He studied medicine at the University of Heidelberg, and at the University of Göttingen under Friedrich Gustav Jakob Henle (1809–1885), at Berlin under Rudolf Virchow (1821–1902) and in Würzburg, where he became an assistant to Friedrich Daniel von Recklinghausen (1833–1910). In 1867 he became a lecturer at the University of Marburg, and in 1872 became a full professor of pathology at the University of Giessen, where he succeeded Ludwig Franz Alexander Winther (1812–1871).
- This article is based on a translation of an article at the German Wikipedia, namely: Historischen Lexikon der Schweiz
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