User:Snoteleks/cladograms/Cercozoa

Cladogram of Phylum Cercozoa, down to Order level^[1]

Cercozoa

	Chlorarachnida

	Minorisida

Granofilosea

	Cryptofilida

	Desmothoracida

	Leucodictyida

	Limnofilida

	Axomonadida

Skiomonadea

	Tremulida

	'Aquavolonida'

	Metopiida

	Metromonadida

Ventrifilosa

◄

Phylum Cercozoa Cavalier-Smith 1998 emend. 2018

Subphylum Filosa Cavalier-Smith & Chao 2003^[a]
- Infraphylum Reticulofilosa Cavaler-Smith 1997
  - Class Chlorarachnea Hibberd & Norris 1984 (as Chlorarachniophyceae)
    - Order Chlorarachnida Ishida et al. 1996 (as Chlorarachniales)
      - Family Chlorarachniidae Ishida et al. 1996 (Bigelowiella, Chlorarachnion, Norisiella, Partenskyella, ?Cryptochlora)
      - Family Gymnochloridae Cavalier-Smith 2018 (Gymnochlora, Amorphochlora, Viridiuvalis)
      - Family Lotharellidae Cavalier-Smith 2018 (Lotharella)
    - Order Minorisida Cavalier-Smith 2018
      - Family Minorisidae Cavalier-Smith 2018 (Minorisa)
  - Class Granofilosea Cavalier-Smith & Bass 2009
    - Order Cryptofilida Cavalier-Smith & Bass 2009
      - Family Nanofilidae Cavalier-Smith & Bass 2009 – Nanofila Cavalier-Smith & Bass 2009
      - Family Mesofilidae Cavalier-Smith & Bass 2009 – Mesofila Cavalier-Smith & Bass 2009
    - Order Desmothoracida Hertwig & Lesser 1874
      - Family Clathrulinidae Claus 1874
    - Order Leucodictyida Cavalier-Smith 1993
      - Family Leucodictyidae Cavalier-Smith 1993 – Leucodictyon, Reticulamoeba
      - Family Massisteriidae Cavalier-Smith 1993 – Massisteria, Minimassisteria
    - Order Limnofilida Cavalier-Smith & Bass 2009
      - Family Limnofilidae Cavalier-Smith & Bass 2009 (Limnofila)
    - Order Axomonadida Cavalier-Smith 2012
      - Family Tetradimorphidae Febvre-Chevalier & Febvre 1984 (Tetradimorpha)
  - Class Skiomonadea Cavalier-Smith 2012
    - Order Tremulida Cavalier-Smith & Howe 2011
      - Family Tremulidae Cavalier-Smith 2018 (Tremula)
    - Order Aquavolonida Bass et al. 2018 (Aquavolon)
- Infraphylum Monadofilosa Cavalier-Smith 1997
  - Superclass Eoglissa Cavalier-Smith 2011 emend. 2018
    - Class Metromonadea Cavalier-Smith 2007
      - Order Metopiida Cavalier-Smith 2003
        Family Metopiidae Cavalier-Smith 2003 (Metopion, Micrometopion)
      - Order Metromonadida Bass & Cavalier-Smith 2004
        Family Metromonadidae Bass & Cavalier-Smith 2004 (Metromonas)
        
        Family Kiitoksiidae Cavalier-Smith 2014
    - Class Helkesea Cavalier-Smith 2018
      - Order Helkesida Cavalier-Smith 2018
        Superfamily Sainouroidea Cavalier-Smith 2009 emend. 2018
        Family Sainouridae Cavalier-Smith 2008 (Sainouron, Cholamonas)
        
        Superfamily Helkesimastigoidea Cavalier-Smith 2018
        Family Helkesimastigidae Cavalier-Smith 2012 emend. 2018 (Helkesimastix)
        
        Family Guttulinopsidae Olive 1970 (Guttulinopsis, Rosculus)
  - Superclass Ventrifilosa Cavalier-Smith 2012 emend. 2018
    - Class Sarcomonadea Cavalier-Smith 1993 stat. nov. 1995 emend. 2018
      - Subclass Paracercomonada Cavalier-Smith 2018
        Order Paracercomonadida Cavalier-Smith 2018
        Family Paracercomonadidae Cavalier-Smith 2012 – Paracercomonas, Nucleocercomonas, Metabolomonas, Brevimastigomonas, Phytocercomonas
      - Subclass Pediglissa Cavalier-Smith 2018
        Order Cercomonadida Poche 1913 emend. Cavalier-Smith
        Family Cavernomonadidae Cavalier-Smith 2012 – Cavernomonas
        
        Family Cercomonadidae Saville Kent 1880-1881, emend. Cavalier-Smith
        
        Order Glissomonadida Howe et al. 2009
        Suborder Allapsina Cavalier-Smith 2018
        Family Allapsidae Howe et al. 2009 – Allapsa, Allantion, Teretomonas
        
