Sexual reproduction

Sexual reproduction is a type of reproduction that results in increasing genetic diversity of the offspring. It is characterized by two processes. The first, meiosis, involves the halving of the number of chromosomes. The second process, fertilization, leads the fusion of two gametes and the restoration of the original number of chromosomes. During meiosis, the chromosomes of each pair usually cross over to achieve genetic recombination.

The evolution of sex is a major puzzle in modern evolutionary biology. The first fossilized evidence of sexually reproducing organisms is from eukaryotes of the Stenian period, about 1.2 to 1 billion years before the present time. Sexual reproduction is the primary method of reproduction for the vast majority of visible organisms, including almost all animals and plants. Bacterial conjugation, the transfer of DNA between two bacteria, is often mistakenly confused with sexual reproduction, because the mechanics are similar.

Contemporary evolutionary thought posits a few explanations for why sexual reproduction persists when parthenogenesis appears to be a superior form of reproduction. Sexual reproduction may persist because of selection pressure on the clade itself- the ability for a population to radiate more rapidly due to a changing environment through sexual recombination than parthenogenesis allows. Alternatively, sexual reproduction may allow for the 'ratcheting' of evolutionary speed as one clade competes with another for a limited resource.

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In the first stage of sexual reproduction, 'meiosis', the number of chromosomes is reduced from a diploid number (2n) to a haploid number (n). During 'fertilization', haploid gametes come together to form a diploid zygote and the original number of chromosomes (2n) is restored.

Sexual reproduction of protists and fungi

Many protists and fungi reproduce sexually. Although they are unicellular, at times of reproduction the "father" cell and the "mother" cell combines together. Next, their genetic information combines together into a new formation, and by cell division the offspring is born.

Reproduction in plants

In flowering plants, a stamen produces gametes called pollen grains, which attach to a pistil, in which the female gametes (ovules) are located. Here, the female gamete is fertilized and develops into a seed. The ovary, which produced the gamete then grows into a fruit, which surrounds the seed(s). Plants may either self-pollinate or cross-pollinate.

Reproduction in archosaurs (reptiles and birds)

Male and female birds and reptiles both have cloacae. The female lays amniotic eggs, fertilized by the male, in which the young gestate.

Reproduction in mammals

In placental mammals, offspring are born as juveniles: complete animals with the sex organs present although non-functional. After several months or years, the sex organs develop further to maturity and the animal becomes sexually mature. Most female mammals are only fertile during certain periods and during those times, they are said to be "in heat". At this point, the animal is ready to mate. Individual male and female mammals meet and carry out copulation. For most mammals, males and females exchange sexual partners throughout their adult lives.

The mammalian male

The male reproductive system contains two main divisions: the penis, which is inserted into the female and carries the sperm inside it, and the testes, which produce the sperm. In humans, both of these organs are outside the abdominal cavity, but they can be primarily housed within the abdomen in other animals (for instance, in dogs, the penis is internal except when mating). Having the testes outside the abdomen best facilitates temperature regulation of the sperm, which require specific temperatures to survive.

Sperm are the smaller of the two gametes and are generally very short-lived, requiring males to produce them continuously from the time of sexual maturity until death. They are motile and swim by chemotaxis.

The mammalian female

The female reproductive system likewise contains two main divisions: the vagina and uterus, which act as the receptacle for the male's sperm, and the ovaries, which produce the female's ova. All of these parts are always internal. The vagina is attached to the uterus through the cervix, while the uterus is attached to the ovaries via the Fallopian tubes. At certain intervals, the ovaries release an ovum (the singular of ova), which passes through the fallopian tube into the uterus.

If, in this transit, it meets with sperm, the sperm penetrate and merge with the egg, fertilizing it. The fertilization usually occurs in the oviducts, but can happen in the uterus itself. The zygote then implants itself in the wall of the uterus, where it begins the processes of embryogenesis and morphogenesis. When developed enough to survive outside the womb, the cervix dilates and contractions of the uterus propel the fetus through the birth canal, which is the vagina.

The ova are larger than sperm and are generally all created by birth. They are for the most part stationary, aside from their transit to the uterus, and contain nutrients for the later zygote and embryo. Over a regular interval, a process of oogenesis matures one ovum to be sent down the Fallopian tube attached to its ovary in anticipation of fertilization. If not fertilized, this egg is flushed out of the system through menstruation in humans and great apes and reabsorbed in all other mammals in the estrus cycle.

Gestation

Gestation, called pregnancy in humans, is the period of time during which the fetus develops, dividing via mitosis inside the female. During this time, the fetus receives all of its nutrition and oxygenated blood from the female, filtered through the placenta, which is attached to the fetus' abdomen via an umbilical cord. This drain of nutrients can be quite taxing on the female, who is required to ingest significantly higher levels of calories. In addition, certain vitamins and other nutrients are required in greater quantities than normal, often creating abnormal eating habits. The length of gestation, called the gestation period, varies greatly from species to species; it is 38 weeks in humans, 56-60 in giraffes and 16 days in hamsters.

Birth

Once the fetus is sufficiently developed, chemical signals start the process of birth, which begins with contractions of the uterus and the dilation of the cervix. The fetus then descends to the cervix, where it is pushed out into the vagina, and eventually out of the female. The newborn, which is called an infant in humans, should typically begin respiration on its own shortly after birth. Not long after, the placenta is passed as well. Most mammals eat this, as it is a good source of protein and other vital nutrients needed for caring for the young. The end of the umbilical cord attached to the young’s abdomen eventually falls off on its own.

Monotremes

Monotremes, only five species of which exist, all from Australia and New Guinea, lay eggs. They have one opening for excretion and reproduction called the cloaca. They hold the eggs internally for several weeks, providing nutrients, and then lay them and cover them like birds. After less than two weeks the young hatches and crawls into its mother’s pouch, much like marsupials, where it nurses for several weeks as it grows.

Marsupials

Marsupials reproduce in essentially the same manner, though their young are born at a far earlier stage of development than other mammals. After birth, marsupial joeys crawl into their mother’s pouch and attach to a teat, where they receive nourishment and finish developing into self-sufficient animals.

References

Pang, K. "Certificate Biology: New Mastering Basic Concepts", Hong Kong, 2004
Journal of Biology of Reproduction, accessed in August 2005.