)

References

^ ^a ^b Provanna Dall, 1918. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna abyssalis Okutani & Fujikura, 2002. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna admetoides Warén & Ponder, 1991. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna buccinoides Warén & Bouchet, 1993. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna chevalieri Warén & Bouchet, 2009. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna glabra Okutani, Tsuchida & Fujikura, 1992. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna goniata Warén & Bouchet, 1986. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna ios Warén & Bouchet, 1986. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna laevis Warén & Ponder, 1991. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna lomana (Dall, 1918). Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna macleani Warén & Bouchet, 1989. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna muricata Warén & Bouchet, 1986. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna nassariaeformis Okutani, 1990. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna pacifica (Dall, 1908). Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna reticulata Warén & Bouchet, 2009. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna sculpta Warén & Ponder, 1991. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna segonzaci Warén & Ponder, 1991. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna shinkaiae Okutani & Fujikura, 2002. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Provanna variabilis Warén & Bouchet, 1986. Retrieved through: World Register of Marine Species on 4 May 2010.
^ Steffen Kiel, New Records and Species of Molluscs from Tertiary Cold-Seep Carbonates in Washington State, USA; Journal of Paleontology, Vol. 80, No. 1 (Jan., 2006), pp. 121-137

Healy, J. M. 1990. Taxonomic affinities of the deep-sea genus Provanna: new evidence from sperm structure; J. Molluscan Stud. 56: 119-122
Warén, A. & Bouchet, P. (1993). New records, species, genera, and a new family of gastropods from hydrothermal vents and hydrocarbon seeps. Zoologica Scripta. 22: 1-90.
Spencer, H.G., Marshall, B.A. & Willan, R.C. 2009 Recent Mollusca. pp 196–219 in: Gordon, D.P. (Ed.), The New Zealand inventory of biodiversity. 1. Kingdom Animalia: Radiata, Lophotrochozoa, Deuterostomia. Canterbury University Press, Christchurch
Saether, K.P., Little, C.T.S. & Campbell, K.A. (2010) A new fossil provannid gastropod from Miocene hydrocarbon seep deposits, East Coast Basin, North Island, New Zealand. Acta Palaeontologica Polonica 55, 507–517.

External links

[WoRMS-1] Provanna Dall, 1918. Retrieved through: World Register of Marine Species on 4 May 2010.

[2] Provanna abyssalis Okutani & Fujikura, 2002. Retrieved through: World Register of Marine Species on 4 May 2010.

[3] Provanna admetoides Warén & Ponder, 1991. Retrieved through: World Register of Marine Species on 4 May 2010.

[4] Provanna buccinoides Warén & Bouchet, 1993. Retrieved through: World Register of Marine Species on 4 May 2010.

[5] Provanna chevalieri Warén & Bouchet, 2009. Retrieved through: World Register of Marine Species on 4 May 2010.

[6] Provanna glabra Okutani, Tsuchida & Fujikura, 1992. Retrieved through: World Register of Marine Species on 4 May 2010.

[7] Provanna goniata Warén & Bouchet, 1986. Retrieved through: World Register of Marine Species on 4 May 2010.

[8] Provanna ios Warén & Bouchet, 1986. Retrieved through: World Register of Marine Species on 4 May 2010.

[9] Provanna laevis Warén & Ponder, 1991. Retrieved through: World Register of Marine Species on 4 May 2010.

[10] Provanna lomana (Dall, 1918). Retrieved through: World Register of Marine Species on 4 May 2010.

[11] Provanna macleani Warén & Bouchet, 1989. Retrieved through: World Register of Marine Species on 4 May 2010.

[12] Provanna muricata Warén & Bouchet, 1986. Retrieved through: World Register of Marine Species on 4 May 2010.

[13] Provanna nassariaeformis Okutani, 1990. Retrieved through: World Register of Marine Species on 4 May 2010.

[14] Provanna pacifica (Dall, 1908). Retrieved through: World Register of Marine Species on 4 May 2010.

[15] Provanna reticulata Warén & Bouchet, 2009. Retrieved through: World Register of Marine Species on 4 May 2010.

[16] Provanna sculpta Warén & Ponder, 1991. Retrieved through: World Register of Marine Species on 4 May 2010.

[17] Provanna segonzaci Warén & Ponder, 1991. Retrieved through: World Register of Marine Species on 4 May 2010.

[18] Provanna shinkaiae Okutani & Fujikura, 2002. Retrieved through: World Register of Marine Species on 4 May 2010.

[19] Provanna variabilis Warén & Bouchet, 1986. Retrieved through: World Register of Marine Species on 4 May 2010.

[20] Steffen Kiel, New Records and Species of Molluscs from Tertiary Cold-Seep Carbonates in Washington State, USA; Journal of Paleontology, Vol. 80, No. 1 (Jan., 2006), pp. 121-137

[1]

[2]

[3]

[4]

[5]

[6]

[7]

[8]

[9]

[10]

[11]

[12]

[13]

[14]

[15]

[16]

[17]

[18]

[19]

[20]