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''Alaskacephale'', due to the lack of fossil remains, is poorly known. Despite this, much of its characteristics such as the presence of bony nodes and structures along the squamosal, prove it was a pachycephalosaur. Later phylogenetic analyses suggest it was a close relative of its southern neighbor ''Pachycephalosaurus''. However, ''Alaskacephale'' is distinguished by traits such as two rows of nodes along the squamosal, whereas other pachycephalosaurs bear one row.
''Alaskacephale'', due to the lack of fossil remains, is poorly known. Despite this, much of its characteristics such as the presence of bony nodes and structures along the squamosal, prove it was a pachycephalosaur. Later phylogenetic analyses suggest it was a close relative of its southern neighbor ''Pachycephalosaurus''. However, ''Alaskacephale'' is distinguished by traits such as two rows of nodes along the squamosal, whereas other pachycephalosaurs bear one row.


It was discovered in the rocks deriving from the Prince Creek Formation, which bears a variety of fossils, many of them similar to other previously known Laramidian taxa. ''Alaskacephale'' coexisted with dinosaurs like the ceratopsian ''Pachyrhinosaurus'', ornithopod ''Edmontosaurus'', and tyrannosaurid ''Nanuqsaurus'' in additon to mammals like ''Unnuakomys<u>.</u>''
It was discovered in the rocks deriving from the Prince Creek Formation, which bears a variety of fossils, many of them similar to other previously known Laramidian taxa. ''Alaskacephale'' coexisted with dinosaurs like the ceratopsian ''Pachyrhinosaurus'', ornithopod ''Edmontosaurus'', and tyrannosaurid ''Nanuqsaurus'' in additon to mammals like ''Unnuakomys.''


==Discovery==
==Discovery==

Revision as of 20:03, 22 September 2023

Alaskacephale
Temporal range: Latest Campanian, 72–71 Ma
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Ornithischia
Clade: Pachycephalosauria
Family: Pachycephalosauridae
Genus: Alaskacephale
Sullivan, 2006
Species:
A. gangloffi
Binomial name
Alaskacephale gangloffi
Sullivan, 2006

Alaskacephale was a genus of pachycephalosaurid, a group of dome-headed, herbivorous ornithischian dinosaurs, that lived during the Campanian stage of the Late Cretaceous period in what is now northern Alaska. Alaskacephale is one of the few known Arctic dinosaurs and was found in the Prince Creek Formation, which preserves a menagerie of fossils. The only known specimen, a squamosal bone, was found in 1999 and later described in 2005. However, Alaskacephale was not formally named until the next year.

Alaskacephale, due to the lack of fossil remains, is poorly known. Despite this, much of its characteristics such as the presence of bony nodes and structures along the squamosal, prove it was a pachycephalosaur. Later phylogenetic analyses suggest it was a close relative of its southern neighbor Pachycephalosaurus. However, Alaskacephale is distinguished by traits such as two rows of nodes along the squamosal, whereas other pachycephalosaurs bear one row.

It was discovered in the rocks deriving from the Prince Creek Formation, which bears a variety of fossils, many of them similar to other previously known Laramidian taxa. Alaskacephale coexisted with dinosaurs like the ceratopsian Pachyrhinosaurus, ornithopod Edmontosaurus, and tyrannosaurid Nanuqsaurus in additon to mammals like Unnuakomys.

Discovery

In 1999 during an expedition by the University of Alaska, an isolated pachycephalosaur squamosal was unearthed from bluffs on the bank of the Colville River in North Slope Borough, Alaska. Strata (rock layers) deriving from this area come from the latest Campanian-aged (72-71 mya) Prince Creek Formation.[1][2] The fossil was then deposited in the University of Alaska Museum and described in 2005 by Roland Gangloff and colleagues as belonging to a pachycephalosaurine closely related to, and possibly, Pachycephalosaurus.[2] However, the next year paleontologist Robert Sullivan described it as a new genus and species, Alaskacephale gangloffi. The genus name comes from Alaska, the state the holotype (name-bearing specimen) was found in, and cephale, from the Greek kephale meaning "head". Its species name is in honor of Roland Gangloff, who published the earlier 2005 description.[3]

The dimensions of this bone suggest that A. gangloffi was about half the size of Pachycephalosaurus wyomingensis or three quarters the size of Prenocephale, and about the same size as Sphaerotholus edmontonensis and Foraminacephale.[2]

Description

The dimensions of this bone suggest that A. gangloffi was about half the size of Pachycephalosaurus wyomingensis or three quarters the size of Prenocephale, and about the same size as Sphaerotholus edmontonensis and Foraminacephale.[2] The specimen was previously described by Gangloff et al. (2005) as an unnamed pachycephalosaurid, possibly a Pachycephalosaurus. Gangloff et al. described the squamosal as having an interdigitated suture with the quadrate, a feature previously described only in Pachycephalosaurus. Sullivan (2006) opined that this "suture" is instead a breakage point in both Alaskacephale and Pachycephalosaurus, so it could not be used to unite the two taxa

Classification

Within the tribe Pachycephalosaurini, Alaskacephale is very closely related to Pachycephalosaurus wyomingensis. Below is a cladogram from Evans et al., 2021.[4]

