List of herbivorous animals: Difference between revisions

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Can you add humans, other primates, lemurs, bears and badgers as plant-eating animals?
Various substantial edits to "Mammals" section; random other edits; WIP on entire Mammals section. 10th edit; finally created an account
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===[[Squamates]] (lizards, snakes, and worm lizards)===
===[[Squamates]] (lizards, snakes, and worm lizards)===
All snakes and the majority of non-snake Squamates ([[lizards]]) are carnivorous. However, some degree of herbivory is relatively common among lizards. Perhaps 12% of lizards have diets which are >10% herbivorous. Dedicated herbivory, with plants comprising >90% of the diet, occurs in perhaps ~1% of lizards, though estimates vary.<ref>{{cite journal |last1=Cooper Jr|first1=William E.|last2=Vitt|first2=Laurie J.|name-list-style=amp |title=Distribution, extent, and evolution of plant consumption by lizards|journal=Journal of Zoology|volume=257|issue=4|pages=487-517|year=2002|doi=10.1017/S0952836902001085 }}</ref>
All snakes and the majority of non-snake Squamates ([[lizards]]) are carnivorous. However, some degree of herbivory is relatively common among lizards. Perhaps 12% of lizards have diets which are >10% herbivorous. Dedicated herbivory, with plants comprising >90% of the diet, occurs in perhaps ~1% of lizards, though estimates vary.<ref>{{cite journal |last1=Cooper Jr|first1=William E.|last2=Vitt|first2=Laurie J.|name-list-style=amp |title=Distribution, extent, and evolution of plant consumption by lizards|journal=Journal of Zoology|volume=257|issue=4|pages=487-517|year=2002|doi=10.1017/S0952836902001085 }}</ref>
* [[Iguana]]s (family Iguanidae) are overwhelmingly herbivores. Members such as the [[rhinoceros iguana]], [[marine iguana]], [[green iguana]], [[Lesser Antillean iguana]], [[chuckwalla]]s, [[desert iguana]], and [[Galápagos land iguana]] are virtually exclusively herbivorous.
* [[Iguana]]s (family Iguanidae) are overwhelmingly herbivores; members such as the [[rhinoceros iguana]], [[marine iguana]], [[green iguana]], [[Lesser Antillean iguana]], [[chuckwalla]]s, [[desert iguana]], and [[Galápagos land iguana]] are virtually exclusively herbivorous
* Members of the family [[Liolaemidae]] are herbivores.
* Members of the family [[Liolaemidae]] are herbivores
* The [[Panay monitor|Panay]], [[Northern Sierra Madre forest monitor|Northern Sierra Madre forest]], and [[Gray's monitor|Gray's]] [[monitor lizards]] are the only three species of herbivorous [[Varanid]].
* The [[Panay monitor|Panay]], [[Northern Sierra Madre forest monitor|Northern Sierra Madre forest]], and [[Gray's monitor|Gray's]] [[monitor lizards]] are the only three species of herbivorous [[Varanid]]
* Species of the [[Agamid]] genus ''[[Uromastyx]]'', the spiny-tailed lizards, are herbivores.
* Species of the [[Agamid]] genus ''[[Uromastyx]]'', the spiny-tailed lizards, are herbivores
* Certain [[skink]]s; the [[Solomon Islands skink]] is exclusively herbivorous. [[King's skink]] is also primarily a herbivore.<ref>{{cite journal |last1=Arena|first1=P. C.|last2=Wooller|first2=R. D. |name-list-style=amp |title=The reproduction and diet of ''Egernia kingii'' (Reptilia : Scincidae) on Penguin Island, Western Australia|journal=Australian Journal of Zoology|volume=51|issue=5|pages=495–504|year=2008|doi=10.1071/zo02040 }}</ref> The extinct [[Cape Verde giant skink]] was herbivorous.
* Certain [[skink]]s; the [[Solomon Islands skink]] is exclusively herbivorous. [[King's skink]] is also primarily a herbivore.<ref>{{cite journal |last1=Arena|first1=P. C.|last2=Wooller|first2=R. D. |name-list-style=amp |title=The reproduction and diet of ''Egernia kingii'' (Reptilia : Scincidae) on Penguin Island, Western Australia|journal=Australian Journal of Zoology|volume=51|issue=5|pages=495–504|year=2008|doi=10.1071/zo02040 }}</ref> The extinct [[Cape Verde giant skink]] was herbivorous.


