Population history of Egypt: Difference between revisions

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There is also evidence that sheep and goats were introduced into [[Nabta]] from [[Southwest Asia]] about 8,000 years ago.<ref name="comp-archaeology.org"/> There is some speculation that this culture is likely to be the predecessor of the Egyptians, based on cultural similarities and social complexity which is thought to be reflective of Egypt's [[Old Kingdom]].<ref>[http://hej3.as.utexas.edu/~www/wheel/africa/nabta_01.htm Ancient Astronomy in Africa]</ref><ref>{{cite book |title=Holocene Settlement of the Egyptian Sahara | first=Fred |last=Wendorf| isbn=0-306-46612-0|publisher=Springer|year=2001|pages=525|url=http://books.google.com/?id=qUk0GyDJRCoC&pg=PA525&dq=nabta+playa+sub-saharan }}</ref>
There is also evidence that sheep and goats were introduced into [[Nabta]] from [[Southwest Asia]] about 8,000 years ago.<ref name="comp-archaeology.org"/> There is some speculation that this culture is likely to be the predecessor of the Egyptians, based on cultural similarities and social complexity which is thought to be reflective of Egypt's [[Old Kingdom]].<ref>[http://hej3.as.utexas.edu/~www/wheel/africa/nabta_01.htm Ancient Astronomy in Africa]</ref><ref>{{cite book |title=Holocene Settlement of the Egyptian Sahara | first=Fred |last=Wendorf| isbn=0-306-46612-0|publisher=Springer|year=2001|pages=525|url=http://books.google.com/?id=qUk0GyDJRCoC&pg=PA525&dq=nabta+playa+sub-saharan }}</ref>


{{main|DNA history of Ancient Egypt}}
==DNA studies==
Contamination from handling and intrusion from microbes create obstacles to the recovery of [[Ancient DNA]].<ref name="books.google.com"/> Consequently most DNA studies have been carried out on modern Egyptian populations with the intent of learning about the influences of historical migrations on the population of Egypt.<ref name="S.O.Y. Keita & A. J. Boyce">{{cite journal|title=Genetics, Egypt, and History: Interpreting Geographical Patterns of Y Chromosome Variation|year=2005|last= [[S.O.Y. Keita]] & A. J. Boyce |doi=10.1353/hia.2005.0013|url= http://wysinger.homestead.com/keita.pdf|first1=S. O. Y.|last2=Boyce|first2=A. J. (Anthony J.)|journal=History in Africa|volume=32|issue=1|pages=221|date=June 2009}}</ref><ref name="Shomarka Keita (2005)">{{cite journal|title=Y-Chromosome Variation in Egypt|last=Shomarka Keita (2005)|url=http://wysinger.homestead.com/African_Archaeological_Revie__June_2005_.pdf|isbn=43700541894 {{Please check ISBN|reason=Invalid length.}}|doi=10.1007/s10437-005-4189-4|year=2005|first1=S. O. Y.|journal=African Archaeological Review|volume=22|issue=2|pages=61}}</ref><ref name="Keita">{{cite journal|title=History in the Interpretation of the Pattern of p49a,f TaqI RFLP Y-Chromosome Variation in Egypt|year=2005|last=Keita|doi=10.1002/ajhb.20428|url=http://wysinger.homestead.com/keita6.pdf|first1=S.O.Y.|journal=American Journal of Human Biology|volume=17|pages=559–67|pmid=16136533|issue=5}}</ref><ref>[http://ngm.nationalgeographic.com/video/player?titleID=1414281487#/?titleID=1414242821&catID=1 Shomarka Keita: What genetics can tell us]</ref>
{{Further|Archaeogenetics of the Near East}}
Attempts to extract ancient DNA or [[aDNA]] from Ancient Egyptian remains have yielded mainly [[Eurasia]]n DNA types from the [[Dakleh Oasis]] cemetery site (from Southern Egypt), and they show a considerable increase in the amount of [[Sub Saharan]] mitchondrial DNA only over the past 2,000 years, suggesting that within this timeframe there was more migration from [[Sub-Saharan Africa]] to the Nile Valley than from [[Eurasia]] to the Nile Valley.<ref name="books.google.com">[http://books.google.com/books?id=XNdgScxtirYC&printsec=frontcover#PPA278,M1 Encyclopedia of the Archaeology of Ancient Egypt By Kathryn A. Bard, Steven Blake Shubert pp 278-279]</ref> One successful study was performed on ancient mummies of the 12th Dynasty, by Paabo and Di Rienzo, which identified multiple lines of descent, which originated in sub-Saharan Africa.<ref>Paabo, S., and A. Di Rienzo, A molecular approach to the study of Egyptian history. In Biological Anthropology and the Study of Ancient Egypt. V. Davies and R. Walker, eds. pp. 86-90. London: British Museum Press. 1993</ref> Contamination from handling and intrusion from microbes have also created obstacles to recovery of [[Ancient DNA]].<ref name="books.google.com"/> Consequently most DNA studies have been carried out on modern Egyptian populations with the intent of learning about the influences of historical migrations on the population of Egypt.<ref name="S.O.Y. Keita & A. J. Boyce">{{cite journal|title=Genetics, Egypt, and History: Interpreting Geographical Patterns of Y Chromosome Variation|year=2005|last= [[S.O.Y. Keita]] & A. J. Boyce |doi=10.1353/hia.2005.0013|url= http://wysinger.homestead.com/keita.pdf|first1=S. O. Y.|last2=Boyce|first2=A. J. (Anthony J.)|journal=History in Africa|volume=32|issue=1|pages=221|date=June 2009}}</ref><ref name="Shomarka Keita (2005)">{{cite journal|title=Y-Chromosome Variation in Egypt|last=Shomarka Keita (2005)|url=http://wysinger.homestead.com/African_Archaeological_Revie__June_2005_.pdf|isbn=43700541894 {{Please check ISBN|reason=Invalid length.}}|doi=10.1007/s10437-005-4189-4|year=2005|first1=S. O. Y.|journal=African Archaeological Review|volume=22|issue=2|pages=61}}</ref><ref name="Keita">{{cite journal|title=History in the Interpretation of the Pattern of p49a,f TaqI RFLP Y-Chromosome Variation in Egypt|year=2005|last=Keita|doi=10.1002/ajhb.20428|url=http://wysinger.homestead.com/keita6.pdf|first1=S.O.Y.|journal=American Journal of Human Biology|volume=17|pages=559–67|pmid=16136533|issue=5}}</ref><ref>[http://ngm.nationalgeographic.com/video/player?titleID=1414281487#/?titleID=1414242821&catID=1 Shomarka Keita: What genetics can tell us]</ref>