        Suborder Sandonina Cavalier-Smith 2018
        Family Bodomorphidae Hollande 1952 – Bodomorpha
        
        Family Sandonidae Howe et al. 2009 – Neoheteromita, Sandona, Flectomonas, Mollimonas
        
        Family Proleptomonadidae Howe et al. 2009 – Proleptomonas
        
        Suborder Pansomonadina Vickerman 2005 stat. nov. Cavalier-Smith 2018
        Family Viridiraptoridae Hess & Melkonian 2013 – Viridiraptor, Orciraptor
        
        Family Agitatidae Cavalier-Smith & Bass 2009 – Agitata
        
        Family Acinetactidae Stokes 1886 – Acinectatis
        
        Family Aurigamonadidae Cavalier-Smith 2011 – Aurigamonas
      - Sarcomonadea incertae sedis
        Family Katabiidae Cavalier-Smith 2012 – Katabia^[3]
    - Class Imbricatea Cavalier-Smith 2003 emend. 2018
      - Subclass Placonuda Cavalier-Smith 2012
        Superorder Nudisarca Cavalier-Smith 2012
        Order Variglissida Cavalier-Smith 2014
        Family Clautriaviidae Cavalier-Smith 2013 (Clautriavia)
        
        Family Nudifilidae Cavalier-Smith & Howe 2011 (Nudifila)
        
        Family Quadriciliidae Cavalier-Smith 2018 (Quadricilia)
        
        Order Marimonadida Cavalier-Smith & Bass 2011
        Family Auranticordidae Cavalier-Smith 2018 (Auranticordis, Rhabdamoeba)
        
        Family Cyranomonadidae Cavalier-Smith 2018 (Cyranomonas)
        
        Family Pseudopirsoniidae Cavalier-Smith 2018 (Pseudopirsonia)
        
        Family Abolliferidae Cavalier-Smith 2018 (Abollifer, Cowlomonas, ?Heterochromonas)
        
        Superorder Euglyphia Cavalier-Smith 2018
        Order Euglyphida Copeland 1956 emend. Cavalier-Smith 1987
        Family Euglyphidae Wallich 1864 (Euglypha)
        
        Family Trinematidae Hoogenraad & De Groot 1940 (Corythion, Trinema)
        
        Family Sphenoderiidae Chatelain et al. 2013 (Sphenoderia, Trachelocorythion)
        
        Family Assulinidae Lara et al. 2006 (Assulina, Placocista)
        
        Family Cyphoderiidae De Saedeleer 1934 (Cyphoderia, Corythionella, Pseudocorythion)
        
        Family Paulinellidae De Saedeleer 1934 (Paulinella, Ovulinata, Micropyxidiella)
        
        Order Zoelucasida Cavalier-Smith 2014
        Family Zoelucasidae Cavalier-Smith 2014 (Zoelucasa)
        
        Superorder Discomonada Cavalier-Smith 2018
        Order Discomonadida Cavalier-Smith 2014
        Family Discomonadidae Cavalier-Smith 2014 (Discomonas)
      - Subclass Placoperla Cavalier-Smith 2012
        Superorder Placofila Cavalier-Smith 2012
        Order Thaumatomonadida Shirkina 1987
        Family Thaumatomonadidae Hollande 1952 (Allas, Ovaloplaca, Reckertia, Scutellomonas, Thaumatomonas, Thaumatospina, Penardeugenia)
        
        Family Peregriniidae Cavalier-Smith 2011 (Peregrinia, Gyromitus)
        
        Family Esquamulidae Shiratori et al. 2012 (Esquamula)
        
        Order Discocelida Cavalier-Smith 1997
        Family Discoceliidae Cavalier-Smith 1993 orthog. emend. 2012 (Discocelia)
        
        Superorder Perlatia Cavalier-Smith 2012
        Order Spongomonadida Hibberd 1983
        Family Spongomonadidae Karpov 1990 (Spongomonas, Rhipidodendron)
        
        Order Perlofilida Cavalier-Smith 2012
        Family Pompholyxophryidae Page 1987 (Pompholyxophrys)
        
        Family Acanthoperlidae Cavalier-Smith 2012 (Acanthoperla)
      - Subclass Krakenia Cavalier-Smith 2018
        Order Krakenida Dumack et al. 2017 ex Cavalier-Smith 2018
        Family Krakenidae Dumack et al. 2017 (Kraken)
    - Class Thecofilosea Cavalier-Smith 2003 emend. 2012
      - Subclass Ventricleftia Cavalier-Smith 2018
        Order Ventricleftida Cavalier-Smith 2011
        Family Ventrifissuridae Cavalier-Smith 2018 (Ventrifissura)
        
