Picea sitchensis
Picea sitchensis Sitka spruce | |
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Quinault Lake Spruce, the largest member of the species according to American Forest by points | |
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Species: | P. sitchensis
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Binomial name | |
Picea sitchensis | |
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Range |
Picea sitchensis, the Sitka spruce, is a large coniferous evergreen tree growing to almost 100 m tall,[1] and with a trunk diameter at breast height that can exceed 5 m (see List of superlative trees. It is by far the largest species of spruce; and the fifth largest conifer in the world (behind giant sequoia, coast redwood, kauri and western redcedar);[2] and the third tallest conifer species (after coast redwood and coast Douglas-fir). The Sitka spruce is one of the few species documented to reach 300 feet in height.[3] It acquires its name from the community of Sitka, Alaska.
Description
![](http://upload.wikimedia.org/wikipedia/commons/thumb/3/3e/Picea_sitchensis1.jpg/220px-Picea_sitchensis1.jpg)
The bark is thin and scaly, flaking off in small circular plates 5–20 cm across. The crown is broad conic in young trees, becoming cylindric in older trees; old trees may not have branches lower than 30–40 m. The shoots are very pale buff-brown, almost white, and glabrous (hairless) but with prominent pulvini. The leaves are stiff, sharp and needle-like, 15–25 mm long, flattened in cross-section, dark glaucous blue-green above with two or three thin lines of stomata, and blue-white below with two dense bands of stomata.
The cones are pendulous, slender cylindrical, 6–10 cm long [4] and 2 cm broad when closed, opening to 3 cm broad. They have thin, flexible scales 15–20 mm long; the bracts just above the scales are the longest of any spruce, occasionally just exserted and visible on the closed cones. They are green or reddish, maturing pale brown 5–7 months after pollination. The seeds are black, 3 mm long, with a slender, 7–9 mm long pale brown wing.
Size
More than a century of logging has left only a remnant of the spruce forest. The largest trees were cut long before careful measurements could be made. Trees over 90 m tall may still be seen in the Pacific Rim National Park and Carmanah Walbran Provincial Park on Vancouver Island, British Columbia (the Carmanah Giant, at 96 m (315 ft) tall is the tallest tree in Canada), and in the Olympic National Park, Washington and Prairie Creek Redwoods State Park, California (USA); two at the last site are just over 96 m (315 ft) tall. The Queets Spruce is the largest in the world with a trunk volume of 337 m3 (11,900 cu ft) it is 75.6 m (248 ft) tall and 455 cm (15 ft) in dbh. It is located near the Queets River in Olympic National Park, about 26 kilometres (16 mi) from the Pacific Ocean.
Age
Sitka spruce is a long-lived tree, with individuals over 700 years old known. Because it grows rapidly under favorable conditions, large size may not indicate exceptional age. The Queets Spruce has been estimated to be only 350 to 450 years old, but adds more than a cubic meter of wood each year (Van Pelt, 2001).
Root system
Living in an extremely wet climate, the Sitka has a shallow root system, [1], with long lateral roots and few branchings.
Fire danger
Fire is not important in Sitka spruce ecology. Their thin bark and shallow root system make them susceptible to fire damage.[5]
Taxonomy
DNA analysis[6][7] has shown that only Picea breweriana has a more basal position than Sitka spruce to the rest of the spruce. The other thirty-three species of spruce are more derived which suggests that Picea originated in North America.
Distribution and habitat
![](http://upload.wikimedia.org/wikipedia/en/thumb/7/70/Sitka_Spruce_forest.jpg/220px-Sitka_Spruce_forest.jpg)
Sitka spruce is native to the west coast of North America, with its northwestern limit on Kodiak Island, Alaska, and its southeastern limit near Fort Bragg in northern California (Griffin & Critchfield 1972). It is closely associated with the temperate rain forests and is found within a few kilometers of the coast in the southern portion of its range. North of Oregon, its range extends inland along river floodplains, but nowhere does its range extend more than 80 km from the Pacific Ocean and its inlets.
