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Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
(unranked): clade Heterobranchia
informal group Lower Heterobranchia
Superfamily: Mathildoidea
Family: Tofanellidae
Bandel, 1995[1]
  • Graphididae J.C. N. Barros, Mello, F. N. Barros, Lima, Santos, Cabral & Padovan, 2003
  • Tofanellinae Bandel, 1995
  • Usedomellinae Gründel, 1998

The Tofanellidae is an taxonomic family of sea snails, marine gastropod mollusks in the informal group Lower Heterobranchia.[2]


The family Tofanellidae used to consist of the two following subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

  • Tofanellinae Bandel, 1995
  • Usedomellinae Gründel, 1998[3]


Genera within the family Tofanellidae include:

Genera brought into synonymy
  • Cioniscus Jeffreys, 1869: synonym of Graphis Jeffreys, 1867
  • Dissopalia Iredale, 1936: synonym of Bouryiscala Cossmann, 1902
  • Rotfanella Gründel, 1998: synonym of Graphis Jeffreys, 1867

Bandel (2005)[4] recognized the genus Graphis Jeffreys, 1867 as a living fossil within the subfamily Usedomellinae.[4]


  1. ^ Bandel K. (1995). "The Mathildoidea from the Late St. Cassian Formation". Scripta Geologica 111: 1-83, 19 Plates. Leiden. pages 21, 39.
  2. ^ a b c Bouchet P.; Rocroi J.-P.; Frýda J.; Hausdorf B.; Ponder W.; Valdés Á. & Warén A. (2005). "Classification and nomenclator of gastropod families". Malacologia: International Journal of Malacology. Hackenheim, Germany: ConchBooks. 47 (1-2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997. 
  3. ^ Gründel J. (1998). "Heterostropha (Gastropoda) aus dem Dogger Norddeutschlands und Nordpolens. II. Weitere Allogastropoda". Freiberger Forschungshefte ser. C., 474(6): 1-37, plates 1-7. abstract.
  4. ^ a b c d e Bandel K. (2005). "Living fossils among tiny Allogastropoda with high and slender shell from the reef environment of the Gulf of Aqaba with remarks on fossil and recent relatives". Mitt. Geol.-Paläont. Inst. Universität Hamburg 89: 1-24. PDF