|Long-tailed tit Aegithalos caudatus|
|Song of the American bushtit,|
The bushtits or long-tailed tits, Aegithalidae, are a family of small, drab passerine birds with moderately long tails. The family contains 13 species in four genera, all but one of which are found in Eurasia. Bushtits are active birds, moving almost constantly while they forage for insects in shrubs and trees. During non-breeding season, birds live in flocks of up to 50 individuals. Several bushtit species display cooperative breeding behavior, also called helpers at the nest.
Distribution and habitat
All the Aegithalidae are forest birds, particularly forest edge and understory habitats. The species in the genus Aegithalos prefer deciduous or mixed deciduous forests, while the tiny pygmy tit is found mostly in montane coniferous forest. The bushtit is found in a wide range of habitats, including on occasion sagebrush and other arid shrublands, but is most common in mixed woodland. Most species in this family live in mountainous habitats in and around the Himalayas, and all are distributed in Eurasia except the American bushtit, which is native to western North America. The long-tailed tit has the most widespread distribution of any species of Aegithalidae, occurring across Eurasia from Britain to Japan. Two species in contrast have tiny distributions, the Burmese bushtit, which is entirely restricted to two mountains in Burma, and the pygmy bushtit, which is restricted to the mountains of western Java. The species in this family are generally not migratory, although the long-tailed tit is prone to dispersing in the northern edges of its range (particularly in Siberia). Many mountainous species move to lower ground during the winter.
They are small birds, measuring 9 to 14 cm (3.5–5.5 in) in length, including the relatively long tail, and weighing just 4.5 to 9 g (0.16–0.32 oz). Their plumage is typically dull grey or brown, although some species have white markings and the long-tailed tit has some pinkish colour. In contrast to the rest of the family the two Leptopoecile tit-warblers are quite brightly coloured, having violet and blue plumage. The crested tit-warbler is the only member of the family to have a crest. The bills in this family are tiny, short and conical in shape. The wings are short and rounded and the legs are relatively long.
Birds in this family live in flocks ranging from 4 to 50 individuals. Flocks form as soon as one breeding season finishes and last until the next one begins. They maintain contact with "churring" calls; their songs are quiet. Other species of birds, such as tits or warblers, will occasionally join the flock to forage.
Diet and feeding
They are omnivorous, primarily eating insects and other invertebrates  such as leafhoppers, treehoppers, aphids, scale insects, and caterpillars. Plant material, such as berries or seeds, is taken occasionally during the winter. The family generally forages arboreally, usually in the shrub layer or canopy, and seldom visits the ground. Prey is generally gleaned from branches, leaves and buds. Less frequently, prey is taken in the air. While foraging, this agile family may hang upside down on branches (although this behaviour is not thought to occur in the tit-warblers) and even manipulate branches and leaves in order to locate hidden food.
The family generally has a monogamous breeding system. Pairs may be aided by helpers, where a related individual (or more than one) helps the established pair raise the young. This has been recorded in at least four of the species; further research is required to see if the behavior carries over to other members of the family. Aegithalids make domed, bag-like, nests of woven cobwebs and lichen, which they line with feathers. They make the nests in trees with thick foliage, making them difficult for predators to find. The clutch comprises 6 to 10 white eggs, which in many of the species have red speckles. Adults incubate the eggs for 13 to 14 days; young stay in the nest for 16 to 17 days. In at least four of the species (the long-tailed tit, the American bushtit, the black-throated bushtit, and silver-throated bushtit), only the female incubates. Young chicks are fed exclusively on insects and spiders.
The family Aegithalidae (as a subfamily Aegithalinae) was introduced by the German naturalist Ludwig Reichenbach in 1850. The name comes from the Ancient Greek word aigithalos for a tit. Aristotle recognised three species: the long-tailed tit, the great tit, and the Eurasian blue tit.
The pygmy bushtit is placed in this family because it moves around in flocks and its nests resemble the long-tailed tits', but information about it is so scanty that the placement is only provisional. The Burmese bushtit is sometimes treated as conspecific with the black-browed bushtit. The American bushtit was once thought to belong to the chickadee family, but it has distinctive behavioral habits, especially when it comes to nesting.
There are 13 species in 3 genera.
- Long-tailed tit, Aegithalos caudatus
- Silver-throated bushtit, Aegithalos glaucogularis
- White-cheeked bushtit, Aegithalos leucogenys
- Black-throated bushtit, Aegithalos concinnus
- White-throated bushtit, Aegithalos niveogularis
- Rufous-fronted bushtit, Aegithalos iouschistos
- Black-browed bushtit, Aegithalos bonvaloti
- Burmese bushtit, Aegithalos sharpei
- Sooty bushtit, Aegithalos fuliginosus
- Pygmy bushtit, Aegithalos exilis
- American bushtit, Psaltriparus minimus
- Li, J.; Lv, L.; Wang, Y.; Xi, B.; Zhang, Z. (2012). "Breeding biology of two sympatric Aegithalos tits with helpers at the nest" (PDF). Journal of Ornithology. 153 (2): 273–283. doi:10.1007/s10336-011-0740-z.
- Harrap, Simon (2008). "Family Aegithalidae (Long-tailed Tits)". In del Hoyo, Josep; Elliott, Andrew; Christie, David. Handbook of the Birds of the World. Volume 13, Penduline-tits to Shrikes. Barcelona: Lynx Edicions. pp. 76–101. ISBN 978-84-96553-45-3.
- Perrins, Christopher M. (2003). "Long-tailed Tits". In Perrins, Christopher. The Firefly Encyclopedia of Birds. Firefly Books. pp. 556–557. ISBN 1-55297-777-3.
- Perrins, C. (1991). Forshaw, Joseph, ed. Encyclopaedia of Animals: Birds. London: Merehurst Press. p. 202. ISBN 1-85391-186-0.
- Kaufman, K. (1996). Lives of North American Birds. Houghton Mifflin: Boston.
- Reichenbach, Ludwig (1850). Die vollständigste Naturgeschichte der Vögel (in German). Apt. II Band I. Dresden: Expedition der Vollständigsten Naturgeschichte. Plate LXII.
- Bock, Walter J. (1994). History and Nomenclature of Avian Family-Group Names. Bulletin of the American Museum of Natural History. Number 222. New York: American Museum of Natural History. pp. 153, 253.
- Jobling, J.A. (2018). del Hoyo, J.; Elliott, A.; Sargatal, J.; Christie, D.A.; de Juana, E., eds. "Key to Scientific Names in Ornithology". Handbook of the Birds of the World Alive. Lynx Edicions. Retrieved 21 March 2018.
- Gill, Frank; Donsker, David, eds. (2017). "Bushtits, leaf warblers, reed warblers". World Bird List Version 8.1. International Ornithologists' Union. Retrieved 21 March 2018.
- Long-tailed Tits videos on the Internet Bird Collection
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