In general, various DNA studies have found that the gene frequencies of [[North African]] populations are intermediate between those of the [[Near East]], the [[Horn of Africa]], southern [[Europe]] and [[Sub Saharan Africa]],<ref>[http://books.google.com/books?id=FrwNcwKaUKoC&printsec=frontcover#PPA136,M1 Cavalli-Sforza, History and Geography of Human Genes], The intermediacy of North Africa and to a lesser extent [[Europe]] is apparent</ref> though Egypt’s NRY frequency distributions appear to be much more similar to those of the [[Middle East]] than to any sub-Saharan African population, suggesting a much larger [[Eurasia]]n genetic component.<ref name="luis">The Levant versus the Horn of Africa: Evidence for Bidirectional Corridors of Human Migrations – Luis; Rowold; Regueiro; Caeiro; Cinnioğlu; Roseman; Underhill; Cavalli-Sforza; and Herrera. – see http://www.pubmedcentral.nih.gov/articlerender.fcgi?artid=1182266</ref><ref name="Cavalli-Sforza">Cavalli-Sforza, L.L., P. Menozzi, and A. Piazza. 1994. The History and Geography of Human Genes. Princeton: Princeton University Press.</ref><ref name="Cavalli-Sforza" /><ref name="Bosch1997">{{cite journal | last1 = Bosch | first1 = E. ''et al.'' | year = 1997 | title = Population history of north Africa: evidence from classical genetic markers | url = | journal = Human Biology | volume = 69 | issue = 3| pages = 295–311 }}</ref><ref>{{cite journal | author=Arredi B, Poloni E, Paracchini S, Zerjal T, Fathallah D, Makrelouf M, Pascali V, Novelletto A, Tyler-Smith C | title=A predominantly Nilo Saharan origin for Y-chromosomal DNA variation in North Africa. | journal=Am J Hum Genet | volume=75 | issue=2 | pages=338–45 | year=2004 | pmid=15202071 | doi = 10.1086/423147 | pmc=1216069}}</ref><ref>{{cite journal | author=Manni F, Leonardi P, Barakat A, Rouba H, Heyer E, Klintschar M, McElreavey K, Quintana-Murci L | title=Y-chromosome analysis in Egypt suggests a genetic regional continuity in Northeastern Africa. | journal=[[Hum Biol]]| volume=74 | issue=5 | pages=645–58 | year=2002 | pmid=12495079 | doi = 10.1353/hub.2002.0054}}</ref><ref>{{cite book|title= The History and Geography of Human Genes|authorlink=Cavalli-Sforza|last=Cavalli-Sforza|chapter=Synthetic maps of Africa|chapterurl=http://books.google.com/books?id=FrwNcwKaUKoC&printsec=frontcover#PPA189,M1|isbn= 0691087504}}The present population of the Sahara is Sudan in the extreme north, with an increase of Negroid component as one goes south</ref>
===DNA studies on modern Egyptians===
Egypt has experienced several invasions during its history. However, these do not seem to account for more than about 10% overall of current Egyptians ancestry when the DNA evidence of the ancient mitochondrial DNA and modern Y chromosomes is considered. While [[Ivan van Sertima]] argue that the Egyptians were primarily [[Africoid]] before the many conquests of Egypt diluted the Africanity of the Egyptian people,<ref name="Ivan van Sertima 1994">{{cite book|title=Egypt, Child of Africa|authorlink=Ivan van Sertima|chapter=|chapterurl=http://books.google.com/books?id=Y7KmBTz2vUoC&printsec=frontcover#PPA2-IA1,M1|year=1994|isbn=1-56000-792-3}}</ref> other scholars such as [[Frank Yurco]] believe that Modern Egyptians are largely representative of the ancient population, and the DNA evidence appears to support this view.<ref name="Frank Yurco 1996. p. 62-100">Frank Yurco, "An Egyptological Review" in Mary R. Lefkowitz and Guy MacLean Rogers, eds. ''Black Athena Revisited''. Chapel Hill: University of North Carolina Press, 1996. p. 62-100</ref>


Blood typing and DNA sampling on ancient Egyptian mummies is scant; however, blood typing of dynastic mummies found ABO frequencies to be most similar to modern Egyptians<ref>Borgognini Tarli S.M., Paoli G. 1982. Survey on paleoserological studies. Homo, 33: 69-89</ref> and some also to Northern [[Haratin]] populations. ABO blood group distribution shows that the Egyptians form a sister group to North African populations, including [[Berber people|Berbers]], [[Nubians]] and [[Canary Island]]ers.<ref>Cavalli-Sforza, L.L., P. Menozzi, and A. Piazza. 1994. ''The History and Geography of Human Genes''. Princeton: Princeton University Press, 169-174</ref> Scholars such as [[Frank Yurco]] believe that Modern Egyptians are largely representative of the ancient population, and the DNA evidence appears to support this view.<ref name="Frank Yurco 1996. p. 62-100">Frank Yurco, "An Egyptological Review" in Mary R. Lefkowitz and Guy MacLean Rogers, eds. ''Black Athena Revisited''. Chapel Hill: University of North Carolina Press, 1996. p. 62-100</ref>
In general, various DNA studies have found that the gene frequencies of modern North African populations are intermediate between those of the [[Horn of Africa]] and [[Eurasia]],<ref>[http://books.google.com/books?id=FrwNcwKaUKoC&printsec=frontcover#PPA136,M1 Cavalli-Sforza, History and Geography of Human Genes], The intermediacy of North Africa and to lesser extent East Africa between [[Africa]] and [[Europe]] is apparent</ref> though possessing a greater genetic affinity with the populations of Eurasia than they do with Africa.<ref name="Cavalli-Sforza">Cavalli-Sforza, L.L., P. Menozzi, and A. Piazza. 1994. The History and Geography of Human Genes. Princeton: Princeton University Press.</ref><ref name="Cavalli-Sforza"/><ref name="Bosch1997">{{cite journal | last1 = Bosch | first1 = E. ''et al.'' | year = 1997 | title = Population history of north Africa: evidence from classical genetic markers | url = | journal = Human Biology | volume = 69 | issue = 3| pages = 295–311 }}</ref><ref>{{cite journal | author=Arredi B, Poloni E, Paracchini S, Zerjal T, Fathallah D, Makrelouf M, Pascali V, Novelletto A, Tyler-Smith C | title=A predominantly neolithic origin for Y-chromosomal DNA variation in North Africa. | journal=Am J Hum Genet | volume=75 | issue=2 | pages=338–45 | year=2004 | pmid=15202071 | doi = 10.1086/423147 | pmc=1216069}}</ref><ref>{{cite journal | author=Manni F, Leonardi P, Barakat A, Rouba H, Heyer E, Klintschar M, McElreavey K, Quintana-Murci L | title=Y-chromosome analysis in Egypt suggests a genetic regional continuity in Northeastern Africa. | journal=[[Hum Biol]]| volume=74 | issue=5 | pages=645–58 | year=2002 | pmid=12495079 | doi = 10.1353/hub.2002.0054}}</ref><ref>{{cite book|title= The History and Geography of Human Genes|authorlink=Cavalli-Sforza|last=Cavalli-Sforza|chapter=Synthetic maps of Africa|chapterurl=http://books.google.com/books?id=FrwNcwKaUKoC&printsec=frontcover#PPA189,M1|isbn= 0-691-08750-4}}The present population of the Sahara is Caucasoid in the extreme north, with a fairly gradual increase of Negroid component as one goes south</ref>