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==[[Mammals]]==
==[[Mammals]]==
Herbivory is quite common among [[mammals]], with herbivores occuring across diverse lineages and in a large variety of sizes, [[bauplan|bodyplans]], and niches. Both large and small herbivorous mammals are often important prey species for various predators, though the largest, such as the so-called [[wiktionary:pachyderm|"pachyderms"]], are generally safe from predation as healthy adults. Large, herbivorous mammals came to dominate global ecosystems in the [[Cenozoic]], and, though [[Quaternary extinction event|vastly reduced in number]], they continue to be key features of certain modern ecosystems, such as the [[Maasai Mara]] and [[Kaziranga National Park]].
'''Mammals''' (formally '''Mammalia''') are a [[class (biology)|class]] of [[vertebrate]], air-breathing [[animal]]s whose females are characterized by the possession of [[mammary gland]]s while both males and females are characterized by [[hair]] and/or fur, three [[middle ear]] [[bone]]s used in [[hearing]], and a [[neocortex]] region in the brain. Herbivorous mammals include:

===[[Marsupials]]===
*Most of the extant [[Diprotodonts]], including:
**the extant [[Vombatiformes]] ([[wombats]] and [[koalas]]), as well as many extinct members such as ''[[Diprotodon]]'', ''[[Zygomaturus]]'', and ''[[Palorchestes]]''
**almost all extant [[Macropodiformes]], such as [[kangaroos]], [[wallabies]], [[wallaroo]]s, [[tree kangaroos]], and the [[Potoroidae|Potoroids]] (though this last group also consumes large amounts of fungi and generally some insects)
**most [[Phalangeriformes]] are mostly or wholly herbivorous, including the [[greater glider]], [[honey possum]], and [[common spotted cuscus]]
*A relatively small number of [[Didelphimorphia|Didelphimorphs]] are primarily herbivorous, such as the [[woolly opossum]]s
*The recently extinct entire family of [[Chaeropodidae|pig-footed bandicoots]] (Chaeropodidae, with one genus, ''Chaeropus'') was probably herbivorous
===[[Placental mammal]]s===
*[[Xenarthran]]s
**Certain extinct [[Cingulata|Cingulates]], such as the [[Pampathere]]s and [[Glyptodont]]s
**All living and extinct [[sloths]] (Folivora), including the so-called [[ground sloth]]s
*[[Boreoeutheria|Boreoeutherians]]
**Both species of [[colugo]] (Dermoptera) are herbivores
**The great majority of [[primate]]s:
***The vast majority of [[lemur]]s are largely or exclusively herbivorous, including the [[ring-tailed lemur]], the [[sifaka]]s, the [[indri]], and the [[bamboo lemur]]s{{Sfn|Birkinshaw|Colquhoun|2003|pp=1207–1220}}
***The three species of [[potto]] have diets which are ~90% herbivorous<ref name="Estes1991">{{Cite book | title = The Behavior Guide to African Mammals | url = https://archive.org/details/isbn_0520080858 | url-access = registration | quote = potto. | first = R. D. | last = Estes | author-link = Richard Despard Estes | publisher = University of California Press | year = 1991 | pages = [https://archive.org/details/isbn_0520080858/page/458 458]–464}}</ref>
***[[Nycticebus|Slow lorises]] are majority (and possibly even nearly exclusively) herbivorous, though study findings differ{{Sfn|Nekaris|Bearder|2007|pp=28–33}}
***Some [[Galagidae|bushbabies]]; [[Euoticus|needle-clawed bushbabies]] are gum specialists, and the [[Bioko Allen's bushbaby]] is primarily frugivorous<ref name="Estes1991">{{Cite book | title = The Behavior Guide to African Mammals | url = https://archive.org/details/isbn_0520080858 | url-access = registration | quote = potto. | first = R. D. | last = Estes | author-link = Richard Despard Estes | publisher = University of California Press | year = 1991 | pages = [https://archive.org/details/isbn_0520080858/page/458 458]–464}}</ref>
***The vast majority of [[ape]] and [[monkey]] species (the [[Simiiformes]]) are wholly or primarily