Quercus palustris
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| Species: | Q. palustris
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Quercus palustris, the pin oak[3] or swamp Spanish oak, is an oak in the red oak section Quercus sect. Lobatae. Pin oak is one of the most commonly used landscaping oaks in its native range due to its ease of transplant, relatively fast growth, and pollution tolerance.[4] Its distinctive shape is considered unique among hardwoods.[4][5][6]
Distribution
Quercus palustris is mainly distributed in the eastern and central United States from Connecticut west to eastern Kansas, and south to Georgia, west to eastern Oklahoma and Kansas.[7] It is also native in the extreme south of Ontario, Canada. The pin oak is also well adapted to life in Australia (where it has been introduced) and is quite widespread across the Australian continent especially in the cooler southern States such as Victoria and New South Wales. Is also well adapted to life in Argentina, especially in the Río de la Plata region.
Pin oak grows primarily on level or nearly level, poorly drained alluvial floodplain and river bottom soils with high clay content. Pin oak is usually found on sites that flood intermittently during the dormant season but do not ordinarily flood during the growing season. It does not grow on the lowest, most poorly drained sites that may be covered with standing water through much of the growing season. However, it does grow extensively on poorly drained upland "pin oak flats" on the glacial till plains of southwestern Ohio, southern Illinois and Indiana, and northern Missouri. The level topography and presence of a claypan in the soil of these areas cause these sites to be excessively wet in winter and spring.[4]
Description
It is a medium-sized deciduous tree growing to 18–22 metres (59–72 ft) tall, with a trunk up to 1 metre (3.3 ft) diameter. It has an 8–14-metre (26–46 ft) spread. A 10-year-old tree will be about 8 metres (26 ft) tall. Young trees have a straight, columnar trunk with smooth bark and a pyramidal canopy. By the time the tree is 40 years old, it develops more rough bark with a loose, spreading canopy. This canopy is considered one of the most distinctive features of the pin oak: the upper branches point upwards, the middle branches are perpendicular to trunk, and the lower branches droop downwards.[4][8]
The leaves are 5–16 centimetres (2.0–6.3 in) long and 5–12 centimetres (2.0–4.7 in) broad, lobed, with five or seven lobes. Each lobe has 5-7 bristle-tipped teeth. The sinuses are typically u-shaped and extremely deep cut. In fact, there is approximately the same amount of sinus area as actual leaf area. The leaf is mostly hairless, except for a very characteristic tuft of pale orange-brown down on the lower surface where each lobe vein joins the central vein. Overall autumn leaf coloration is generally bronze, though individual leaves may be red for a time, and is not considered particularly distinctive.[5] The acorns, borne in a shallow, thin cap, are hemispherical, 10–16 millimetres (0.39–0.63 in) long and 9–15 millimetres (0.35–0.59 in) broad, green maturing pale brown about 18 months after pollination. The acorn is unpalatable because the kernel is very bitter.[8]
A fast-growing pioneer or riparian species, pin oak is relatively short-lived, with a maximum lifespan of 120 years against many oaks which can live several centuries. It is naturally a wetland tree, and develops a shallow, fibrous root system, unlike many oaks, which have a strong, deep taproot when young.[4] It is confined to acidic soils, and does not tolerate limestone or sandy Florida soil, and grows at low altitudes from sea level up to 350 m (1,148 ft). The Latin specific epithet palustris means "of swamps".[9][8]
A characteristic shared by a few other oak species, and also some beeches and hornbeams, is the retention of leaves through the winter on juvenile trees, a natural phenomenon referred to as marcescence. Young trees under 6 metres (20 ft) will often be covered with leaves year-round, though the leaves die in the fall, remaining attached to the shoots until the new leaves appear in the spring. As with many other oak species, dead pin oak branches will stay on the tree for many years.[4][8]
Flowering and fruiting
Like all oaks, flowering and leaf-out occur in late spring when all frost danger has passed. The flowers are monoecious catkins which, being self-incompatible, require the presence of another oak for pollination. Any species in the red oak group can serve as a pollinator, but in Pin Oak's natural range, this will usually be Northern Red Oak or Scarlet Oak. Inter-species hybridization occurs freely. The acorns take two growing seasons to develop.[8]
Associated forest cover
Pin oak is a major species in only one forest cover type, pin oak-sweetgum, which is found on bottom lands and some upland sites throughout the central portion of the pin oak range. Associated species in this type include red maple (Acer rubrum), American elm (Ulmus americana), black tupelo (Nyssa sylvatica), swamp white oak (Quercus bicolor), willow oak (Quercus phellos), overcup oak (Quercus lyrata), bur oak (Quercus macrocarpa), green ash (Fraxinus pennsylvanica), Nuttall oak (Quercus nuttallii), swamp chestnut oak (Quercus michauxii), and shellbark (Carya laciniosa) and shagbark (Carya ovata) hickories. Pin oak and sweetgum (Liquidambar styraciflua) vary in their relative proportions in this cover type. Large areas of almost pure pin oak occur on the "pin oak flats" of the upland glacial till plains or in the bottom lands of the lower Ohio and central Mississippi River valleys.[4]
Pin oak is an associated species in silver maple-American elm in the bottom lands along the Ohio, Wabash, Mississippi, and Missouri Rivers. A variant of this type, silver maple-American elm-pin oak-sweetgum, is found along major streams in southern Illinois and Indiana.[4]
Pin oak also occurs in black ash-American elm-red maple in poorly drained bottom lands in northern Ohio and Indiana along with silver maple (Acer saccharinum), swamp white oak, sycamore (Platanus occidentalis), black tupelo, and eastern cottonwood (Populus deltoides).[4]
Reaction to competition
Pin oak is classed as intolerant of shade. It is less tolerant than elm, boxelder (Acer negundo), sweetgum, hackberry (Celtis occidentalis), and ash, but is more tolerant than eastern cottonwood and black willow. Pin oak usually grows in even-aged stands of dominant and co-dominant trees. Intermediate and suppressed trees in such stands usually die within a few years of being overtopped. Single pin oaks in mixed stands usually are dominants. Pin oak is considered a subclimax species. It persists on heavy, wet soils because it produces an abundance of acorns which, if released, grow faster on these sites than most of its competitors.[4]
Pin oaks are also classified as "allelopathic," meaning they crowd out or otherwise successfully compete with understory vegetation to survive.