A study by Krings et al. (1999) on [[mitochondrial DNA]] [[Cline (biology)|clines]] along the Nile Valley found that a [[Eurasia]]n cline runs from [[Northern Egypt]] to [[Southern Sudan]] and a Sub-Saharan cline from [[Southern Sudan]] to [[Northern Egypt]].<ref name="kings">{{cite journal|year=1992|last=Kings|title=mtDNA Analysis of Nile River Valley Populations: Genetic Corridor or a Barrier to Migration?|first9=A|last9=Di Rienzo|first8=D|last8=Welsby|first7=C|last7=Simon|first6=L|last6=Chaix|first5=AK|last5=Malek|first4=H|last4=Geisert|first3=K|last3=Bauer|first2=AE|pmid=10090902|url=http://genapps.uchicago.edu/labweb/pubs/krings.pdf|last2=Salem|pmc=1377841|first1=T|volume=64|issue=5|pages=1116–76|journal=Am J Hum Genet.|doi=10.1086/302314}}</ref> Another mtDNA study of modern Egyptians from the [[Kurna|Gurna]] region near [[Thebes, Egypt|Thebes]] in Southern Egypt by Stevanovicth et al. 2004 revealed that [[Eurasia]]n [[Recent African origin of modern humans|Out of Africa]] haplogroups represented 61.8% of the population, with the remainder being of [[Sub-Saharan]] (20.6%) and with a high frequency (17.6%) of haplogroup M1. According to the authors "This sedentary population presented similarities to the Ethiopian population by the L1 and L2 macrohaplogroup frequency (20.6%), by the West Eurasian component (defined by haplogroups H to K and T to X) and particularly by a high frequency (17.6%) of haplogroup M1... Our results suggest that the Gurna population has conserved the trace of an ancestral genetic structure from an ancestral East African population, characterized by a high M1 haplogroup frequency". The oral tradition of the Gurna people indicates that they, like most modern day Egyptians, descend from the [[Ancient Egyptians]] <ref name="stevanovitch">{{cite journal|title=Mitochondrial DNA Sequence Diversity in a Sedentary Population from Egypt |year=2004 |doi=10.1046/j.1529-8817.2003.00057.x|url=http://www3.interscience.wiley.com/cgi-bin/fulltext/118745570/HTMLSTART|last=Stevanovitch|first1=A.|last2=Gilles|first2=A.|last3=Bouzaid|first3=E.|last4=Kefi|first4=R.|last5=Paris|first5=F.|last6=Gayraud|first6=R. P.|last7=Spadoni|first7=J. L.|last8=El-Chenawi|first8=F.|last9=Beraud-Colomb|first9=E.|journal=Annals of Human Genetics|volume=68|pages=23–39|pmid=14748828|issue=Pt 1}}</ref>

Luis et al. (2004) found that the male haplogroups in a sample of 147 Egyptians were [[Haplogroup E1b1b (Y-DNA)|E1b1b]] (36.1%, predominantly [[Haplogroup E1b1b1a (Y-DNA)|E-M78]]), [[Haplogroup J (Y-DNA)|J]] (32.0%), [[Haplogroup G (Y-DNA)|G]] (8.8%), [[Haplogroup T (Y-DNA)|T]](8.2%), and [[Haplogroup R (Y-DNA)|R]] (7.5%). E1b1b and its subclades are characteristic of some [[Afro-Asiatic]] speakers and are believed to have originated in either the [[Near East]], [[North Africa]], or the [[Horn of Africa]]. Cruciani et al. (2007) suggests that E-M78, E1b1b predominant subclade in Egypt, originated in "Northeastern Africa", which in the study refers specifically to [[Egypt]] and [[Libya]] <ref name="luis">The Levant versus the Horn of Africa: Evidence for Bidirectional Corridors of Human Migrations – Luis; Rowold; Regueiro; Caeiro; Cinnioğlu; Roseman; Underhill; Cavalli-Sforza; and Herrera. – see http://www.pubmedcentral.nih.gov/articlerender.fcgi?artid=1182266</ref><ref>Underhill (2002), Bellwood and Renfrew, ed., Inference of Neolithic Population Histories using Y-chromosome Haplotypes, Cambridge: McDonald Institute for Archaeological Research, ISBN 1-902937-20-1
</ref>

The results of these genetic studies is consistent with the historical record, which records significant bidirectional contact between Egypt and Nubia, and the [[Levant]]/[[Near East]] within the last few thousand years, but with general population continuity from the [[Early Dynastic]] period up to the modern day era.<ref name="krings">{{cite journal|year=1992|last=Krings|title=mtDNA Analysis of Nile River Valley Populations: Genetic Corridor or a Barrier to Migration?|first9=A|last9=Di Rienzo|first8=D|last8=Welsby|first7=C|last7=Simon|first6=L|last6=Chaix|first5=AK|last5=Malek|first4=H|last4=Geisert|first3=K|last3=Bauer|first2=AE|pmid=10090902|url=http://genapps.uchicago.edu/labweb/pubs/krings.pdf|last2=Salem|pmc=1377841|first1=T|volume=64|issue=5|pages=1116–76|journal=Am J Hum Genet.|doi=10.1086/302314}}</ref><ref name="lucotte">{{cite journal|title=Brief communication: Y-chromosome haplotypes in Egypt |year=2001|last=Lucotte|doi=10.1002/ajpa.10190|url=http://wysinger.homestead.com/haplotypes_in_egypt.pdf|first1=G.|last2=Mercier|first2=G.|journal=American Journal of Physical Anthropology|volume=121|pages=63–6|pmid=12687584|issue=1}}</ref>

