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{{short description|Clade comprising turtles, birds and crocodilians}}
{{short description|Clade comprising turtles, birds and crocodilians}}
{{For|an explanation of very similar terms|Archosauromorpha}}
{{Automatic taxobox
{{Automatic taxobox
| fossil_range = {{Fossil range|Lopingian|Recent|earliest=Guadalupian}} {{Small|Possible [[Guadalupian]] record}}
| fossil_range = {{Fossil range|Lopingian|Recent|earliest=Capitanian}} {{Small|Possible [[Capitanian]] record}}
| image = Proganochelys Quenstedti.jpg
| image = Proganochelys Quenstedti.jpg
| image_caption = ''[[Proganochelys quenstedti]]''
| image_caption = ''[[Proganochelys quenstedti]]''
| taxon = Archelosauria
| taxon = Archelosauria
| authority = Crawford ''et al.'', 2015<ref name="Crawford2015"/>
| authority = Crawford ''et al.'', 2015<ref name="Crawford2015"/>
| subdivision_ranks = Extant subclades
| subdivision_ranks = Subgroups
| subdivision = * [[Pantestudines]]
| subdivision =
* [[Pantestudines]]<br />{{Small|(Includes [[turtle]]s and their extinct relatives)}}
⚫
* [[Archosauromorpha]]
* [[Archosauromorpha]]<br />{{Small|(Includes [[archosaur]]s and their extinct relatives)}}
}}
}}


'''Archelosauria''' is a [[clade]] grouping [[turtle]]s and [[archosaur]]s (birds and crocodilians) and their fossil relatives.<ref name="Crawford2015"/> Multiple [[sequence alignment]]s of DNA and protein sequences and [[phylogenetic]] inferences showed that turtles are the closest living relatives to birds and crocodilians.<ref name="Shen2011"/><ref name="Tzika2011"/><ref name="Chiari2012"/> There are about 1000 [[ultra-conserved element]]s in their genome that are unique to turtles and archosaurs, but which are not found in [[lepidosaur]]s (tuatara, lizards and snakes).<ref name="Crawford2012"/> Other genome-wide analyses also support this grouping.<ref name="Wang2013"/><ref name="Field2014"/>
'''Archelosauria''' is a [[clade]] grouping [[turtle]]s and [[archosaur]]s (birds and crocodilians) and their fossil relatives, to the exclusion of [[lepidosaur]]s (the clade containing [[lizard]]s, [[snake]]s and the [[tuatara]]). The majority of [[phylogenetic]] analyses based on molecular data (e.g. [[DNA]] and [[protein]]s) have supported a sister-group relationship between turtles and archosaurs. On the other hand, Archelosauria has not been supported by most [[Morphology (biology)|morphological]] analyses, which have instead found turtles to either be descendants of [[parareptile]]s, early-diverging [[diapsid]]s outside of [[Sauria]], or close relatives of lepidosaurs within the clade [[Ankylopoda]].


⚫
==Classification==
⚫
Analyses based on [[Morphology (biology)|morphological]] data have generally recovered turtles either as non-[[diapsid]] reptiles nested within [[Parareptilia]], as early-diverging diapsids outside of [[Sauria]] (the clade formed by archosaurs and lepidosaurs), or as close relatives of Lepidosauria. A 2022 analysis by Simões and colleagues found a monophyletic Archelosauria using only morphological data for the first time, thus agreeing with most molecular analyses. Archelosauria was diagnosed by two unambiguous [[synapomorphies]] (shared derived traits): a [[sagittal crest]] on the [[supraoccipital]] bone, and the lack of an [[entepicondylar foramen]] on the [[humerus]].<ref name="Simões2022" />
Multiple [[sequence alignment]]s of [[DNA]] and [[protein]] sequences and [[phylogenetic]] inferences have shown that turtles are the closest living relatives to birds and crocodilians.<ref name="Shen2011"/><ref name="Tzika2011"/><ref name="Chiari2012"/> There are about 1000 [[ultra-conserved element]]s in the genome that are unique to turtles and [[archosaur]]s, but which are not found in [[lepidosaur]]s.<ref name="Crawford2012"/> Other genome-wide analyses also support this grouping.<ref name="Wang2013"/><ref name="Field2014"/>