herbivorous,<ref name="Milton">{{cite journal |last1=Milton |first1=Katharine |date=6 November 1999 |title=Nutritional Characteristics of Wild Primate Foods: Do the Diets of Our Closest Living Relatives Have Lessons for Us? |url=https://nature.berkeley.edu/miltonlab/pdfs/nutritionalchar.pdf |journal=Nutrition |volume=15 |issue=6 |pages=489 |doi=10.1016/s0899-9007(99)00078-7 |access-date=9 July 2023}}</ref> including:
****The majority of the [[New World monkeys]] (though several taxa consume insects, some as a primary food source); predominantly or wholly herbivorous New World monkeys include the [[spider monkey]]s, [[muriqui]]s, [[woolly monkeys]], [[howler monkey]]s, [[saki monkeys|saki]]s, and [[uakari]]s,<ref>{{cite journal |last1=Rosenberger |first1=Alfred L. |date=August 1992 |title=Evolution of feeding niches in new world monkeys |url=https://onlinelibrary.wiley.com/doi/abs/10.1002/ajpa.1330880408 |journal=American Journal of Biological Anthropology |volume=88 |issue=4 |pages=525-562 |doi=10.1002/ajpa.1330880408 |access-date=9 July 2023}}</ref> among others
****Virtually all [[Cercopithecidae|Old World monkey]]s (''sensu stricto'', excluding apes)<ref>{{cite book |editor=Macdonald, D. |author1=Brandon-Jones, Douglas |author2=Rowell, Thelma E. |name-list-style=amp |year=1984 |title=The Encyclopedia of Mammals |publisher=Facts on File |location=New York |pages=[https://archive.org/details/encyclopediaofma00mals_0/page/370 370–405] |isbn=0-87196-871-1 |url-access=registration |url=https://archive.org/details/encyclopediaofma00mals_0/page/370 }}</ref>:
*****[[Vervet monkeys]]<ref>{{Cite book |author1=Skinner, J.D. |author2=Smithers, R.H.N. |name-list-style=amp |year=1990 |title=The mammals of the southern African subregion |edition=2nd |location=Pretoria (South Africa) |publisher=University of Pretoria |page=771 |isbn=0-86979-802-2}}</ref><ref>{{Cite book |editor=Napier, P.H. |year=1981 |title=Catalogue of primates in the British Museum (Natural History) and elsewhere in the British Isles |chapter=Part II: Family Cercopithecidae, Subfamily Cercopithecinae |location=London |publisher=British Museum (Natural History) |page=203}}</ref>
*****Several [[macaque]]s are overwhelmingly herbivorous, such as the [[Tibetan macaque|Tibetan]]<ref>[http://www.arkive.org/tibetan-macaque/macaca-thibetana/#text=Threats Tibetan macaque videos, photos and facts – Macaca thibetana] {{Webarchive|url=https://web.archive.org/web/20120823153523/http://www.arkive.org/tibetan-macaque/macaca-thibetana/#text=Threats#text=Threats |date=2012-08-23 }}. ARKive (2006-12-22). Retrieved on 2012-08-21.</ref>, [[Japanese macaque|Japanese]], [[Barbary macaque|Barbary]]<ref>{{cite journal |last1=Hanya |first1=Goro |last2=Ménard |first2=Nelly |date=22 February 2011 |title=Dietary adaptations of temperate primates: comparisons of Japanese and Barbary macaques |url=https://link.springer.com/article/10.1007/s10329-011-0239-5 |journal=Primates |volume=52 |pages=192 |doi=10.1007/s10329-011-0239-5 |access-date=9 July 2023}}</ref>, and [[Assam macaque]]s<ref>{{cite journal |last1=Huang |first1=Zhonghao |last2=Huang |first2=Chengming |date=17 September 2014 |title=Dietary adaptations of Assamese macaques (Macaca assamensis) in limestone forests in Southwest China |url=https://onlinelibrary.wiley.com/doi/abs/10.1002/ajp.22320 |journal=American Journal of Primatology |volume=77 |issue=2 |pages=171-185 |doi=10.1002/ajp.22320 |access-date=9 July 2023}}</ref>
*****Although [[baboons]] are famously regarded as omnivores and will occasionally consume animals, studies consistently find species, including the [[yellow baboon|yellow]]<ref>{{cite journal |last1=Post |first1=David G. |date=December 1982 |title=Feeding behavior of yellow baboons (Papio cynocephalusin) the Amboseli National Park, Kenya |url=https://link.springer.com/article/10.1007/BF02693741# |journal=International