        Family Verrucomonadidae Cavalier-Smith 2018 (Verrucomonas)
      - Subclass Eothecia Cavalier-Smith 2012
        Order Matazida Cavalier-Smith 2012
        Family Matazidae Cavalier-Smith 2012
        
        Order Ebriida Deflandre 1936 (Ebria, Hermesinum, Botuliforma)
        
        Order Cryomonadida Cavalier-Smith 1993
        Family Cryomonadidae Cavalier-Smith 1993 (Cryothecomonas)
        
        Family Protaspidae Cavalier-Smith 1993 (Protaspa)
        
        Family Rhogostomidae Dumack et al. 2017 (Rhogostoma, Capsellina, Sacciforma)
      - Subclass Phaeodaria Haeckel 1879 (=Tripylea Hertwig 1879)
        Order Eodarida Cavalier-Smith 2012
        Suborder Phaeogymnocellina (=Phaeogymnocellida) Cachon & Cachon 1985
        Family Phaeosphaeridae Cachon-Enjumet 1961 – Phaeopyla, Phaeodactylis, Phaeosphaera
        
        Family Phaeodinidae Cachon-Enjumet 1961 – Phaeodina
        
        Family Atlanticellidae Cachon-Enjumet 1961 – Gymnocelia, Halocelia, Lobocelia, Miracelia, Planktonetta
        
        Suborder Phaeocystina (=Phaeocystida) Haeckel 1887
        Family Aulacanthidae Haeckel 1887 – Aulacantha
        
        Family Astracanthidae Haeckel 1887 – Astracantha, Castanella, Castanissa
        
        Order Opaloconchida Cavalier-Smith 2012
        Suborder Phaeosphaeria (=Phaeocystida) Haeckel 1887
        Family Aulosphaeridae Haeckel 1887 – Aulosphaera, Aularia, Aulotractus
        
        Family Cannosphaeridae Haeckel 1887 – Coelocantha
        
        Family Sagosphaeridae Haeckel 1887 – Sagenoarium, Sagenoscena, Sagoscena
        
        Suborder Phaeocalpia (=Phaeocalpida) Haeckel 1887
        Family Castanellidae Haecker 1906 – Castanea
        
        Family Circoporidae Haeckel 1887 – Circoporus, Circospathis, Haeckeliana
        
        Family Tuscaroridae Haeckel 1887 – Tuscarora, Tuscarilla, Tuscaretta
        
        Family Porospathidae Borgert 1900 – Porospathis
        
        Family Polypyramidae Reschetnjak 1966 – Polypyramis
        
        Suborder Phaeogromia (=Phaeogromida) Haeckel 1887
        Family Challengeridae Murray 1886 – Challengeria, Challengeron
        
        Family Medusettiidae Haeckel 1887 – Euphysetta, Gazelletta, Medusetta
        
        Family Lirellidae Loeblich & Tappan 1961 – Borgertella, Lirella
        
        Suborder Phaeoconchia (=Phaeoconchida) Haeckel 1887
        Family Concharidae Haeckel 1887 – Conchidium
        
        Suborder Phaeodendria (=Phaeodendrida) Haeckel 1887
        Family Coelodendridae Haeckel 1887 – Coelodendrum, Coelographis
      - Subclass Tectosia Cavalier-Smith 2012
        Order Tectofilosida Cavalier-Smith 2003
        Family Chlamydophryidae De Saedeleer 1934
        
        Family Psammonobiotidae Golemansky 1974
        
        Family Volutellidae Sudzuki 1979 (Volutella)
        
        Suborder Fiscullina Cavalier-Smith 2018
        Family Fiscullidae Dumack, Mausback & Bonkowski 2017 (Fisculla)
        