Archelosauria was named in a 2015 article by Crawford ''et al''. The name is meant to evoke the archosaurs and chelonians (turtles), the two living subgroups of the clade. Crawford ''et al.'' defined Archelosauria as the clade formed by the descendants of the [[most recent common ancestor]] of ''[[Crocodylus niloticus]]'' (the Nile crocodile) and ''[[Testudo graeca]]'' (the Greek tortoise).<ref name="Crawford2015"/> A 2021 article by Joyce ''et al.'' modified the definition to specifically exclude the [[lizard]] ''[[Lacerta agilis]]'' from the group.<ref name="Joyce2021" />
==Etymology==
The [[clade]] name Archelosauria was coined in 2015 to evoke its two subclades, the '''''Ar'''''cho'''''sauria''''' and the '''''Chelo'''''nia.<ref name="Crawford2015"/>


Below is the phylogeny from Crawford ''et al.'', showing interrelationships of Testudines at family level down to [[Durocryptodira]]. Archelosauria was grouped within [[Sauria]] (the clade formed by archosaurs and lepidosaurs), as the sister branch to Lepidosauria, the clade containing lizards, [[snake]]s and the [[tuatara]].<ref name="Crawford2015"/>
⚫
==Classification==
Archelosauria is a group used to unite [[archosaurs]] with [[turtles]]. It was found that Archelosauria was grouped within [[Sauria]], as the sister branch to [[lizards]] and ''[[Sphenodon]]''. Below is the phylogeny from Crawford ''et al.'', showing interrelationships of Testudines at family level down to [[Durocryptodira]].<ref name="Crawford2015"/>


⚫
{{clade
{{clade| style=font-size:75%; line-height:75%
|label1=[[Sauria]]
|label1=[[Sauria]]
|1={{clade
|1={{clade
Line 36: Line 35:
|label1=[[Archosauria]]
|label1=[[Archosauria]]
|1={{clade
|1={{clade
|1=''[[Dinosauria]]''
|1=''[[Crocodylus]]''
|2=''[[Pseudosuchia]]'' }}
|2=''[[Gallus]]'' }}
|label2=[[Pantestudines]]
|label2=[[Testudines]]
|2={{clade
|2={{clade
|label1=[[Testudines]]
|label1=[[Pleurodira]]
|1={{clade
|1={{clade
|label1=[[Pleurodira]]
|1=[[Chelidae]]
|label2=[[Pelomedusoides]]
⚫
|1={{clade
|1=[[Chelidae]]
|2={{clade
|label2=[[Pelomedusoides]]
|1=[[Pelomedusidae]]
|2={{clade
|2=[[Podocnemididae]] }} }}
⚫
|1=[[Pelomedusidae]]
|2=[[Podocnemididae]] }} }}
|label2=[[Cryptodira]]
|label2=[[Cryptodira]]
|2={{clade
|2={{clade
Line 55: Line 52:
|1=''[[Carettochelys]]''
|1=''[[Carettochelys]]''
|2=[[Trionychidae]] }}
|2=[[Trionychidae]] }}
|2=[[Durocryptodira]] }} }} }} }} }} }}
|2=[[Durocryptodira]] }} }} }} }} }}