Damaging agents
Although pin oak is very tolerant of dormant-season flooding, it is much less tolerant of growing-season flooding. Trees may be injured or killed by intermittent growing season flooding over several successive years. The trees can usually survive one growing season of continuous flooding but will be killed by continuous flooding over 2 or 3 consecutive years. Pin oak is rated as "intermediately tolerant" to growing season flooding. Also, since the bark of pin oak is relatively thin, the species is especially susceptible to damage by fire and decay associated with fire wounds.[4]
Uses
In its native range, Pin Oak is the most commonly used landscaping oak along with Northern Red Oak due to its ease of transplant, relatively fast growth, and pollution tolerance. However, as it is naturally adapted to wet, acidic soils, it may develop a condition known as iron chlorosis on less suitable locations, causing the tree to shed leaves during the growing season and rot from the top down. Mature pin oaks are often too big to treat and this nutrient deficiency on alkaline soil may eventually kill them. The drooping lower branches can also be a problem, interfering with access for traffic and pedestrians. It is also cultivated in parks and large gardens in the United Kingdom, and has gained the Royal Horticultural Society's Award of Garden Merit.[10]
The wood is generally marketed as red oak, but is of significantly inferior quality, being somewhat weaker, often with many small knots.[4] The wood is hard and heavy and is used in general construction and for firewood.
The bark was used by some Native American tribes to make a drink for treatment of intestinal pain.
The name "pin oak" is possibly due to the many small, slender twigs, but may also be from the historical use of the hard wood for pins in wooden building construction.[11]
The pin oak is the only known foodplant of Bucculatrix domicola caterpillars.
Associated species
Due to similarity in leaf shape, the pin oak is often confused with scarlet oak and black oak, and occasionally, red oak. However, it can be distinguished by its distinctive dead branches on the lower trunk ("pins"), and its uniquely shaped crown. The sinuses on Pin Oak leaves are also deeply cut, often covering just as much area as the leaf itself.
References
- ^ Münchhausen, Otto von 1770. Der Hausvater 5(1): 253-254 diagnosis in Latin, description in German in Teutonic script
- ^ The Plant List, Quercus palustris Münchh.
- ^ "BSBI List 2007". Botanical Society of Britain and Ireland. Archived from the original (xls) on 2015-02-25. Retrieved 2014-10-17.
- ^ a b c d e f g h i j k l McQuilkin, Robert A. "Quercus palustris Muenchh". http://www.na.fs.fed.us/spfo/pubs/silvics_manual/volume_2/quercus/palustris.htm, accessed September 25, 2014.
- ^ a b "Oak, Pin Quercus palustris", http://www.arborday.org/treeguide/TreeDetail.cfm?id=19
- ^ "Pin Oak", http://www.treehelp.com/pin-oak/
- ^ Biota of North America Program 2014 county distribution map
- ^ a b c d e Flora of North America: Quercus palustris
- ^ Harrison, Lorraine (2012). RHS Latin for gardeners. United Kingdom: Mitchell Beazley. p. 224. ISBN 9781845337315.
- ^ "Quercus palustris". Royal Horticultural Society. Retrieved 23 July 2013.
- ^ Harlow, W. M. (1942). Trees of the Eastern and Central United States and Canada.
External Links
- Quercus palustris images from Vanderbilt University
- Pin Oak Cirrus Image, Diagnostic photographs and information.
- United States Department of Agriculture, National Forest Service, Silvics Manual
- photo of herbarium specimen at Missouri Botanical Garden, collected in Missouri in 1924