Journal of Primatology |volume=3 |pages=403-430 |doi=10.1007/BF02693741 |access-date=9 July 2023}}</ref>, [[olive baboon|olive]]<ref>{{cite journal |last1=Swedell |first1=Larissa |last2=Johnson |first2=Caley A. |date=9 December 2011 |title=Feeding ecology of olive baboons (Papio anubis) in Kibale National Park, Uganda: preliminary results on diet and food selection |url=https://onlinelibrary.wiley.com/doi/10.1111/j.1365-2028.2011.01316.x |journal=African Journal of Ecology |volume=50 |issue=3 |pages=367-370 |doi=10.1111/j.1365-2028.2011.01316.x |access-date=9 July 2023}}</ref>, [[Chacma baboon|chacma]]<ref>{{cite journal |last1=Tew |first1=Eleanor |last2=Landman |first2=Marietjie |date=17 April 2018 |title=The contribution of the chacma baboonto seed dispersal in the eastern Karoo,South Africa |url=https://journals.co.za/doi/epdf/10.3957/056.048.023002 |journal=African Journal of Wildlife Research |volume=48 |issue=2 |pages=3 |doi=10.3957/056.048.023002 |access-date=9 July 2023}}</ref>, and [[hamadryas baboon]]s<ref>{{cite journal |last1=Swedell |first1=Larissa |last2=Schreier |first2=Amy |date=8 February 2008 |title=Composition and Seasonality of Diet in Wild Hamadryas Baboons: Preliminary Findings from Filoha |url=https://brill.com/view/journals/ijfp/79/6/article-p476_5.xml?ebody=article%20details |journal=Folia Primatologica |volume=79 |issue=6 |pages=476-490 |doi=10.1159/000164431 |access-date=9 July 2023}}</ref>, to have diets which are ~99% herbivorous
*****[[Mandrill]]<ref>{{cite journal|author1=Tutin, C. E.|author2=Ham, R. M.|author3=White, L. J.|author4=Harrison, M. J.|pmid=9108968|title=The primate community of the Lopé Reserve, Gabon: diets, responses to fruit scarcity, and effects on biomass|year=1997|volume=42|issue=1|pages=1–24|doi=10.1002/(SICI)1098-2345(1997)42:1<1::AID-AJP1>3.0.CO;2-0|journal=American Journal of Primatology|s2cid=37902903}}</ref> and [[Drill (animal)|drill]]<ref>{{cite journal |last1=Owens |first1=Jacob R. |last2=Honarvar |first2=Shaya |date=16 September 2015 |title=From frugivore to folivore: Altitudinal variations in the diet and feeding ecology of the Bioko Island drill (Mandrillus leucophaeus poensis) |url=https://onlinelibrary.wiley.com/doi/abs/10.1002/ajp.22479 |journal=American Journal of Primatology |volume=77 |issue=12 |pages=1270 |doi=10.1002/ajp.22479 |access-date=9 July 2023}}</ref> diets are both >95% herbivorous
*****[[Gelada]]s<ref>{{cite web |url=https://www.britannica.com/animal/gelada |title=Gelada |website=www.britannica.com |publisher=Britannica |access-date=9 July 2023}}</ref>
*****All of the [[Colobinae|Colobines]] are (or are nearly) purely herbivorous<ref>{{cite web |url=https://www.britannica.com/animal/primate-mammal/Diet#ref876307 |title=Primate - Diet |website=www.britannica.com |publisher=Britannica |access-date=9 July 2023}}</ref>, including the [[black-and-white colobus]]es, [[gray langur]]s, [[snub-nosed monkeys]], and the [[proboscis monkey]]
****All wild [[apes]]:
*****[[Gibbon]] (family Hylobatidae) diets are ~90% herbivorous<ref>{{cite journal |last1=Elder |first1=Alice A. |date=1 January 2009 |title=Hylobatid Diets Revisited: The Importance of Body Mass, Fruit Availability, and Interspecific Competition |url=https://link.springer.com/chapter/10.1007/978-0-387-88604-6_8 |journal=The Gibbons |pages=133-159 |doi=10.1007/978-0-387-88604-6_8 |access-date=9 July 2023}}</ref>
*****[[Orangutan]] diets can range from <90% to ~99% herbivorous, depending on various factors<ref>{{cite journal |last1=Wich |first1=S. A. |date=6 December 2006 |title=Dietary and Energetic Responses of Pongo abelii to Fruit Availability Fluctuations |url=https://link.springer.com/article/10.1007/s10764-006-9093-5 |journal=International Journal of Primatology |volume=27 |pages=1544 |doi=10.1007/s10764-006-9093-5 |access-date=9 July 2023}}</ref>

