He obtained doctorates in medicine (1894) and sciences (1899) at the University of Caen. From 1904 to 1923, he was chair of zoology at the University of Montpellier, afterwards attaining the chair of marine biology at the Sorbonne. At the same time, he was also named director of the Arago laboratory in Banyuls-sur-Mer, and in 1931 became manager of the biological station at Villefranche-sur-Mer.
- La glande venimeuse de la scolopendre, 1894 – The venomous gland of Scolopendra.
- Recherches sur les chilopodes, 1899 – Research on chilopods.
- Titres et travaux scientifiques, 1904 – Scientific works.
- Études sur la sexualité chez les grégarines, 1909 (with Louis Jules Léger) – Studies on the sexuality of gregarines.