The wingspan is 5.4–6.6 millimetres (0.21–0.26 in). There are two generations per year with late instar larvae being encountered throughout the year.
The larvae feed on Prunus ilicifolia. They mine the leaves of their host plant. The mine is located on the upper-surface and is linear or serpentine and convoluted terminally and in this region frequently crossing itself or forming a blotch. Later it gradually increasing in width throughout its length. The frass is deposited centrally as a continuous line.
- "Stigmella braunella (Jones, 1933)". Moth Photographers Group. Retrieved November 20, 2013.
- Annette F. Braun (1914). Notes on the North American species of Nepticula with descriptions of new species (Lepidoptera). 46. Canadian Entomologist. pp. 17–27.
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