Jennings was so struck by the continued production of hereditarily diverse clones at conjugation, even after many successive inbreedings, that he undertook to examine the matter mathematically. As a result, general formulae for the results of diverse systems of mating were published in a series of papers between 1912 and 1917; these were one of the main seeds from which the whole field of mathematical genetics developed.
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Barkan, E (1991), "Reevaluating progressive eugenics: Herbert Spencer Jennings and the 1924 immigration legislation.", Journal of the history of biology24 (1): 91–112, doi:10.1007/BF00130475, PMID11612743
Sonneborn, T M (1974), "Herbert Spencer Jennings.", Biographical memoirs. National Academy of Sciences (U.S.)47: 143–223, PMID11615625