Paleoenvironment

Alaskacephale lived alongside many other dinosaurs during latest interval of the Campanian stage. Having resided at an estimated 80°–85°N paleolatitude, the area Alaskacephale thrived in experienced climatic extremes unlike that experienced by most other dinosaurs. The temperature of this Northern environment would've ranged from around 10 to 12°C during the warmer months and about -2°C ± 3.9°C during the colder months. As well as this, the environment would have faced 120 days of continuous low-light conditions during the Winter. In contrast to the contemporary large herbivores Edmontosaurus and Pachyrhinosaurus, which apparently preferred coastal lowland and upland environments respectively, Nanuqsaurus appears to have been fairly ubiquitous throughout the Prince Creek landscape.[5][2]

Other animals that lived alongside Nanuqsaurus include the following: unnamed leptoceratopsid, the hadrosaurid Edmontosaurus (similar to E. regalis),[6] an unnamed lambeosaurine, an unnamed thescelosaurine (mentioned to be similar to Parkosaurus and Thescelosaurus), an unnamed orodromine (mentioned to be similar to Orodromeus), the ceratopsid Pachyrhinosaurus perotorum, the pachycephalosaurid Alaskacephale, a large troodontid referred to the dubious genus Troodon, the dromaeosaurid Dromaeosaurus, the dromaeosaurid Richardoestesia (similar to R. isosceles), an unnamed saurornitholestine, an unnamed ornithomimosaur, and multiple avialans.[7] As well as this, several mammals, including the metatherian Unnuakomys,[8] the eutherian Gypsonictops, both an unnamed and named multituberculate, the latter being Cimolodon, and finally an indeterminate marsupial. Interestingly, due to the cooler conditions of this habitat, many otherwise common ectothermic clades lack representation entirely in the Prince Creek Formation, suggesting that all the animals that did thrive in these extreme latitudes were indeed endotherms to some degree.

See also

References

  1. ^ Fiorillo, Anthony R.; Tykoski, Ronald S. (2014-03-12). "A Diminutive New Tyrannosaur from the Top of the World". PLOS ONE. 9 (3): e91287. doi:10.1371/journal.pone.0091287. ISSN 1932-6203. PMC 3951350. PMID 24621577.{{cite journal}}: CS1 maint: PMC format (link) CS1 maint: unflagged free DOI (link)
  2. ^ a b c d e Gangloff, R.A.; A.R. Fiorillo; D.W. Norton (2005). "The first pachycephalosaurine (Dinosauria) from the Paleo-Arctic of Alaska and its paleogeographic implications". Journal of Paleontology. 79 (5): 997–1001. doi:10.1666/0022-3360(2005)079[0997:tfpdft]2.0.co;2. S2CID 130669713.
  3. ^ Sullivan, RM (2006). "A taxonomic review of the Pachycephalosauridae (Dinosauria: Ornithischia)". New Mexico Museum of Natural History and Science Bulletin (35): 347–365.
  4. ^ Evans, David; Brown, Caleb M.; You, Hailu; Campione, Nicolás E. (2021). "Description and revised diagnosis of Asia's first recorded pachycephalosaurid, Sinocephale bexelli gen. nov., from the Upper Cretaceous of Inner Mongolia, China". Canadian Journal of Earth Sciences. 58 (10): 981–992. Bibcode:2021CaJES..58..981E. doi:10.1139/cjes-2020-0190. S2CID 244227050.
  5. ^ Herman, Alexei B.; Spicer, Robert A.; Spicer, Teresa E. V. (2016-01-01). "Environmental constraints on terrestrial vertebrate behaviour and reproduction in the high Arctic of the Late Cretaceous". Palaeogeography, Palaeoclimatology, Palaeoecology. Selected papers based on Geological Society of America, Annual Meeting, Theme Session 241, Ancient Polar Ecosystems and Climate History in Deep Time, Denver, Colorado, USA, 30 October 2013. 441: 317–338. doi:10.1016/j.palaeo.2015.09.041. ISSN 0031-0182.
  6. ^ Takasaki, Ryuji; Fiorillo, Anthony R.; Tykoski, Ronald S.; Kobayashi, Yoshitsugu (2020-05-06). "Re-examination of the cranial osteology of the Arctic Alaskan hadrosaurine with implications for its taxonomic status". PLOS ONE. 15 (5): e0232410. Bibcode:2020PLoSO..1532410T. doi:10.1371/journal.pone.0232410. ISSN 1932-6203. PMC 7202651. PMID 32374777.
  7. ^ Cite error: The named reference :1 was invoked but never defined (see the help page).
  8. ^ Eberle, Jaelyn J.; Clemens, William A.; McCarthy, Paul J.; Fiorillo, Anthony R.; Erickson, Gregory M.; Druckenmiller, Patrick S. (2019-11-02). "Northernmost record of the Metatheria: a new Late Cretaceous pediomyid from the North Slope of Alaska". Journal of Systematic Palaeontology. 17 (21): 1805–1824. doi:10.1080/14772019.2018.1560369. ISSN 1477-2019.