⚫
Analyses based on [[Morphology (biology)|morphological]] data have generally recovered turtles either as non-[[diapsid]] reptiles nested within [[Parareptilia]] (a group of [[Basal (phylogenetics)|basal]] reptiles that lived from the [[Carboniferous]] to the [[Triassic]]), as early-diverging diapsids outside of Sauria, or as close relatives of Lepidosauria. The hypothetical clade formed by turtles and lepidosaurs to the exclusion of archosaurs is known as [[Ankylopoda]].<ref name="Joyce2021" /> A 2022 analysis by Simões ''et al.'' found a monophyletic Archelosauria using only morphological data for the first time, thus agreeing with most molecular analyses. Archelosauria was diagnosed by two unambiguous [[synapomorphies]] (shared derived traits): a [[sagittal crest]] on the [[supraoccipital]] bone, and the lack of an [[entepicondylar foramen]] on the [[humerus]]. A cladogram adapted from their analysis is shown below:<ref name="Simões2022" />
==See also==

* [[Ankylopoda]], an alternative clade that places turtles as sister taxon to lepidosaurs.
{{Clade
|label1=[[Neodiapsida]]
|1={{Clade
|1=[[Younginiformes]]
|2={{Clade
⚫
|1=''[[Eunotosaurus]]''
|2={{Clade
|1=''[[Coelurosauravus]]''
|label2=[[Sauria]]
|2={{Clade
|label1=[[Lepidosauromorpha]]
|1={{Clade
|1=[[Sphenodontia]]
|2=[[Squamata]]
}}
|label2='''Archelosauria'''
|2={{Clade
|label1=[[Turtle|Testudines]]
|1={{Clade
|1=''[[Pappochelys]]''
|2={{Clade
|1=''[[Odontochelys]]''
|2={{Clade
|1=''[[Kayentachelys]]''
|2=''[[Proganochelys]]''
}} }} }}
⚫
|label2=[[Archosauromorpha]]
|2={{Clade
|1={{Clade
|1=[[Ichthyosauromorpha]]
|2={{Clade
|1=[[Sauropterygia]]
|2=[[Thalattosauria]]
}} }}
|2={{Clade
|1=[[Protorosauria]]
|2={{Clade
|1={{Clade
|1=[[Allokotosauria]]
|2=[[Rhynchosauria]]
}}
|2={{Clade
|1=''[[Prolacerta]]''
|2=[[Archosauriformes]]
}} }} }} }} }} }} }} }} }} }}