Uses
![](http://upload.wikimedia.org/wikipedia/commons/thumb/b/b3/Bucked_Spruce.jpg/220px-Bucked_Spruce.jpg)
Sitka spruce is of major importance in forestry for timber and paper production. Outside of its native range, it is particularly valued for its fast growth on poor soils and exposed sites where few other trees can be grown successfully; in ideal conditions young trees may grow 1.5 m per year. It is naturalized in some parts of Ireland and Great Britain where it was introduced in 1831 (Mitchell, 1978) and New Zealand, though not so extensively as to be considered invasive. Sitka spruce is also planted extensively in Denmark, Norway and Iceland.[8][9] In Norway, Sitka spruce was introduced in the early 1900s. An estimated 50,000 hectares have been planted in Norway, mainly along the coast from Vest-Agder in the south to Troms in the north. It is more tolerant to wind and saline ocean air, and grows faster than the native Norway spruce.[10]
Sitka spruce is used widely in piano, harp, violin, and guitar manufacture, as its high strength-to-weight ratio and regular, knot-free rings make it an excellent conductor of sound. For these reasons, the wood is also an important material for sailing boat spars, aircraft wing spars (including flying models), and the nosecones of Trident missiles.[11] The Wright brothers' Flyer was built using Sitka spruce, as were many aircraft before World War II; during that war, aircraft such as the British Mosquito used it as a substitute for strategically important aluminium.
Newly grown tips of Sitka spruce branches are used to flavour spruce beer and are boiled to make syrup.
The root bark of Sitka spruce trees is used in Native Alaskan basket-weaving designs.
Culture
A unique specimen with golden foliage that used to grow on Haida Gwaii, known as Kiidk'yaas or "The Golden Spruce", is sacred to the Haida Native American people. It was illegally felled in 1997 by Grant Hadwin, although saplings grown from cuttings can now be found near its original site.
Chemistry
The stilbene glucosides astringin, isorhapontin (isorhapontigenin glucoside) and piceid can be found in the bark of Picea sitchensis.[12][13]
See also
References and external links
![](http://upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png)
- ^ Rushforth, Keith (1986) [1980]. Bäume (in German) (2nd ed.). Bern: Hallwag AG. ISBN 3-444-70130-6.
{{cite book}}
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suggested) (help) - ^ "Agathis australis". Conifers. Retrieved 9 April 2012.
- ^ "Tallest Sitka Spruce". Landmark Trees. Retrieved 9 April 2012.
- ^ "Picea sitchensis". Oregon State University. Retrieved 9 April 2012.
- ^ http://www.fs.fed.us/database/feis/plants/tree/picsit/all.html
- ^ Ran, J.-H., Wei, X.-X. & Wang, X.-Q. 2006. Molecular phylogeny and biogeography of Picea (Pinaceae): Implications for phylogeographical studies using cytoplasmic haplotypes. Mol Phylogenet Evol. 41(2): 405–19.
- ^ Sigurgeirsson, A. & Szmidt, A.E. 1993. Phylogenetic and biogeographic implications of chloroplast DNA variation in Picea. Nordic Journal of Botany 13(3): 233–246.
- ^ Dammert, L (2001). Dressing the landscape: afforestation efforts on Iceland, Unasylva Vol. 52, No. 207.
- ^ Hermann, R (1987). North American Tree Species in Europe, Journal of Forestry.
- ^ Sitkagran - utbredelse, egenskaper og anvendelse (Sitka spruce - propagation, properties and uses) by Kjell Vadla, Norwegian Forest and Landscape Institute.
- ^ "Ballistic missile", National Museum of American History
- ^ Stilbene glucosides in the bark of Picea sitchensis. Masakazu Aritomi, Dervilla M.X. Donnelly, Phytochemistry, Volume 15, Issue 12, 1976, Pages 2006–2008, doi:10.1016/S0031-9422(00)88881-0
- ^ Astringin and isorhapontin distribution in Sitka spruce trees. Claudia D. Toscano Underwood and Raymond B. Pearce, Phytochemistry, Volume 30, Issue 7, 1991, Pages 2183–2189, doi:10.1016/0031-9422(91)83610-W
- Template:IUCN2006
- Van Pelt, R. 2001. Forest Giants of the Pacific Coast. University of Washington Press. ISBN 0-295-98140-7.
- Griffin, J. R. & Critchfield, W. B. 1976. Distribution of forest trees in California. USDA Forest Service Research Paper PSW-82: 23–24, 75.
- Mitchell, A. 1978. Trees of Britain & Northern Europe. Collins Field Guide. HarperCollins. London. ISBN 0-00-219213-6
- Gymnosperm Database
- Flora of North America
- Arboretum de Villardebelle - photos of cones of Picea sitchensis and related spruces
- Prof Stephen Sillett's webpage with photos taken during canopy research.
- Description of Sitka Spruce in forestry by US Department of Agriculture.
- the Picea Sitchinesis 'Octopus tree'
- IUCN Red List least concern species
- Use dmy dates from August 2011
- Picea
- Trees of the West Coast (U.S.)
- Trees of California
- Trees of Oregon
- Trees of Washington (state)
- Trees of Alaska
- Trees of British Columbia
- Trees of mild maritime climate
- Least concern flora of California
- Least concern flora of the United States
- Trees of the United States
- Trees of Canada