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Pieter Dirk Nieuwkoop (July 17, 1917 - September 18 1996) was a Dutch embryologist and developmental biologist who studied the early embryonic development of amphibians and reptiles. He is noted for the discovery and description of the process of endomesoderm^[a] induction in the amphibian blastula.

Personal life and Career

Pieter Nieuwkoop was born in Enshele, The Netherlands, the second son of a secondary school teacher. He began his studies at the State University of Utrecht in 1934, gaining a BSc in 1937 and an MSc in 1940. He carried out his doctoral studies there under Chris P. Raven during the German occupation of The Netherlands, but was unable to graduate during the war as he opted to write his thesis in English. He graduated Cum Laude in July 1946.

From 1942-49 Nieuwkoop worked as a scientific assistant in the department of Zoology at Utrecht. While holding this position he was appointed as deputy director of the Hubrecht Laboratory.

Research

Publications

Notes

^ An embryonic cell layer which is destined to give rise to both endodermal and mesodermal tissues.^[1]

References

^ Test

[2] An embryonic cell layer which is destined to give rise to both endodermal and mesodermal tissues.^[1]

[endomesoderm_def-1] Test

[a]

[1]

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 ==Personal life and Career==
-Pieter Nieuwkoop was born in Enshele, The Netherlands, the second son of a secondary school teacher. He began his studies at the State University of Utrecht in 1934, gaining a BSc in 1937 and an MSc in 1940. He carried out his doctoral studies there under Chris P. Raven during the German occupation of The Netherlands, but was unable to graduate during the war as he opted to write his thesis in English. He graduated Cum Laude in July 1946. From 1942-49 he worked as a scientific assistant in the department of Zoology at Utrecht. While holding this position he was appointed as deputy director of the Hubrecht Laboratory.
+Pieter Nieuwkoop was born in Enshele, The Netherlands, the second son of a secondary school teacher. He began his studies at the State University of Utrecht in 1934, gaining a BSc in 1937 and an MSc in 1940. He carried out his doctoral studies there under Chris P. Raven during the German occupation of The Netherlands, but was unable to graduate during the war as he opted to write his thesis in English. He graduated Cum Laude in July 1946.
+From 1942-49 Nieuwkoop worked as a scientific assistant in the department of Zoology at Utrecht. While holding this position he was appointed as deputy director of the Hubrecht Laboratory.
 ==Research==