==References==
==References==
{{Reflist | refs=
{{Reflist | refs=
<ref name="Chiari2012">{{Cite journal |last1=Chiari |first1=Ylenia |last2=Cahais |first2=Vincent |last3=Galtier |first3=Nicolas |last4=Delsuc |first4=Frédéric |date=2012 |title=Phylogenomic analyses support the position of turtles as the sister group of birds and crocodiles (Archosauria) |url= |journal=BMC Biology |language=en |volume=10 |issue=1 |pages=65 |doi=10.1186/1741-7007-10-65 |issn=1741-7007 |pmc=3473239 |pmid=22839781}}</ref>
<ref name="Chiari2012">{{Cite journal |last1=Chiari |first1=Y. |last2=Cahais |first2=V. |last3=Galtier |first3=N. |last4=Delsuc |first4=F. |date=2012 |title=Phylogenomic analyses support the position of turtles as the sister group of birds and crocodiles (Archosauria) |url= |journal=BMC Biology |language=en |volume=10 |issue=1 |pages=65 |doi=10.1186/1741-7007-10-65 |issn=1741-7007 |pmc=3473239 |pmid=22839781}}</ref>
<ref name="Crawford2012">{{Cite journal |last1=Crawford |first1=Nicholas G. |last2=Faircloth |first2=Brant C. |last3=McCormack |first3=John E. |last4=Brumfield |first4=Robb T. |last5=Winker |first5=Kevin |last6=Glenn |first6=Travis C. |date=2012-10-23 |title=More than 1000 ultraconserved elements provide evidence that turtles are the sister group of archosaurs |url= |journal=Biology Letters |volume=8 |issue=5 |pages=783–786 |doi=10.1098/rsbl.2012.0331 |pmc=3440978 |pmid=22593086}}</ref>
<ref name="Crawford2012">{{Cite journal |last1=Crawford |first1=N. G. |last2=Faircloth |first2=B. C. |last3=McCormack |first3=J. E. |last4=Brumfield |first4=R. T. |last5=Winker |first5=K. |last6=Glenn |first6=T. C. |date=2012-10-23 |title=More than 1000 ultraconserved elements provide evidence that turtles are the sister group of archosaurs |url= |journal=Biology Letters |volume=8 |issue=5 |pages=783–786 |doi=10.1098/rsbl.2012.0331 |pmc=3440978 |pmid=22593086}}</ref>
<ref name="Crawford2015">{{Cite journal |last1=Crawford |first1=Nicholas G. |last2=Parham |first2=James F. |last3=Sellas |first3=Anna B. |last4=Faircloth |first4=Brant C. |last5=Glenn |first5=Travis C. |last6=Papenfuss |first6=Theodore J. |last7=Henderson |first7=James B. |last8=Hansen |first8=Madison H. |last9=Simison |first9=W. Brian |date=2015-02-01 |title=A phylogenomic analysis of turtles |url=http://www.sciencedirect.com/science/article/pii/S1055790314003819 |journal=Molecular Phylogenetics and Evolution |language=en |volume=83 |pages=250–257 |doi=10.1016/j.ympev.2014.10.021 |pmid=25450099 |issn=1055-7903}}</ref>
<ref name="Crawford2015">{{Cite journal |last1=Crawford |first1=N. G. |last2=Parham |first2=J. F. |last3=Sellas |first3=A. B. |last4=Faircloth |first4=B. C. |last5=Glenn |first5=T. C. |last6=Papenfuss |first6=T. J. |last7=Henderson |first7=J. B. |last8=Hansen |first8=M. H. |last9=Simison |first9=W. B. |date=2015-02-01 |title=A phylogenomic analysis of turtles |url=http://www.sciencedirect.com/science/article/pii/S1055790314003819 |journal=Molecular Phylogenetics and Evolution |language=en |volume=83 |pages=250–257 |doi=10.1016/j.ympev.2014.10.021 |pmid=25450099 |issn=1055-7903}}</ref>
<ref name="Field2014">{{cite journal |last1=Field |first1=Daniel J. |last2=Gauthier |first2=Jacques A. |last3=King |first3=Benjamin L. |last4=Pisani |first4=Davide |last5=Lyson |first5=Tyler |last6=Peterson |first6=Kevin J. |title=Toward consilience in reptile phylogeny: miRNAs support an archosaur, not lepidosaur, affinity for turtles |journal=Evolution & Development |date=July–August 2014 |volume=16 |issue=4 |pages=189–196 |url=http://danieljfield.com/Home/Publications_files/Field%20et%20al.%202014%20Toward%20consilience%20in%20reptile%20phylogeny%3A%20miRNAs%20support%20an%20archosaur,%20not%20lepidosaur,%20affinity%20for%20turtle.pdf |access-date=5 July 2015 |doi=10.1111/ede.12081 |pmid=24798503 |pmc=4215941}}</ref>
<ref name="Field2014">{{cite journal |last1=Field |first1=D. J. |last2=Gauthier |first2=J. A. |last3=King |first3=B. L. |last4=Pisani |first4=D. |last5=Lyson |first5=T. |last6=Peterson |first6=K. J. |title=Toward consilience in reptile phylogeny: miRNAs support an archosaur, not lepidosaur, affinity for turtles |journal=Evolution & Development |date=July–August 2014 |volume=16 |issue=4 |pages=189–196 |doi=10.1111/ede.12081 |pmid=24798503 |pmc=4215941}}</ref>