====Y-Dna haplogroups====
{| class="wikitable sortable" style="text-align:center; font-size: 75%"; border="1"
| align="center" style="background:#f0f0f0;"|'''Population'''
| align="center" style="background:#f0f0f0;"|'''Nb'''
| align="center" style="background:#f0f0f0;"|'''A/B'''
| align="center" style="background:#f0f0f0;"|'''E(xE1b1b)'''
| align="center" style="background:#f0f0f0;"|'''E1b1b1 (M35)'''
| align="center" style="background:#f0f0f0;"|'''E1b1b1a (M78)'''
| align="center" style="background:#f0f0f0;"|'''E1b1b1b (M81)'''
| align="center" style="background:#f0f0f0;"|'''E1b1b1c (M123)'''
| align="center" style="background:#f0f0f0;"|'''F'''
| align="center" style="background:#f0f0f0;"|'''K'''
| align="center" style="background:#f0f0f0;"|'''G'''
| align="center" style="background:#f0f0f0;"|'''I'''
| align="center" style="background:#f0f0f0;"|'''J1'''
| align="center" style="background:#f0f0f0;"|'''J2'''
| align="center" style="background:#f0f0f0;"|'''R1a'''
| align="center" style="background:#f0f0f0;"|'''R1b'''
| align="center" style="background:#f0f0f0;"|'''Other'''
| align="center" style="background:#f0f0f0;"|'''Study'''
|-
| 1 Egyptians ||147||2.7%||2.7%||0||18.4%||8.2%||9.5%||0||7.5%||9.5%||0||19.7%||12.2%||3.4% ||4.1%||2.1%||Luis et al. (2004)<ref>Luis JR, Rowold DJ, Regueiro M, Caeiro B, Cinnioglu C, Roseman C, Underhill PA, Cavalli-Sforza LL, Herrera RJ (2004) [http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1182266/ The Levant versus the Horn of Africa: evidence for bidirectional corridors of human migrations]. Am J Hum Genet 74:532–544</ref>
|-
| 2 Egyptians from El-Hayez Oasis (Western Desert)||35||0||5.70%||5.7%||28.6%||28.6%||0||0||0||0||0||31.4%||0||0||0||0||Kujanová et al. (2009)<ref>Martina Kujanová, Luísa Pereira, Verónica Fernandes, Joana B. Pereira, Viktor Černý, [http://onlinelibrary.wiley.com/doi/10.1002/ajpa.21078/abstract Near Eastern Neolithic genetic input in a small oasis of the Egyptian Western Desert], American Journal of Physical Anthropology, Volume 140, Issue 2, pages 336–346, October 2009</ref>
|-
| 3 Egyptians from [[Siwa Oasis]] (Western Desert)||93||28.0%||6.5%||2.2%||6.5%||1.1%||2.2%||0||0||3.2%||0||7.5%|| 6.5%||0||28.0%||8.3%||Dugoujon et al. (2009)<ref>Dugoujon J.M., Coudray C., Torroni A., Cruciani F., Scozzari F., Moral P., Louali N., Kossmann M. ''The Berber and the Berbers: Genetic and linguistic diversities''</ref>
|-
| 4 Northern Egyptians ||44||2.3%||0||4.5%||27.3%||11.4%||9.1%||6.8%||2.3%||0||0||9.1%||9.1%||2.3%||9.9%||6.8%||Arredi et al. (2004)
|-
| 5 Southern Egyptians ||29||0.0%||0||0||17.2%||6.9%||6.9%||17.2%||10.3%||0||3.4%||20.7%||3.4%||0||13.8%||0||Arredi et al. (2004)
|-
|}

;Distribution of E1b1b1a (E-M78) and its subclades
{| class="wikitable sortable" style="text-align:center; font-size: 70%"; border="1"
| align="center" style="background:#f0f0f0;"|'''Population'''
| align="center" style="background:#f0f0f0;"|'''N'''
| align="center" style="background:#f0f0f0;"|'''E-M78'''
| align="center" style="background:#f0f0f0;"|'''E-M78*'''
| align="center" style="background:#f0f0f0;"|'''E-V12*'''
| align="center" style="background:#f0f0f0;"|'''E-V13'''
| align="center" style="background:#f0f0f0;"|'''E-V22'''
| align="center" style="background:#f0f0f0;"|'''E-V32'''
| align="center" style="background:#f0f0f0;"|'''E-V65'''
| align="center" style="background:#f0f0f0;"|'''Study'''
|-
| Southern Egyptians||79||50.6%||||44.3%||1.3%||3.8%||||1.3%|| Cruciani et al. (2007)<ref>Cruciani, F.; La Fratta, R.; Trombetta, B.; Santolamazza, P.; Sellitto, D.; Colomb, E. B.; Dugoujon, J.-M.; Crivellaro, F. et al. (2007), "[http://mbe.oxfordjournals.org/content/24/6/1300.full.pdf+html Tracing Past Human Male Movements in Northern/Eastern Africa and Western Eurasia: New Clues from Y-Chromosomal Haplogroups E-M78 and J-M12]", Molecular Biology and Evolution 24 (6): 1300–1311, DOI:10.1093/molbev/msm049, PMID 17351267</ref>
|-
| Egyptians from Bahari||41||41.4%||||14.6%||2.4%||21.9%||||2.4%|| Cruciani et al. (2007)
|-
| Northern Egyptians (Delta)||72||23.6%||||5.6%||1.4%||13.9%||2.8%|||| Cruciani et al. (2007)
|-
| Egyptians from Gurna Oasis||34||17.6%||5.9%||8.8%||||||2.9%|||| Cruciani et al. (2007)
|-
| Egyptian from Siwa Oasis||93||6.4%||||2.1%||||||||4.3%|| Cruciani et al. (2007)
|-
|}

==== Autosomal DNA ====
In 13 January 2012, an exhaustive genetic study of North Africa's human populations was published.<ref>Henn BM, Botigué LR, Gravel S, Wang W, Brisbin A, et al. (2012) [http://www.plosgenetics.org/article/info%3Adoi%2F10.1371%2Fjournal.pgen.1002397 Genomic Ancestry of North Africans Supports Back-to-Africa Migrations]. PLoS Genet 8(1): e1002397. {{doi|10.1371/journal.pgen.1002397}}</ref> The researchers analyzed around 800,000 genetic markers, distributed throughout the entire genome in 125 North African individuals belonging to seven representative populations in the whole region (Saharawi, South Moroccans, North Moroccans, Algerians, Tunisians Berbers, Libyans and Egyptians) and the information obtained was compared with the information from the neighbouring populations. The results of this study show that there is a native genetic component ("Maghrebi") which defines North Africans. The study identified mainly two distinct, opposite gradients of ancestry: an east-to-west increase of this native North African ancestry and an east-to-west decrease in likely Near Eastern Arabic ancestry.

The study also reveals that the genetic composition of North Africa's human populations is very complex and is the result of five distinct ancestries : a local component (Maghrebi) dating back thirteen thousand years and the varied genetic influence of neighbouring populations on North African groups during successive migrations (European, Near Eastern, eastern and western sub-Saharan Africa). According to the authors, the people inhabiting North Africa today are not descendants of either the earliest occupants of this region fifty thousand years ago, or descendants of the most recent Neolithic populations. The data shows that the ancestors of today's North Africans were a group of populations which already lived in the region around thirteen thousand years ago. Furthermore, this local North African genetic component is very different from the one found in the populations in the south of the Sahara, which shows that the ancestors of today's North Africans were members of a subgroup of humanity who left Africa to conquer the rest of the world and who subsequently returned to the north of the continent to settle in the region. As well as this local component, North African populations were also observed to share genetic markers with all the neighbouring regions, as a result of more recent migrations, although these appear in different proportions. There is an influence from the Middle East, which becomes less marked as the distance from the Arabian Peninsula increases, similar proportions of European influence in all North African populations, and, in some populations (South Moroccans, Saharawi...), there are even individuals who present a large proportion of recent influence from the South of the Sahara in their genome.

