Tambach Formation

Tambach Formation
Tambach Formation
Stratigraphic range: Early Permian, Artinskian PreꞒ Ꞓ O S D C P T J K Pg N
Type	Geological formation
Lithology
Primary	Conglomerate, sandstone, mudstone
Location
Region	Free State of Thuringia
Country	Germany
Type section
Named for	Tambach-Dietharz village

The Tambach Formation is an Early Permian-age geologic formation in central Germany. It consists of terrestrial lacustrine (lake) and fluvial (river) deposits. Many vertebrate fossils have been found from the formation, including those of early amphibians like Rotaryus and Tambaroter and early reptiles like Eudibamus and Thuringothyris.

Paleobiota

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Amphibians

Taxon	Member	Material	Notes
Georgenthalia^[1]	Tambach-Sandstein Member	MNG 11135 (holotype), a small but complete skull.	An amphibamid dissorophoid temnospondyl.
Orobates^[2]	Tambach-Sandstein Member	Four individuals are known from the specimens MNG 10181 (holotype), MNG 8760, 8980, 11133, 11134 (paratypes).	A diadectid reptiliomorph.
Rotaryus^[3]	Tambach-Sandstein Member	MNG 10182 (holotype), articulated partial well-preserved skull and both mandibles and a closely associated partial postcranial skeleton.	A trematopid dissorophoid temnospondyl.
Seymouria sanjuanensis^[4]	Tambach-Sandstein Member	MNG 7727, 8759, 10553, 10554 (referred)	A seymouriid reptiliomorph.
Silvadectes^[5]^[6]	Tambach-Sandstein Member	Four individuals are known from the specimens MNG 8853 (holotype), MNG 7721, 8747, 8978 (paratypes).	A diadectid reptiliomorph.
Tambachia^[7]	Tambach-Sandstein Member	MNG 7722 (holotype), a crushed skull and much of the postcranial skeleton.	A trematopid dissorophoid temnospondyl.
Tambaroter^[8]	Finsterbergen conglomerate Member	MNG 14708 (holotype), an almost complete skull.	An ostodolepid microsaur.

Reptiles

Taxon	Member	Material	Notes	Images
Eudibamus^[9]	Tambach-Sandstein Member	MNG 8852 (holotype), an almost complete cranial and postcranial skeleton.	A bolosaurid parareptile.
Thuringothyris^[10]	Tambach-Sandstein Member	At least eight individuals are known from the specimens MNG 7729 (holotype), MNG 10183, 10647, 10652, 11191 (referred).	The basalmost known captorhinid eureptile.^[11]

Synapsids

Taxon	Member	Material	Notes	Images
Dimetrodon teutonis^[12]	Tambach-Sandstein Member	MNG 10598 (holotype), partial vertebral column.^[12] The referred specimens MNG 10654, 10655, 10693 represent much of the postcranial skeleton. The referred specimen MNG 13433 represents a right maxilla.^[13]	A sphenacodontid.