*[[Hominidae]] ([[human]]s)
*[[Hominidae]] ([[human]]s)
*[[Strepsirrhini]] ([[lemur]]s, [[galago]]s, [[potto]]s)
*[[Ungulata]]
*[[Ungulata]]
**[[Odd-toed ungulates]], including [[Equidae|horses, asses, and zebras]], [[rhinoceros]]es, and [[tapir]]s. Like many mammalian herbivores, they are [[hindgut fermenter]]s.
**[[Odd-toed ungulates]], including [[Equidae|horses, asses, and zebras]], [[rhinoceros]]es, and [[tapir]]s. Like many mammalian herbivores, they are [[hindgut fermenter]]s.
**[[Even-toed ungulates]] such as [[Camelidae|camel]]s, [[Suidae|pig]]s, [[hippopotamus]]es, [[Cervidae|deer]], [[bovid]]s, and [[giraffe]]s. Many are [[ruminant]]s, animals with compartmentalized stomachs for processing plant material.
**[[Even-toed ungulates]] such as [[Camelidae|camel]]s, [[Suidae|pig]]s, [[hippopotamus]]es, [[Cervidae|deer]], [[bovid]]s, and [[giraffe]]s. Many are [[ruminant]]s, animals with compartmentalized stomachs for processing plant material.
*[[Paenungulata]] ([[Proboscidea]], [[Sirenia]], [[Hyracoidea]])
*[[Paenungulata]] ([[Proboscidea]], [[Sirenia]], [[Hyracoidea]])
*[[Carnivora]] ([[bear]]s, [[badger]]s)
*[[Carnivora]] ([[bear]]s)
*[[Castorimorpha]] ([[Heteromyidae|kangaroo rat]]s, [[Geomyidae|gopher]]s, [[beaver]]s)
*[[Castorimorpha]] ([[Heteromyidae|kangaroo rat]]s, [[Geomyidae|gopher]]s, [[beaver]]s)
*[[Lagomorpha]] ([[Leporidae]], [[Ochotonidae]])
*[[Lagomorpha]] ([[Leporidae]], [[Ochotonidae]])
*[[Sloths]]
*[[Marsupialia]]
**[[Macropodidae]] ([[kangaroo]], [[wallaby]])
**[[Vombatiformes]] ([[wombats]], [[koala]])