        Family Pseudodifflugiidae De Saedeleer 1934
        
        Family Rhizaspididae Skuja 1948 (Rhizaspis)
Subphylum Endomyxa Cavalier-Smith 2002^[a]
- Superclass Marimyxia Cavalier-Smith 2018
  - Class Gromiidea Cavalier-Smith 2003 emend. 2018^[b]^[1]
    - Order Gromiida Claparède & Lachmann 1856
      - Family Gromiidae Reuss 1862 – Gromia
    - Order Reticulosida Cavalier-Smith 2003 emend. 2018^[1]
      - Family Filoretidae Cavalier-Smith & Bass 2009^[4] – Filoreta
  - Class Ascetosporea Sprague 1979 stat. nov. Cavalier-Smith 2002
    - Order Claustrosporida Cavalier-Smith 2003
      - Family Claustrosporidiidae Larsson 1987 – Claustrosporidium
    - Order Haplosporida Caullery & Mesnil 1889 orthog. emend. Lühe 1900 [=Balanosporida Sprague 1979]
      - Family Haplosporidiidae Sprague 1982 [=Haplosporidae] – Bonamia, Haplosporidium, Minchinia
      - Family Urosporidiidae Sprague 1979 [=Anurosporidiidae] – Urosporidium (=Anurosporidium)
    - Order Mikrocytida Hartikainen et al. 2014
      - Family Mikrocytidae Cavalier-Smith 2018^[1] – Mikrocytos, Paramikrocytos
    - Order Paradinida Cavalier-Smith 2009^[4]
      - Family Paradiniidae Schiller 1935 – Paradinium
    - Order Paramyxida Chatton 1911
      - Family Marteiliidae Sprague 1979 – Marteilia, Paramarteilia
      - Family Paramyxidae Chatton 1911 – Paramyxa
- Superclass Proteomyxia Lankester 1885 ex Cavalier-Smith 2018^[1]
  - Class Vampyrellidea Cavalier-Smith 2018^[1]
    - Order Vampyrellida West 1901) ex Cavalier-Smith 2018^[1] [=Aconchulinida De Saedeleer 1934]
      - Family Vampyrellidae Zopf 1885 – Vampyrella
      - Family Leptophryidae Hess et al. 2012 – Leptophrys, Platyreta, Theratromyxa, Vernalophrys
      - Family Arachnulidae Page 1967 – Arachnula
      - Family Hyalodiscidae Poche 1913 – Hyalodiscus
      - Vampyrellida genera incertae sedis – Thalassomyxa
  - Class Phytomyxea Engler & Prantl 1897
    - Order Phagomyxida Cavalier-Smith 1993 – Phagomyxa, Maullinia
      - Family Phagomyxidae Cavalier-Smith 1993 – Phagomyxa
    - Order Plasmodiophorida Cook 1928 [=Plasmodiophorales Engl. 1892;^[5] phylum Plasmodiophoromycota R.H. Whittaker 1969^[6]]
      - Family Plasmodiophoridae Zopf 1884 [=Plasmodiophoraceae Zopf ex Berl. 1888] – Plasmodiophora (=Ostenfeldiella), Spongospora, Tetramyxa (=Molliardia), Hillenburgia^[7]
    - Phytomyxea incertae sedis – Marinomyxa^[8]

Notes

^ ^a ^b The taxonomy of Cercozoa was revised in 2018 to exclude Endomyxa to Retaria, making Cercozoa synonymous with the subphylum Filosa.^[1] However, this page maintains the subphylum Endomyxa inside of Cercozoa in accordance to the latest phylogenetic results.^[2]
^ Probably paraphyletic.

References

^ ^a ^b ^c ^d ^e ^f ^g ^h Cite error: The named reference RhizariaPhylo2018 was invoked but never defined (see the help page).
^ Cite error: The named reference MonoCercozoa was invoked but never defined (see the help page).
^ Karpov SA, Ekelund F, Moestrup Ø (2003), "Katabia gromovi nov. gen., nov. sp. a new soil flagellate with affinities to Heteromita (Cercomonadida)", Protistology, 3: 30–41
^ ^a ^b Cite error: The named reference NovelNakedFilose was invoked but never defined (see the help page).
^ Cite error: The named reference Engler1903 was invoked but never defined (see the help page).
^ Cite error: The named reference Whittaker69 was invoked but never defined (see the help page).
^ Cite error: The named reference Hillenburgia was invoked but never defined (see the help page).
^ Cite error: The named reference Marinomyxa was invoked but never defined (see the help page).

[rhizaria-3] The taxonomy of Cercozoa was revised in 2018 to exclude Endomyxa to Retaria, making Cercozoa synonymous with the subphylum Filosa.^[1] However, this page maintains the subphylum Endomyxa inside of Cercozoa in accordance to the latest phylogenetic results.^[2]

[5] Probably paraphyletic.

[RhizariaPhylo2018-1] ^ ^a ^b ^c ^d ^e ^f ^g ^h Cite error: The named reference RhizariaPhylo2018 was invoked but never defined (see the help page).

[MonoCercozoa-2] Cite error: The named reference MonoCercozoa was invoked but never defined (see the help page).

[Katabia-4] Karpov SA, Ekelund F, Moestrup Ø (2003), "Katabia gromovi nov. gen., nov. sp. a new soil flagellate with affinities to Heteromita (Cercomonadida)", Protistology, 3: 30–41

[NovelNakedFilose-6] Cite error: The named reference NovelNakedFilose was invoked but never defined (see the help page).

[Engler1903-7] Cite error: The named reference Engler1903 was invoked but never defined (see the help page).

[Whittaker69-8] Cite error: The named reference Whittaker69 was invoked but never defined (see the help page).

[Hillenburgia-9] Cite error: The named reference Hillenburgia was invoked but never defined (see the help page).

[Marinomyxa-10] Cite error: The named reference Marinomyxa was invoked but never defined (see the help page).

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