References

^ Jason S. Anderson; Amy C. Henrici; Stuart S. Sumida; Thomas Martens; David S. Berman (2008). "Georgenthalia clavinasica, A New Genus and Species of Dissorophoid Temnospondyl from the Early Permian of Germany, and the Relationships of the Family Amphibamidae". Journal of Vertebrate Paleontology. 28 (1): 61–75. doi:10.1671/0272-4634(2008)28[61:GCANGA]2.0.CO;2. {{cite journal}}: Unknown parameter |last-author-amp= ignored (|name-list-style= suggested) (help)
^ Berman, D. Berman, D. S, Henrici, AC, Kissel, R., Sumida, SS, and Martens, T. S, Henrici, AC, Kissel, R., Sumida, SS, and Martens, T. (2004): A new diadectid (Diadectomorpha), Orobates pabsti, from the Early Permian of Central Germany. Bulletin of the Carnegie Museum of Natural History No 35: pp 1-37. abstract
^ "Rotaryus gothae, a New Trematopid (Temnospondyli: Dissorophoidea) from the Lower Permian of Central Germany". Annals of Carnegie Museum. 80 (1): 49–65. 2011. doi:10.2992/007.080.0106. {{cite journal}}: Unknown parameter |authors= ignored (help)
^ "Redescription of Seymouria sanjuanensis (Seymouriomorpha) from the Lower Permian of Germany based complete, mature specimens with a discussion of Paleoecology of Bromacker locality assemblage". Journal of Vertebrate Paleontology. 20 (2): 253–268. 2000. doi:10.1671/0272-4634(2000)020[0253:rosssf]2.0.co;2. {{cite journal}}: Unknown parameter |authors= ignored (help)
^ Berman, D.S.; Sumida, S.S.; Martens, T. (1998). "Diadectes (Diadectomorpha: Diadectidae) from the Early Permian of central Germany, with description of a new species". Annals of the Carnegie Museum. 67: 53–93.
^ Kissel, R. (2010). Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha). Toronto: University of Toronto Press. p. 185. hdl:1807/24357.
^ Sumida, S.S; Berman, D.S.; Martens, T. (1998). "A new trematopid amphibian from the Lower Permian of central Germany" (PDF). Palaeontology. 41 (4): 605–629. Archived from the original (PDF) on 2012-02-24. {{cite journal}}: Unknown parameter |deadurl= ignored (|url-status= suggested) (help)
^ Henrici, A.C.; Martens, T.; Berman, D.S.; Sumida, S.S. (2011). "An ostodolepid 'microsaur' (Lepospondyli) from the Lower Permian Tambach Formation of central Germany". Journal of Vertebrate Paleontology. 31 (5): 997–1004. doi:10.1080/02724634.2011.596601.
^ David S. Berman, Robert R. Reisz, Diane Scott, Amy C. Henrici, Stuart S. Sumida and Thomas Martens (2000). "Early Permian Bipedal Reptile". Science. 290 (5493): 969–972. Bibcode:2000Sci...290..969B. doi:10.1126/science.290.5493.969. PMID 11062126.{{cite journal}}: CS1 maint: multiple names: authors list (link)
^ "The basal reptile Thuringothyris mahlendorffae (Amniota: Eureptilia) from the Lower Permian of Germany". Journal of Paleontology. 80 (4): 726–739. 2006. doi:10.1666/0022-3360(2006)80[726:TBRTMA]2.0.CO;2. {{cite journal}}: Unknown parameter |authors= ignored (help)
^ Robert R. Reisz, Jun Liu, Jin-Ling Li and Johannes Müller (2011). "A new captorhinid reptile, Gansurhinus qingtoushanensis, gen. et sp. nov., from the Permian of China". Naturwissenschaften. 98 (5): 435–441. Bibcode:2011NW.....98..435R. doi:10.1007/s00114-011-0793-0. PMID 21484260.{{cite journal}}: CS1 maint: multiple names: authors list (link)
^ ^a ^b David S. Berman, Robert R. Reisz, Thomas Martens and Amy C. Henrici (2001). "A new species of Dimetrodon (Synapsida: Sphenacodontidae) from the Lower Permian of Germany records first occurrence of genus outside of North America" (PDF). Canadian Journal of Earth Sciences. 38 (5): 803–812. Bibcode:2001CaJES..38..803B. doi:10.1139/cjes-38-5-803.{{cite journal}}: CS1 maint: multiple names: authors list (link)
^ David S. Berman, Amy C. Henrici, Stuart S. Sumida and Thomas Martens (2004). "New materials of Dimetrodon teutonis (Synapsida: Sphenacodontidae) from the Early Permian of central Germany" (PDF). Annals of the Carnegie Museum of Natural History. 73 (2): 48–56.{{cite journal}}: CS1 maint: multiple names: authors list (link)

[Georgenthalia-1] Jason S. Anderson; Amy C. Henrici; Stuart S. Sumida; Thomas Martens; David S. Berman (2008). "Georgenthalia clavinasica, A New Genus and Species of Dissorophoid Temnospondyl from the Early Permian of Germany, and the Relationships of the Family Amphibamidae". Journal of Vertebrate Paleontology. 28 (1): 61–75. doi:10.1671/0272-4634(2008)28[61:GCANGA]2.0.CO;2. {{cite journal}}: Unknown parameter |last-author-amp= ignored (|name-list-style= suggested) (help)

[Orobates-2] Berman, D. Berman, D. S, Henrici, AC, Kissel, R., Sumida, SS, and Martens, T. S, Henrici, AC, Kissel, R., Sumida, SS, and Martens, T. (2004): A new diadectid (Diadectomorpha), Orobates pabsti, from the Early Permian of Central Germany. Bulletin of the Carnegie Museum of Natural History No 35: pp 1-37. abstract