<ref name="Joyce2021">{{cite journal |last1=Joyce |first1=W. G. |last2=Anquetin |first2=J. |last3=Cadena |first3=E.-A. |last4=Claude |first4=J. |last5=Danilov |first5=I. G. |last6=Evers |first6=S. W. |last7=Ferreira |first7=G. S. |last8=Gentry |first8=A. D. |last9=Georgalis |first9=G. L. |last10=Lyson |first10=T. R. |last11=Pérez-García |first11=A. |last12=Rabi |first12=M. |last13=Sterli |first13=J. |last14=Vitek |first14=N. S. |last15=Parham |first15=J. F. |title=A nomenclature for fossil and living turtles using phylogenetically defined clade names |journal=Swiss Journal of Palaeontology |date=2021 |volume=140 |issue=1 |pages=5 |doi=10.1186/s13358-020-00211-x}}</ref>
<ref name="Shen2011">{{Cite journal |last1=Shen |first1=X.-X. |last2=Liang |first2=D. |last3=Wen |first3=J.-Z. |last4=Zhang |first4=P. |date=2011-12-01 |title=Multiple Genome Alignments Facilitate Development of NPCL Markers: A Case Study of Tetrapod Phylogeny Focusing on the Position of Turtles |url=https://academic.oup.com/mbe/article-lookup/doi/10.1093/molbev/msr148 |journal=Molecular Biology and Evolution |language=en |volume=28 |issue=12 |pages=3237–3252 |doi=10.1093/molbev/msr148 |pmid=21680872 |issn=0737-4038|doi-access=free }}</ref>
<ref name="Shen2011">{{Cite journal |last1=Shen |first1=X.-X. |last2=Liang |first2=D. |last3=Wen |first3=J.-Z. |last4=Zhang |first4=P. |date=2011-12-01 |title=Multiple Genome Alignments Facilitate Development of NPCL Markers: A Case Study of Tetrapod Phylogeny Focusing on the Position of Turtles |url=https://academic.oup.com/mbe/article-lookup/doi/10.1093/molbev/msr148 |journal=Molecular Biology and Evolution |language=en |volume=28 |issue=12 |pages=3237–3252 |doi=10.1093/molbev/msr148 |pmid=21680872 |issn=0737-4038|doi-access=free }}</ref>
<ref name="Simões2022">{{cite journal |last1=Simões |first1=T. R. |last2=Kammerer |first2=C. F. |last3=Caldwell |first3=M. W. |last4=Pierce |first4=S. E. |title=Successive climate crises in the deep past drove the early evolution and radiation of reptiles |journal=Science Advances |date=2022 |volume=8 |issue=33 |pages=eabq1898 |doi=10.1126/sciadv.abq1898}}</ref>
<ref name="Simões2022">{{cite journal |last1=Simões |first1=T. R. |last2=Kammerer |first2=C. F. |last3=Caldwell |first3=M. W. |last4=Pierce |first4=S. E. |title=Successive climate crises in the deep past drove the early evolution and radiation of reptiles |journal=Science Advances |date=2022 |volume=8 |issue=33 |pages=eabq1898 |doi=10.1126/sciadv.abq1898}}</ref>
<ref name="Tzika2011">{{Cite journal |last1=Tzika |first1=Athanasia C |last2=Helaers |first2=Raphaël |last3=Schramm |first3=Gerrit |last4=Milinkovitch |first4=Michel C |date=2011 |title=Reptilian-transcriptome v1.0, a glimpse in the brain transcriptome of five divergent Sauropsida lineages and the phylogenetic position of turtles |url= |journal=EvoDevo |language=en |volume=2 |issue=1 |pages=19 |doi=10.1186/2041-9139-2-19 |issn=2041-9139 |pmc=3192992 |pmid=21943375}}</ref>
<ref name="Tzika2011">{{Cite journal |last1=Tzika |first1=A. C. |last2=Helaers |first2=R. |last3=Schramm |first3=G. |last4=Milinkovitch |first4=M. C. |date=2011 |title=Reptilian-transcriptome v1.0, a glimpse in the brain transcriptome of five divergent Sauropsida lineages and the phylogenetic position of turtles |url= |journal=EvoDevo |language=en |volume=2 |issue=1 |pages=19 |doi=10.1186/2041-9139-2-19 |issn=2041-9139 |pmc=3192992 |pmid=21943375}}</ref>
<ref name="Wang2013">{{cite journal |last1=Wang |title=The draft genomes of soft-shell turtle and green sea turtle yield insights into the development and evolution of the turtle-specific body plan |journal=Nature Genetics |date=27 March 2013 |volume=45 |issue=701–706 |pages=701–6 |doi=10.1038/ng.2615 |first1=Zhuo |pmid=23624526 |pmc=4000948}}</ref>
<ref name="Wang2013">{{cite journal |last1=Wang |first1=Z. |title=The draft genomes of soft-shell turtle and green sea turtle yield insights into the development and evolution of the turtle-specific body plan |journal=Nature Genetics |date=27 March 2013 |volume=45 |issue=701–706 |pages=701–6 |doi=10.1038/ng.2615 |pmid=23624526 |pmc=4000948}}</ref>
}}
}}



