Sebecidae

Sebecids; Temporal range: Late Cretaceous - Middle Miocene, ; 67.6–11.8 Ma PreꞒ Ꞓ O S D C P T J K Pg N ;
	Skull of Sebecus icaeorhinus
	Skeleton of Ogresuchus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	Archosauria
Clade:	Pseudosuchia
Clade:	Crocodylomorpha
Clade:	Crocodyliformes
Clade:	†Notosuchia
Clade:	†Sebecosuchia
Clade:	†Sebecia
Family:	†Sebecidae; Simpson, 1937
Subgroups
	†Ayllusuchus; †Barinasuchus; †Bretesuchus; †Dentaneosuchus; †Eremosuchus?; †Ilchunaia?; †Langstonia; †Lorosuchus; †Ogresuchus; †Sahitisuchus; †Sebecus; †Zulmasuchus;
Synonyms
	Bretesuchidae Gasparini, Fernandez & Powell, 1993;

Sebecidae is an extinct family of prehistoric terrestrial sebecosuchian crocodylomorphs, known from the Late Cretaceous and Cenozoic of Europe and South America. They were the latest surviving group of non-crocodilian crocodylomorphs.

The oldest known member of the group is Ogresuchus furatus known from the Upper Cretaceous (Maastrichtian) Tremp Formation (Spain).^[2] Other records of the group are known from the Eocene of Europe.^[3] Sebecids were diverse, abundant and broadly distributed in South America (mostly in Argentina, Brazil and Bolivia) during the Cenozoic, from the Paleocene until the Middle Miocene;^[4] although it has been suggested that at least some forms could have survived until the Miocene-Pliocene boundary in Brazil.^[5]

This group included many medium- and large-sized genera, from Sebecus to the giant 6-metre-long (20 ft) Barinasuchus from the Miocene.^[6]

Phylogeny

Juan Leardi and colleagues in 2024 defined Sebecidae in PhyloCode as "the least inclusive clade containing Sebecus icaeorhinus, Bretesuchus bonapartei, Barinasuchus arveloi, and Sahitisuchus fluminensis".^[7] The following cladogram simplified after Diego Pol and Jaime E. Powell (2011).^[4]

	Pabwehshi

	Stratiotosuchus

	Baurusuchus pachecoi

	Baurusuchus salgadoensis

Sebecidae

	Ayllusuchus

	Bretesuchus

	Lumbrera form

	Langstonia

	Sebecus

	Zulmasuchus

References

^ Martin, J. E.; Pochat-Cottilloux, Y.; Laurent, Y.; Perrier, V.; Robert, E.; Antoine, P.-O. (2023). "Anatomy and phylogeny of an exceptionally large sebecid (Crocodylomorpha) from the middle Eocene of southern France". Journal of Vertebrate Paleontology. 42 (4): e2193828. doi:10.1080/02724634.2023.2193828. S2CID 258361595.
^ Sellés, A. G.; Blanco, A.; Vila, B.; Marmi, J.; López-Soriano, F. J.; Llácer, S.; Frigola, J.; Canals, M.; Galobart, À. (2020). "A small Cretaceous crocodyliform in a dinosaur nesting ground and the origin of sebecids". Scientific Reports. 10 (1): Article number 15293. Bibcode:2020NatSR..1015293S. doi:10.1038/s41598-020-71975-y. PMC 7499430. PMID 32943663.
^ Martin, Jeremy E.; Pochat-Cottilloux, Yohan; Laurent, Yves; Perrier, Vincent; Robert, Emmanuel; Antoine, Pierre-Olivier (2022-10-28). "Anatomy and phylogeny of an exceptionally large sebecid (Crocodylomorpha) from the middle Eocene of southern France". Journal of Vertebrate Paleontology. 42 (4). Bibcode:2022JVPal..42E3828M. doi:10.1080/02724634.2023.2193828. ISSN 0272-4634.
^ ^a ^b Diego Pol and Jaime E. Powell (2011). "A new sebecid mesoeucrocodylian from the Rio Loro Formation (Palaeocene) of north-western Argentina". Zoological Journal of the Linnean Society. 163: S7–S36. doi:10.1111/j.1096-3642.2011.00714.x. hdl:11336/69518.
^ Liccardo, Antonio, and Luiz Carlos Weinschütz. "Registro inédito de fósseis de vertebrados na Bacia Sedimentar de Curitiba (PR) Archived 2015-06-02 at the Wayback Machine." Revista Brasileira de Geociências 40.3 (2010): 330-338.
^ Salias-Gismondi, R.; Antoine, P. O.; Baby, P.; Brusset, S.; Benammi, M.; Espurt, N.; de Franceschi, D.; Pujos, F.; et al. (2007). Middle Miocene Crocodiles From the Fitzcarrald Arch, Amazonian Peru (PDF). Instituto Geológical y Minero de España. p. 4. ISBN 978-84-7840-707-1. Archived from the original (PDF) on July 4, 2009. Retrieved May 12, 2010.
^ Leardi, J. M.; Pol, D.; Montefeltro, F.; Marinho, T. S.; Ruiz, J. V.; Bravo, G. G.; Pinheiro, A. E. P.; Godoy, P. L.; Nicholl, C. S. C.; Lecuona, A.; Larsson, H. C. E. (2024). "Phylogenetic nomenclature of Notosuchia (Crocodylomorpha; Crocodyliformes)". Bulletin of Phylogenetic Nomenclature. 1 (3): 44–82. doi:10.11646/bpn.1.3.2.