=====Admixture analysis=====
Recent genetic analysis of North African populations have found that, despite the complex admixture genetic background, there is an autochthonous genomic component which is likely derived from "back-to-Africa" gene flow older than 12,000 years ago (ya) (i.e., prior to the Neolithic migrations). This local population substratum seems to represent a genetic discontinuity with the earliest modern human settlers of North Africa (those with the Aterian industry) given the estimated ancestry is younger than 40,000 years ago. North Morocco, Libya and Egypt carry high proportions of European and Near Eastern ancestral components, whereas Tunisian Berbers and Saharawi are those populations with highest autochthonous North African component.<ref>[http://www.plosone.org/article/info:doi/10.1371/journal.pone.0047765?imageURI=info:doi/10.1371/journal.pone.0047765.g001 North African Populations Carry the Signature of Admixture with Neandertals]. Sánchez-Quinto F, Botigué LR, Civit S, Arenas C, Ávila-Arcos MC, et al. (2012) North African Populations Carry the Signature of Admixture with Neandertals. PLoS ONE 7(10): e47765. doi:10.1371/journal.pone.0047765</ref>

;Average ancestry proportions in North African populations estimated by ADMIXTURE for k = 4 different ancestries (October 2012)
{| class="wikitable sortable" border=1
| align="center" style="background:#f0f0f0;"|'''Population'''
| align="center" style="background:#f0f0f0;"|'''N'''
| align="center" style="background:#f0f0f0;"|'''Maghreb'''
| align="center" style="background:#f0f0f0;"|'''Europe'''
| align="center" style="background:#f0f0f0;"|'''Near East'''
| align="center" style="background:#f0f0f0;"|'''Sub-Saharan Africa'''
|-
| Tunisia (Berbers)||18||93%||4%||2%||1%
|-
| Saharawi||18||55%||17%||10%||18%
|-
| Morocco North||18||44%||31%||14%||11%
|-
| Morocco South||16||44%||13%||10%||33%
|-
| Algeria||19||39%||27%||16%||18%
|-
| Libya||17||31%||28%||25%||16%
|-
| Egypt||19||19%||37%||30%||14%
|-
|}

===DNA studies on King Tut and His Relatives===
[[Switzerland|Swiss]] scientists at Zurich-based DNA genealogy centre, iGENEA, reconstructed DNA profile of Tutankhamun, based on a film that was made for the Discovery Channel, which showed that Tutankhamun has [[Haplogroup R1b (Y-DNA)|Haplogroup R1b1a2]], to which more than 50% of European men and less than 1% of modern-day Egyptians belong to.<ref>[http://uk.reuters.com/article/2011/08/01/oukoe-uk-britain-tutankhamun-dna-idUKTRE7704OR20110801 Half of European men share King Tut's DNA]. [[Reuters]], 1 August 2011. Retrieved on 6 August 2011</ref> However the Y-chromosome of King Tut has never been published and DNA profile displayed in Discovery Channel documentary may not belong to the Pharaoh. According to Carsten Pusch, a geneticist at Germany's University of Tübingen who was part of the team that unraveled Tut's DNA from samples taken from his mummy and mummies of his family members, iGENEA's claims are "simply impossible." <ref>[http://www.igenea.com/en/index.php?c=62 The Tutankhamun DNA Project], Igenea,</ref><ref>[http://www.livescience.com/15388-discovery-channel-tutankhamen-dna.html King Tut Related to Half of European Men? Maybe Not], LiveScience, 3 August 2011</ref> R1b also shows up in parts of northern Africa, particularly some regions in Algeria, where tests have found it in 11.8% of subjects.<ref name=Robino>{{Cite journal| title=Analysis of Y-chromosomal SNP haplogroups and STR haplotypes in an Algerian population sample| first9=C| last9=Torre| first8=S| last8=Inturri| first7=A| last7=Piazza| first6=N| last6=Cerutti| first5=S| last5=Benhamamouch| first4=A| last4=Bekada| first3=C| last3=Di Gaetano| journal=Journal International Journal of Legal Medicine| first2=F| volume=122|issue=3| last2=Crobu|year=2008| pmid=17909833|doi=10.1007/s00414-007-0203-5| author=Robino et al.| pages=251–5| postscript=<!-- Bot inserted parameter. Either remove it; or change its value to "." for the cite to end in a ".", as necessary. -->}}</ref> It is also found in central Africa around Chad and Cameroon,<ref>{{Cite journal|pmid=11910562 |pmc=447595 |doi=10.1086/340257 |year=2002 |last1=Cruciani |first1=F |last2=Santolamazza |last3=Shen |last4=Macaulay |last5=Moral |last6=Olckers |last7=Modiano |last8=Holmes |last9=Destro-Bisol |title=A back migration from Asia to sub-Saharan Africa is supported by high-resolution analysis of human Y-chromosome haplotypes. |volume=70 |issue=5 |pages=1197–214 |journal=American Journal of Human Genetics |first2=P |first3=P |first4=V |first5=P |first6=A |first7=D |first8=S |first9=G}}, pp. 13–14</ref> but the Chadic-speaking area in Africa is dominated by the branch known as R1b1c (R-V88).<ref name=Cruciani2010>{{Cite journal|author=Cruciani et al.|year=2010|journal=European Journal of Human Genetics | doi=10.1038/ejhg.2009.231|title=Human Y chromosome haplogroup R-V88: a paternal genetic record of early mid Holocene trans-Saharan connections and the spread of Chadic languages|pmc=2987365|url=http://www.nature.com/ejhg/journal/vaop/ncurrent/abs/ejhg2009231a.html|pmid=20051990|last2=Trombetta|first2=B|last3=Sellitto|first3=D|last4=Massaia|first4=A|last5=Destro-Bisol|first5=G|last6=Watson|first6=E|last7=Beraud Colomb|first7=E|last8=Dugoujon|first8=JM|last9=Moral|first9=P|volume=18|issue=7|pages=800–7|postscript=<!--None-->}}</ref>

In December 2011, the private genetics research company DNA Tribes released an analysis, based on 8 forensic autosomal STR markers. The study in particular analyzed the DNA of the Amarna Pharaohs and reported that "Average MLI scores in Table 1 indicate the STR profiles of the Amarna mummies would be most frequent in present day populations of several African regions". (Top MLI (Match Likelihood Index) scores for Amarna mummies based on the world regions identified by DNA Tribes® STR analysis. Each MLI score identifies the likelihood of occurrence of an STR profile in that region versus the likelihood of occurrence in the world as a whole.). These regional matches do not necessarily indicate an exclusively African ancestry for the
Amarna pharaonic family. However, results indicate these ancient individuals inherited some alleles that
today are more frequent in populations of Africa than in other parts of the world.
<ref>{{cite web|last=Digest|first=DNATribes|title=Last of the Amarna Pharaohs: King Tut and His Relatives|url=http://www.dnatribes.com/dnatribes-digest-2012-01-01.pdf|publisher=DNATribes}}</ref>