[Rotaryus-3] "Rotaryus gothae, a New Trematopid (Temnospondyli: Dissorophoidea) from the Lower Permian of Central Germany". Annals of Carnegie Museum. 80 (1): 49–65. 2011. doi:10.2992/007.080.0106. {{cite journal}}: Unknown parameter |authors= ignored (help)

[S.sanjuanensisrev-4] "Redescription of Seymouria sanjuanensis (Seymouriomorpha) from the Lower Permian of Germany based complete, mature specimens with a discussion of Paleoecology of Bromacker locality assemblage". Journal of Vertebrate Paleontology. 20 (2): 253–268. 2000. doi:10.1671/0272-4634(2000)020[0253:rosssf]2.0.co;2. {{cite journal}}: Unknown parameter |authors= ignored (help)

[BST98-5] Berman, D.S.; Sumida, S.S.; Martens, T. (1998). "Diadectes (Diadectomorpha: Diadectidae) from the Early Permian of central Germany, with description of a new species". Annals of the Carnegie Museum. 67: 53–93.

[KR10-6] Kissel, R. (2010). Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha). Toronto: University of Toronto Press. p. 185. hdl:1807/24357.

[SBM98-7] Sumida, S.S; Berman, D.S.; Martens, T. (1998). "A new trematopid amphibian from the Lower Permian of central Germany" (PDF). Palaeontology. 41 (4): 605–629. Archived from the original (PDF) on 2012-02-24. {{cite journal}}: Unknown parameter |deadurl= ignored (|url-status= suggested) (help)

[Hetal11-8] Henrici, A.C.; Martens, T.; Berman, D.S.; Sumida, S.S. (2011). "An ostodolepid 'microsaur' (Lepospondyli) from the Lower Permian Tambach Formation of central Germany". Journal of Vertebrate Paleontology. 31 (5): 997–1004. doi:10.1080/02724634.2011.596601.

[Eudibamus-9] David S. Berman, Robert R. Reisz, Diane Scott, Amy C. Henrici, Stuart S. Sumida and Thomas Martens (2000). "Early Permian Bipedal Reptile". Science. 290 (5493): 969–972. Bibcode:2000Sci...290..969B. doi:10.1126/science.290.5493.969. PMID 11062126.{{cite journal}}: CS1 maint: multiple names: authors list (link)

[Thuringorev-10] "The basal reptile Thuringothyris mahlendorffae (Amniota: Eureptilia) from the Lower Permian of Germany". Journal of Paleontology. 80 (4): 726–739. 2006. doi:10.1666/0022-3360(2006)80[726:TBRTMA]2.0.CO;2. {{cite journal}}: Unknown parameter |authors= ignored (help)

[Gansurhinus-11] Robert R. Reisz, Jun Liu, Jin-Ling Li and Johannes Müller (2011). "A new captorhinid reptile, Gansurhinus qingtoushanensis, gen. et sp. nov., from the Permian of China". Naturwissenschaften. 98 (5): 435–441. Bibcode:2011NW.....98..435R. doi:10.1007/s00114-011-0793-0. PMID 21484260.{{cite journal}}: CS1 maint: multiple names: authors list (link)

[D.teutonis-12] David S. Berman, Robert R. Reisz, Thomas Martens and Amy C. Henrici (2001). "A new species of Dimetrodon (Synapsida: Sphenacodontidae) from the Lower Permian of Germany records first occurrence of genus outside of North America" (PDF). Canadian Journal of Earth Sciences. 38 (5): 803–812. Bibcode:2001CaJES..38..803B. doi:10.1139/cjes-38-5-803.{{cite journal}}: CS1 maint: multiple names: authors list (link)

[D.teutonis04-13] David S. Berman, Amy C. Henrici, Stuart S. Sumida and Thomas Martens (2004). "New materials of Dimetrodon teutonis (Synapsida: Sphenacodontidae) from the Early Permian of central Germany" (PDF). Annals of the Carnegie Museum of Natural History. 73 (2): 48–56.{{cite journal}}: CS1 maint: multiple names: authors list (link)

[1]

[2]

[3]

[4]

[5]

[6]

[7]

[8]

[9]

[10]

[11]

[12]

[13]