{{Chordata}}
{{Chordata}}
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[[Category:Reptile taxonomy]]
[[Category:Reptile taxonomy]]
[[Category:Saurians]]
[[Category:Saurians]]
[[Category:Turtles]]

Revision as of 19:27, 17 September 2022

Archelosauria
Temporal range: Lopingian–Recent Possible Capitanian record
Proganochelys quenstedti
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Sauria
Clade: Archelosauria
Crawford et al., 2015[1]
Subgroups

Archelosauria is a clade grouping turtles and archosaurs (birds and crocodilians) and their fossil relatives, to the exclusion of lepidosaurs (the clade containing lizards, snakes and the tuatara). The majority of phylogenetic analyses based on molecular data (e.g. DNA and proteins) have supported a sister-group relationship between turtles and archosaurs. On the other hand, Archelosauria has not been supported by most morphological analyses, which have instead found turtles to either be descendants of parareptiles, early-diverging diapsids outside of Sauria, or close relatives of lepidosaurs within the clade Ankylopoda.

Classification

Multiple sequence alignments of DNA and protein sequences and phylogenetic inferences have shown that turtles are the closest living relatives to birds and crocodilians.[2][3][4] There are about 1000 ultra-conserved elements in the genome that are unique to turtles and archosaurs, but which are not found in lepidosaurs.[5] Other genome-wide analyses also support this grouping.[6][7]

Archelosauria was named in a 2015 article by Crawford et al. The name is meant to evoke the archosaurs and chelonians (turtles), the two living subgroups of the clade. Crawford et al. defined Archelosauria as the clade formed by the descendants of the most recent common ancestor of Crocodylus niloticus (the Nile crocodile) and Testudo graeca (the Greek tortoise).[1] A 2021 article by Joyce et al. modified the definition to specifically exclude the lizard Lacerta agilis from the group.[8]

Below is the phylogeny from Crawford et al., showing interrelationships of Testudines at family level down to Durocryptodira. Archelosauria was grouped within Sauria (the clade formed by archosaurs and lepidosaurs), as the sister branch to Lepidosauria, the clade containing lizards, snakes and the tuatara.[1]

Sauria

Analyses based on morphological data have generally recovered turtles either as non-diapsid reptiles nested within Parareptilia (a group of basal reptiles that lived from the Carboniferous to the Triassic), as early-diverging diapsids outside of Sauria, or as close relatives of Lepidosauria. The hypothetical clade formed by turtles and lepidosaurs to the exclusion of archosaurs is known as Ankylopoda.[8] A 2022 analysis by Simões et al. found a monophyletic Archelosauria using only morphological data for the first time, thus agreeing with most molecular analyses. Archelosauria was diagnosed by two unambiguous synapomorphies (shared derived traits): a sagittal crest on the supraoccipital bone, and the lack of an entepicondylar foramen on the humerus. A cladogram adapted from their analysis is shown below:[9]