{| class="wikitable sortable" style="text-align:center; font-size: 70%"; border="1"
| align="center" style="background:#f0f0f0;"|'''World Region MLI'''
| align="center" style="background:#f0f0f0;"|'''Thuya'''
| align="center" style="background:#f0f0f0;"|'''Yuya'''
| align="center" style="background:#f0f0f0;"|'''KV35EL'''
| align="center" style="background:#f0f0f0;"|'''Amen-hotepIII'''
| align="center" style="background:#f0f0f0;"|'''KV55'''
| align="center" style="background:#f0f0f0;"|'''KV35YL'''
| align="center" style="background:#f0f0f0;"|'''Tut'''
| align="center" style="background:#f0f0f0;"|'''Average'''
|-
| Southern African||359.72||34.48||20.73||108.53||174.9||71.17||'''1,519.03'''||326.94
|-
| African Great Lakes||233.49||35.53||20.87||222.53||381.3||44.58||1,328.01||323.76
|-
| Tropical West African||142.84||8.91||6.93||37.43||53.03||22.99||314||83.74
|-
| Horn Of Africa||14.65||0.79||5.17||12.03||4.54||22||44.35||14.79
|-
| Sahelian||39.14||0.74||5.76||2.97||4.4||16.85||30.41||14.33
|-
| Levantine||0.4||1.56||0.66||10.3||6.07||8.4||21.08||6.92
|-
| Aegean||0.12||0.35||0.87||9.06||7.05||20.16||9.85||6.78
|-
| Arabian||0.12||0.42||0.7||5.58||2.83||21.41||10.91||6
|-
| Northwest European||0.21||0.28||1.26||3.99||10.41||15.01||5.33||5.21
|-
| Mediterranean||0.08||0.23||0.74||4.54||5.81||16.8||6.04||4.89
|-
| North African||2.22||0.21||0.75||3.39||3.25||12.63||6.55||4.14
|-
| Mesopotamian||0.06||0.41||0.63||6.24||2.69||11.54||5.27||3.84
|-
|}

===Ramesses III===
A 2012 study done on the mummified remains of [[Ramesses III]] determined that the pharaoh's [[y-chromosome]] belonged to [[Haplogroup E1b1a (Y-DNA)]]. The pharaoh's y-chromosome belongs to the most frequent haplogroup among [[Sub-Saharan African]] y-chromosomes.<ref>{{cite journal|title=Revisiting the harem conspiracy and death of Ramesses III: anthropological, forensic, radiological, and genetic study|doi=10.1136/bmj.e8268|year=2012|last=Hawass|publisher=BMJ}}</ref>


==Anthropometric indicators==
==Anthropometric indicators==

Revision as of 09:49, 18 January 2013

The land currently known as Egypt has a long and involved population history. This is partly due to its geographical location at the crossroads of several major cultural areas: the Mediterranean, the Middle East, the Sahara and East Africa. In addition Egypt has experienced several invasions during its long history, including by the Canaanites, the Libyans, the Nubians, the Assyrians, the Kushites, the Persians, the Greeks, the Romans and the Arabs. The various conquests over time have made the relationship between Modern Egyptians and Ancient Egyptians a topic of investigation.

Prehistory

During the Paleolithic the Nile Valley was inhabited by various hunter gatherer populations. About 10,000 years ago the Sahara Desert had a wet phase. People from the surrounding areas moved from the Sahara, and evidence suggests that the populations of the Nile Valley reduced in size.[1] About 5,000 years ago the wet phase of the Sahara came to end. Saharan population retreated to the south towards the Sahel, and East towards the Nile Valley. It was these populations, in addition to Neolithic farmers from the Near East, that played a major role in the formation of the Egyptian state as they brought their food crops, sheep, goats and cattle to the Nile Valley.[1][2]

Predynastic Egypt

The Predynastic period dates to the end of the fourth millennium BC. From about 4800 to 4300BC the Merimde culture flourished in Lower Egypt.[3] This culture, among others, has links to the Levant.[4] The pottery of the Buto Maadi culture, best known from the site at Maadi near Cairo, also shows connections to the southern Levant.[5]

In Upper Egypt the predynastic Badarian culture was followed by the Naqada culture. The origins of these people is still not fully understood.

Biogeographic origin based on cultural data

Located in the extreme north-east corner of Africa, Ancient Egyptian society was at a crossroads between the African and Near Eastern regions. Early proponents of the Dynastic Race Theory based their hypothesis on the increased novelty and seemingly rapid change in Predynastic pottery and noted trade contacts between ancient Egypt and the Middle East.[6] This is no longer the dominant view in Egyptology, however the evidence on which it was based still suggests influence from these regions.[7] Fekri Hassan and Edwin et al. point to mutual influence from both inner Africa as well as the Levant.[8] However according to one author this influence seems to have had minimal impact on the indigenous populations already present.[9]

One author has stated that the Naqada phase of Predynastic Egyptians in Upper Egypt shared an almost identical culture with A-group peoples of the Lower Sudan.[10] Based in part on the similarities at the royal tombs at Qustul, some scholars have even proposed an Egyptian origin in Nubia among the A-group.[11][12] In 1996 Lovell and Prowse reported the presence of individual rulers buried at Naqada in what they interpreted to be elite, high status tombs, showing them to be more closely related morphologically to populations in Northern Nubia than those in Southern Egypt.[13] Most scholars however, have rejected this hypothesis and cite the presence of royal tombs that are contemporaneous with that of Qustul and just as elaborate, together with problems with the dating techniques.[14]

Toby Wilkinson, in his book "Genesis of the Pharaohs", proposes an origin for the Egyptians somewhere in the Eastern Desert.[15] He presents evidence that much of predynastic Egypt duplicated the traditional African cattle-culture typical of Southern Sudanese and East African pastoralists of today. Kendall agrees with Wilkinson's interpretation that ancient rock art in the region may depict the first examples of the royal crowns, while also pointing to Qustul in Nubia as a likely candidate for the origins of the white crown, being that the earliest known example of it was discovered in this area.

There is also evidence that sheep and goats were introduced into Nabta from Southwest Asia about 8,000 years ago.[2] There is some speculation that this culture is likely to be the predecessor of the Egyptians, based on cultural similarities and social complexity which is thought to be reflective of Egypt's Old Kingdom.[16][17]

Contamination from handling and intrusion from microbes create obstacles to the recovery of Ancient DNA.[18] Consequently most DNA studies have been carried out on modern Egyptian populations with the intent of learning about the influences of historical migrations on the population of Egypt.[19][20][21][22]

In general, various DNA studies have found that the gene frequencies of North African populations are intermediate between those of the Near East, the Horn of Africa, southern Europe and Sub Saharan Africa,[23] though Egypt’s NRY frequency distributions appear to be much more similar to those of the Middle East than to any sub-Saharan African population, suggesting a much larger Eurasian genetic component.[24][25][25][26][27][28][29]

Blood typing and DNA sampling on ancient Egyptian mummies is scant; however, blood typing of dynastic mummies found ABO frequencies to be most similar to modern Egyptians[30] and some also to Northern Haratin populations. ABO blood group distribution shows that the Egyptians form a sister group to North African populations, including Berbers, Nubians and Canary Islanders.[31] Scholars such as Frank Yurco believe that Modern Egyptians are largely representative of the ancient population, and the DNA evidence appears to support this view.[32]

Anthropometric indicators

Craniofacial criteria

Ancient Egyptian skull, thought to be that of Akhenaten.

