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Temporal range: Cretaceous(?)[1] - Recent
An apertural view of a shell of Angiola fasciata.
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Subclass: Caenogastropoda
Superfamily: Cerithioidea
Family: Planaxidae
Gray, 1850

Planaxidae, commonly called planaxids or clusterwinks, are a taxonomic family of small and minute sea snails, pantropical marine gastropod molluscs in the superfamily Cerithoidea. They live on rocky shores in the littoral zone of the tropics and subtropics.[2]


Planaxids are known as clusterwinks because of their habit of clustering together in concealed, moist locations when the tide is out. They have conical shells resembling periwinkles, except for the wide, shallow anterior canals. They brood their embryos in a chamber behind their heads, releasing them into the sea as veliger larvae to form part of the plankton.[2]


The following two subfamilies are recognized in the taxonomy of Bouchet & Rocroi (2005):

  • Planaxinae Gray, 1850
  • Fossarinae A. Adams, 1860 - previously in a family of its own, named Fossaridae


The family Planaxidae comprises the following genera:[3] The following genera are recognised: Planaxinae

  • Angiola Dall, 1926: synonym of Hinea Gray, 1847
  • Leucostoma Swainson, 1840: synonym of Fissilabia MacGillivray, 1836 (invalid: junior homonym of Leucostoma Meigen, 1803 [Diptera])
  • Quoyia Gray, 1839: synonym of Fissilabia MacGillivray, 1836


  • Fossar Gray, 1847: synonym of Fossarus Philippi, 1841 (invalid: unjustified emendation of Fossarus)
  • Maravignia Aradas & Maggiore, 1844: synonym of Fossarus Philippi, 1841


  1. ^ Mahmoud, L.G. Etudes paleontologiques sur la fauna Cretacique du Massif du Moghara (Sinai-Egypte). Publications de I'Institut du Desert d'Egypte 8, 1956
  2. ^ a b Beechey, Des (2008). "Family Planaxidae". The Seashells of New South Wales. Retrieved 2014-12-21.
  3. ^ Houbrick R. S. (1987). "Anatomy, Reproductive Biology, and Phylogeny of the Planaxidae (Cerithiacea: Prosobranchia)". Smithsonian Contributions to Zoology 445.
  • Strong E.E., Colgan D.J., Healy J.M., Lydeard C., Ponder W.F. & Glaubrecht M. (2011) Phylogeny of the gastropod superfamily Cerithioidea using morphology and molecules. Zoological Journal of the Linnean Society 162: 43–89.
  • Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1–526.

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