- EMS, UK, 1944-45
- Lecturer, Department of Physiology, Witwatersrand University, 1947-53
- Scientific Staff, National Institute for Medical Research, London, 1954-60
- Reader, Department of Immunology, St. Mary's Hospital Medical School, 1960-65
- Professor of Chemical Pathology, Guy's Hospital Medical School, 1965-86.
Other positions held
He was a member of the Medical Research Council (MRC) and chairman of its Tropical Medicine Research Board, 1974-76. He was the chairman of the World Health Organization Scientific Group on Immunity to Malaria, 1976-81, and a member of the WHO expert adv. panel on malaria, 1977-89. He helped to found the Royal College of Pathologists in 1964. He was elected a fellow of the Royal Society and awarded the CBE in 1978. He was on the Council of the Royal Society, 1981-83, and the Royal Society Assessor on the MRC, 1982-84.
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