Craniofacial criteria are no longer universally accepted as reliable indicators of population grouping or ethnicity. In 1912 Franz Boas demonstrated that cranial shape is heavily influenced by environmental factors, and can change within a few generations if conditions change, and therefore cranial measurements cannot be a reliable indicator of inherited influences such as ethnicity.[33] This conclusion was supported in 2003 in a paper by Gravlee, Bernard and Leonard.[34][35] A study by Beals, Smith, and Dodd (1984) found that "race" and cranial variation had low correlations, and that cranial variation was instead strongly correlated with climate variables.[36] This view is also supported by Kemp.[37] Other studies have shown that the typical cranial shapes of some African (Sudanic and Ethiopic), Arab and Berber ethnic groups are largely the same.[38][39]

A craniofacial study by C. Loring Brace et al. (1993) concluded that: "The Predynastic of Upper Egypt and the Late Dynastic of Lower Egypt are more closely related to each other than to any other population. As a whole, they show ties with the European Neolithic, North Africa, modern Europe, and, more remotely, India, but not at all with sub-Saharan Africa, eastern Asia, Oceania, or the New World. Adjacent people in the Nile valley show similarities in trivial traits in an unbroken series from the delta in the north southward through Nubia and all the way to Somalia at the equator. At the same time, the gradient in skin color and body proportions suggests long-term adaptive response to selective forces appropriate to the latitude where they occur. An assessment of “race” is as useless as it is impossible. Neither clines nor clusters alone suffice to deal with the biological nature of a widely distributed population. Both must be used."[40] He also commented, "We conclude that the Egyptians have been in place since back in the Pleistocene and have been largely unaffected by either invasions or migrations. As others have noted, Egyptians are Egyptians, and they were so in the past as well." The results of this study, however, have been criticized and contradicted by the works of other anthropologists such as S.O.Y. Keita and Sonia Zakrzewski.

A survey cited by Kemp (2005) of pooled ancient Egyptian crania spanning all time periods found that the Egyptian population as a whole clusters more closely to modern Egyptians than to other groups, but apart from modern Egyptians, they cluster closest to Nubian and "Ethiopic" populations than they do to Middle Easterners or Europeans. In Kemp's unpooled dendrogram it details that the Pre-Dynastic Egyptians (El Bardi and Naqada) samples cluster closest to ancient Nubians and modern Ethiopic populations, and conversely that Late Kingdom and modern Egyptians cluster with Middle Eastern and modern European populations. Kemp also noted that Egypt conquered and settled Nubia beginning in the 1st Dynasty.[41]

Anthropologist Nancy Lovell states the following:

"There is now a sufficient body of evidence from modern studies of skeletal remains to indicate that the ancient Egyptians, especially southern Egyptians, exhibited physical characteristics that are within the range of variation for ancient and modern indigenous peoples of the Sub Sahara and tropical Africa.. In general, the inhabitants of Upper Egypt and Nubia had the greatest biological affinity to people of the Sahara and more southerly areas." "must be placed in the context of hypotheses informed by archaeological, linguistic, geographic and other data. In such contexts, the physical anthropological evidence indicates that early Nile Valley populations can be identified as part of an African lineage, but exhibiting local variation. This variation represents the short and long term effects of evolutionary forces, such as gene flow, genetic drift, and natural selection, influenced by culture and geography."[42]

This view was also shared by the late Egyptologist Frank Yurco.[43]

A 2005 study by Keita of predynastic Badarian (Southern Egyptian) crania found that the Badarian samples cluster more closely with East African (Ethiopic) samples than they do with Northern European (Berg and Norse) samples, though importantly no Asian and Southern Africa samples were included in the study.[44] Keita has also said that the predyastic crania are different to the lower Egyptian samples, which display a mean part way between modern Sub Saharans Africans and Ethiopians.

Sonia Zakrzewski in 2007 noted that population continuity occurs over the Egyptian Predynastic into the Greco-Roman periods, and that a relatively high level of genetic differentiation was sustained over this time period. She concluded therefore that the process of state formation itself may have been mainly an indigenous process, but that it may have occurred in association with in-migration, particularly during the Early Dynastic and Old Kingdom periods.[45]

In 2008 Keita found that the early predynastic groups in Southern Egypt were similar craniometrically to Nile valley groups of Ethiopic extraction, and as a whole the dynastic Egyptians (includes both Upper and Lower Egyptians) show much closer affinities more southerly Northeast African populations. He also concluded that more material is needed to make a firm conclusion about the relationship between the early Holocene Nile valley populations and later ancient Egyptians.[46]

Limb ratios

Anthropologist C. Loring Brace points out that limb elongation is "clearly related to the dissipation of metabolically generated heat" in areas of higher ambient temperature. He also stated that "skin color intensification and distal limb elongation is apparent wherever people have been long-term residents of the tropics". He also points out that the term "super negroid" is inappropriate, as it is also applied to non negroid populations. These features have been observed among Egyptian samples.[47] According to Robins and Shute the average limb elongation ratios among ancient Egyptians is higher than that of modern West Africans who reside much closer to the equator. Robins and Shute therefore term the ancient Egyptians to be "super-negroid" but state that although the body plans of the ancient Egyptians were closer to those of modern negroes than for modern whites, "this does not mean that the ancient Egyptians were negroes".[48] Anthropologist S.O.Y. Keita criticized Robins and Shute, stating they do not interpret their results within an adaptive context, and stating that they imply "misleadingly" that early southern Egyptians were not a "part of the Saharo-tropical group, which included Negroes".[49] Gallagher et al. also points out that "body proportions are under strong climatic selection and evidence remarkable stability within regional lineages".[50] Zakrzewski (2003) studied skeletal samples from the Badarian period to the Middle Kingdom. She confirmed the results of Robins and Shute that Ancient Egyptians in general had "tropical body plans" but that their proportions were actually "super-negroid".[51]

Trikhanus (1981) found Egyptians to plot closest to tropical Africans and not Mediterranean Europeans residing in a roughly similar climatic area.[52] A more recent study compared ancient Egyptian osteology to that of African-Americans and White Americans, and found that the stature of the Ancient Egyptians was more similar to the stature of African-Americans, although it was not identical:[53]

Our results confirm that, although ancient Egyptians are closer in body proportion to modern American Blacks than they are to American Whites, proportions in Blacks and Egyptians are not identical.