Neodiapsida

References

  1. ^ a b c Crawford, N. G.; Parham, J. F.; Sellas, A. B.; Faircloth, B. C.; Glenn, T. C.; Papenfuss, T. J.; Henderson, J. B.; Hansen, M. H.; Simison, W. B. (2015-02-01). "A phylogenomic analysis of turtles". Molecular Phylogenetics and Evolution. 83: 250–257. doi:10.1016/j.ympev.2014.10.021. ISSN 1055-7903. PMID 25450099.
  2. ^ Shen, X.-X.; Liang, D.; Wen, J.-Z.; Zhang, P. (2011-12-01). "Multiple Genome Alignments Facilitate Development of NPCL Markers: A Case Study of Tetrapod Phylogeny Focusing on the Position of Turtles". Molecular Biology and Evolution. 28 (12): 3237–3252. doi:10.1093/molbev/msr148. ISSN 0737-4038. PMID 21680872.
  3. ^ Tzika, A. C.; Helaers, R.; Schramm, G.; Milinkovitch, M. C. (2011). "Reptilian-transcriptome v1.0, a glimpse in the brain transcriptome of five divergent Sauropsida lineages and the phylogenetic position of turtles". EvoDevo. 2 (1): 19. doi:10.1186/2041-9139-2-19. ISSN 2041-9139. PMC 3192992. PMID 21943375.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  4. ^ Chiari, Y.; Cahais, V.; Galtier, N.; Delsuc, F. (2012). "Phylogenomic analyses support the position of turtles as the sister group of birds and crocodiles (Archosauria)". BMC Biology. 10 (1): 65. doi:10.1186/1741-7007-10-65. ISSN 1741-7007. PMC 3473239. PMID 22839781.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  5. ^ Crawford, N. G.; Faircloth, B. C.; McCormack, J. E.; Brumfield, R. T.; Winker, K.; Glenn, T. C. (2012-10-23). "More than 1000 ultraconserved elements provide evidence that turtles are the sister group of archosaurs". Biology Letters. 8 (5): 783–786. doi:10.1098/rsbl.2012.0331. PMC 3440978. PMID 22593086.
  6. ^ Wang, Z. (27 March 2013). "The draft genomes of soft-shell turtle and green sea turtle yield insights into the development and evolution of the turtle-specific body plan". Nature Genetics. 45 (701–706): 701–6. doi:10.1038/ng.2615. PMC 4000948. PMID 23624526.
  7. ^ Field, D. J.; Gauthier, J. A.; King, B. L.; Pisani, D.; Lyson, T.; Peterson, K. J. (July–August 2014). "Toward consilience in reptile phylogeny: miRNAs support an archosaur, not lepidosaur, affinity for turtles". Evolution & Development. 16 (4): 189–196. doi:10.1111/ede.12081. PMC 4215941. PMID 24798503.
  8. ^ a b Joyce, W. G.; Anquetin, J.; Cadena, E.-A.; Claude, J.; Danilov, I. G.; Evers, S. W.; Ferreira, G. S.; Gentry, A. D.; Georgalis, G. L.; Lyson, T. R.; Pérez-García, A.; Rabi, M.; Sterli, J.; Vitek, N. S.; Parham, J. F. (2021). "A nomenclature for fossil and living turtles using phylogenetically defined clade names". Swiss Journal of Palaeontology. 140 (1): 5. doi:10.1186/s13358-020-00211-x.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  9. ^ Simões, T. R.; Kammerer, C. F.; Caldwell, M. W.; Pierce, S. E. (2022). "Successive climate crises in the deep past drove the early evolution and radiation of reptiles". Science Advances. 8 (33): eabq1898. doi:10.1126/sciadv.abq1898.
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