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[Martin2023-1] Martin, J. E.; Pochat-Cottilloux, Y.; Laurent, Y.; Perrier, V.; Robert, E.; Antoine, P.-O. (2023). "Anatomy and phylogeny of an exceptionally large sebecid (Crocodylomorpha) from the middle Eocene of southern France". Journal of Vertebrate Paleontology. 42 (4): e2193828. doi:10.1080/02724634.2023.2193828. S2CID 258361595.

[2] Sellés, A. G.; Blanco, A.; Vila, B.; Marmi, J.; López-Soriano, F. J.; Llácer, S.; Frigola, J.; Canals, M.; Galobart, À. (2020). "A small Cretaceous crocodyliform in a dinosaur nesting ground and the origin of sebecids". Scientific Reports. 10 (1): Article number 15293. Bibcode:2020NatSR..1015293S. doi:10.1038/s41598-020-71975-y. PMC 7499430. PMID 32943663.

[3] Martin, Jeremy E.; Pochat-Cottilloux, Yohan; Laurent, Yves; Perrier, Vincent; Robert, Emmanuel; Antoine, Pierre-Olivier (2022-10-28). "Anatomy and phylogeny of an exceptionally large sebecid (Crocodylomorpha) from the middle Eocene of southern France". Journal of Vertebrate Paleontology. 42 (4). Bibcode:2022JVPal..42E3828M. doi:10.1080/02724634.2023.2193828. ISSN 0272-4634.

[Lorosuchus-4] Diego Pol and Jaime E. Powell (2011). "A new sebecid mesoeucrocodylian from the Rio Loro Formation (Palaeocene) of north-western Argentina". Zoological Journal of the Linnean Society. 163: S7–S36. doi:10.1111/j.1096-3642.2011.00714.x. hdl:11336/69518.

[5] Liccardo, Antonio, and Luiz Carlos Weinschütz. "Registro inédito de fósseis de vertebrados na Bacia Sedimentar de Curitiba (PR) Archived 2015-06-02 at the Wayback Machine." Revista Brasileira de Geociências 40.3 (2010): 330-338.

[6] Salias-Gismondi, R.; Antoine, P. O.; Baby, P.; Brusset, S.; Benammi, M.; Espurt, N.; de Franceschi, D.; Pujos, F.; et al. (2007). Middle Miocene Crocodiles From the Fitzcarrald Arch, Amazonian Peru (PDF). Instituto Geológical y Minero de España. p. 4. ISBN 978-84-7840-707-1. Archived from the original (PDF) on July 4, 2009. Retrieved May 12, 2010.

[7] Leardi, J. M.; Pol, D.; Montefeltro, F.; Marinho, T. S.; Ruiz, J. V.; Bravo, G. G.; Pinheiro, A. E. P.; Godoy, P. L.; Nicholl, C. S. C.; Lecuona, A.; Larsson, H. C. E. (2024). "Phylogenetic nomenclature of Notosuchia (Crocodylomorpha; Crocodyliformes)". Bulletin of Phylogenetic Nomenclature. 1 (3): 44–82. doi:10.11646/bpn.1.3.2.

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