Dental morphology

Modern studies on ancient Egyptian dentition clusters the Ancient Egyptians with Caucasoids (Europeans, Western Asians) who have small teeth, as opposed to Negroids (Western Sub-Saharan Africans) who have megadont/large teeth.[54][55]

A 2006 bioarchaeological study on the dental morphology of ancient Egyptians by Prof. Joel Irish shows dental traits characteristic of current indigenous North Africans and to a lesser extent Middle Eastern and southern European populations, but not at all to Sub-Saharan populations. Among the samples included in the study is skeletal material from the Hawara tombs of Fayum, (from the Roman period) which clustered very closely with the Badarian series of the predynastic period. All the samples, particularly those of the Dynastic period, were significantly divergent from a neolithic West Saharan sample from Lower Nubia. Biological continuity was also found intact from the dynastic to the post-pharaonic periods. According to Irish:

[The Egyptian] samples [996 mummies] exhibit morphologically simple, mass-reduced dentitions that are similar to those in populations from greater North Africa (Irish, 1993, 1998a–c, 2000) and, to a lesser extent, western Asia and Europe (Turner, 1985a; Turner and Markowitz, 1990; Roler, 1992; Lipschultz, 1996; Irish, 1998a).[56]

Anthropologist Shomarka Keita takes issue with the suggestion of Irish that Egyptians and Nubians were not primary descendants of the African epipaleolithic and Neolithic populations. Keita also criticizes him for ignoring the possibility that the dentition of the ancient Egyptians could have been caused by "in situ microevolution" driven by dietary change, rather than by racial admixture.[57] However Keita himself has observed population continuity from the Pleistocene to the present in modern Egyptians.

The language element

The Edwin Smith papyrus, the world's oldest surviving surgical document. Written in Hieratic script in Ancient Egypt around 1600 B.C.

Ancient Egyptian languages are classified into six major chronological divisions; Archaic Egyptian, Old Egyptian, Middle Egyptian, Late Egyptian, Demotic Egyptian, and Coptic, the latter of which was still used as a working language until the 18th Century AD, and is still used as a liturgical language by Egyptian Copts to this day.[58]

Origins

The Ancient Egyptian language has been classified as a member of the Afro-Asiatic language family. There is no agreement on when and where these languages originated, though the language is generally believed to have originated somewhere in or near the region stretching from the Levant in the Near East to northern Kenya, and from the Eastern Sahara in North Africa to the Red Sea, or Southern Arabia, Ethiopia and Sudan.[59][60][61][62][63] The language of the neighbouring Nubian people is one of the Nilo-Saharan languages, and is not one of the Afro-Asiatic languages.[64][65]

Scripts

Most surviving texts in the Egyptian language are primarily written in the Hieroglyphic script. However, the Hieratic script was used in parallel although mostly reserved for priests[66] but also for magical purposes and administrative and legal documents. This script used cursive and simplified forms of the Hieroglyphic writing to ease and speed writing. It was mostly written on papyrus, wood, leather and ostraca.[67] It lasted until the ninth century BCE when it was replaced by the "Abnormal Hieratic" in southern Egypt.[67] Another third script was the Demotic which was developed in Lower Egypt during the later part of the 25th Dynasty.

See also

References

  1. ^ a b Ancient Egyptian Origins
  2. ^ a b http://www.comp-archaeology.org/WendorfSAA98.html
  3. ^ Bogucki, Peter I. (1999). The origins of human society. Wiley-Blackwell. p. 355. ISBN 1-57718-112-3.
  4. ^ Josef Eiwanger: Merimde Beni-salame, In: Encyclopedia of the Archaeology of Ancient Egypt. Compiled and edited by Kathryn A. Bard. London/New York 1999, p. 501-505
  5. ^ Jürgen Seeher. Ma'adi and Wadi Digla. in: Encyclopedia of the Archaeology of Ancient Egypt. Compiled and edited by Kathryn A. Bard. London/New York 1999, 455-458
  6. ^ Hoffman. "Egypt before the pharaohs: the prehistoric foundations of Egyptian civilization", pp267
  7. ^ Redford, Egypt, Israel, p. 17.
  8. ^ Edwin C. M et. al, "Egypt and the Levant", pp514
  9. ^ Toby A.H. Wilkinson. " Early Dynastic Egypt: Strategies, Society and Security", pp.15
  10. ^ Hunting for the Elusive Nubian A-Group People - by Maria Gatto, archaeology.org
  11. ^ Egypt and Sub-Saharan Africa: Their Interaction - Encyclopedia of Precolonial Africa, by Joseph O. Vogel, AltaMira Press, (1997), pp. 465-472
  12. ^ Journal of Near Eastern Studies, Vol. 46, No. 1 (Jan., 1987), pp. 15-26
  13. ^ Tracy L. Prowse, Nancy C. Lovell. Concordance of cranial and dental morphological traits and evidence for endogamy in ancient Egypt, American Journal of Physical Anthropology, Vol. 101, Issue 2, October 1996, Pages: 237-246
  14. ^ Wegner, J. W. 1996. Interaction between the Nubian A-Group and Predynastic Egypt: The Significance of the Qustul Incense Burner. In T. Celenko, Ed., Egypt in Africa: 98-100. Indianapolis: Indianapolis Museum of Art/Indiana University Press.
  15. ^ Genesis of the Pharaohs: Genesis of the ‘Ka’ and Crowns? - Review by Timothy Kendall, American Archaeologist
  16. ^ Ancient Astronomy in Africa
  17. ^ Wendorf, Fred (2001). Holocene Settlement of the Egyptian Sahara. Springer. p. 525. ISBN 0-306-46612-0.
  18. ^ Cite error: The named reference books.google.com was invoked but never defined (see the help page).
  19. ^ S.O.Y. Keita & A. J. Boyce, S. O. Y.; Boyce, A. J. (Anthony J.) (June 2009). "Genetics, Egypt, and History: Interpreting Geographical Patterns of Y Chromosome Variation" (PDF). History in Africa. 32 (1): 221. doi:10.1353/hia.2005.0013. {{cite journal}}: Check date values in: |year= / |date= mismatch (help)
  20. ^ Shomarka Keita (2005), S. O. Y. (2005). "Y-Chromosome Variation in Egypt" (PDF). African Archaeological Review. 22 (2): 61. doi:10.1007/s10437-005-4189-4. ISBN [[Special:BookSources/43700541894 |43700541894 [[Category:Articles with invalid ISBNs]]]]. {{cite journal}}: Check |isbn= value: invalid character (help)CS1 maint: numeric names: authors list (link)
  21. ^ Keita, S.O.Y. (2005). "History in the Interpretation of the Pattern of p49a,f TaqI RFLP Y-Chromosome Variation in Egypt" (PDF). American Journal of Human Biology. 17 (5): 559–67. doi:10.1002/ajhb.20428. PMID 16136533.
  22. ^ Shomarka Keita: What genetics can tell us
  23. ^ Cavalli-Sforza, History and Geography of Human Genes, The intermediacy of North Africa and to a lesser extent Europe is apparent
  24. ^ The Levant versus the Horn of Africa: Evidence for Bidirectional Corridors of Human Migrations – Luis; Rowold; Regueiro; Caeiro; Cinnioğlu; Roseman; Underhill; Cavalli-Sforza; and Herrera. – see http://www.pubmedcentral.nih.gov/articlerender.fcgi?artid=1182266
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Further reading

External links