== See also ==
== See also ==

Revision as of 13:06, 9 July 2023

Types of herbivorous feeding strategies
Feeding strategy Diet
Algivores Algae
Frugivores Fruit
Folivores Leaves
Nectarivores Nectar
Nucivores Nuts
Granivores Seeds
Palynivores Pollen
Mucivores Plant fluids, i.e. sap
Xylophages Wood

Herbivores are animals that eat plants. Herbivory is a form of consumption in which a heterotrophic organism consumes other organisms, principally autotrophs[1][page needed] such as plants, algae and photosynthesizing bacteria. More generally, organisms that feed on autotrophs in general are known as 1st level consumers.

Invertebrates

Insects

Herbivory is of extreme ecological importance and prevalence among insects. Perhaps one third (or 500,000) of all described species are herbivores.[2]

Other invertebrates

Fish

Herbivorous fish play a key role in maintaining healthy ecosystems, especially in tropical reefs, where they promote a balance between corals and macroalgae.[15] Herbivorous fish include:

Amphibians

Unusually for tetrapods, herbivory is rare among extant adult lissamphibians. There are, however, many larval and a few adult amphibians which take significant plant matter:

Reptiles

Squamates (lizards, snakes, and worm lizards)

All snakes and the majority of non-snake Squamates (lizards) are carnivorous. However, some degree of herbivory is relatively common among lizards. Perhaps 12% of lizards have diets which are >10% herbivorous. Dedicated herbivory, with plants comprising >90% of the diet, occurs in perhaps ~1% of lizards, though estimates vary.[21]

Chelonians (turtles/tortoises)

Dinosaurs

Mammals

Herbivory is quite common among mammals, with herbivores occuring across diverse lineages and in a large variety of sizes, bodyplans, and niches. Both large and small herbivorous mammals are often important prey species for various predators, though the largest, such as the so-called "pachyderms", are generally safe from predation as healthy adults. Large, herbivorous mammals came to dominate global ecosystems in the Cenozoic, and, though vastly reduced in number, they continue to be key features of certain modern ecosystems, such as the Maasai Mara and Kaziranga National Park.

Marsupials

Placental mammals

See also

References

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  2. ^ Hardy, Nate B.; Kaczvinsky, Chloe; Bird, Gwendolyn; Normark, Benjamin B. (2020-11-02). "What We Don't Know About Diet-Breadth Evolution in Herbivorous Insects". Annual Review of Ecology, Evolution, and Systematics. 51 (1). Annual Reviews: 103–122. doi:10.1146/annurev-ecolsys-011720-023322. ISSN 1543-592X. S2CID 225521141.
  3. ^ "Order Ephemeroptera - Mayflies". www.bugguide.net. Retrieved 23 February 2023.
  4. ^ "Orthoptera". www.genent.cals.ncsu.edu. Retrieved 26 February 2023.
  5. ^ Melissa Barrows. "Buprestidae". ants.biology.utah.edu. Retrieved 24 February 2023.
  6. ^ Kapur, A. P. (10 July 2009). "The Biology and external Morphology of the Larvae of Epilachninae (Coleoptera, Coccinellidae)". Bulletin of Entomological Research. 41 (1): 161–208. doi:10.1017/S0007485300027565. Retrieved 24 February 2023.
  7. ^ "Family Chrysomelidae - Leaf Beetles". www.bugguide.net. Retrieved 12 June 2023.
  8. ^ "Weevils". www.orkincanada.ca. Retrieved 26 February 2023.
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  10. ^ "Suborder Heteroptera - True Bugs". www.bugguide.net. Retrieved 23 February 2023.
  11. ^ "Family Coreidae - Leaf-footed Bugs". www.bugguide.net. Retrieved 23 February 2023.
  12. ^ "Superfamily Lygaeoidea". www.bugguide.net. Retrieved 23 February 2023.
  13. ^ "Family Miridae - Plant Bugs". www.bugguide.net. Retrieved 23 February 2023.
  14. ^ "Superfamily Pentatomoidea". www.bugguide.net. Retrieved 23 February 2023.
  15. ^ a b c d Herbivory in Fish
  16. ^ Lieske, E., and Myers, R. (1999). Coral Reef Fishes. 2nd edition. Princeton University Press. ISBN 0-691-00481-1
  17. ^ What Marine Animals are Herbivores?
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  22. ^ Arena, P. C. & Wooller, R. D. (2008). "The reproduction and diet of Egernia kingii (Reptilia : Scincidae) on Penguin Island, Western Australia". Australian Journal of Zoology. 51 (5): 495–504. doi:10.1071/zo02040.
  23. ^ Discovering Dinosaurs. Curriculum Corporation. 2001. ISBN 9781876973063. Retrieved 2010-08-08.
  24. ^ Birkinshaw & Colquhoun 2003, pp. 1207–1220.
  25. ^ a b Estes, R. D. (1991). The Behavior Guide to African Mammals. University of California Press. pp. 458–464. potto.
  26. ^ Nekaris & Bearder 2007, pp. 28–33.
  27. ^ Milton, Katharine (6 November 1999). "Nutritional Characteristics of Wild Primate Foods: Do the Diets of Our Closest Living Relatives Have Lessons for Us?" (PDF). Nutrition. 15 (6): 489. doi:10.1016/s0899-9007(99)00078-7. Retrieved 9 July 2023.
  28. ^ Rosenberger, Alfred L. (August 1992). "Evolution of feeding niches in new world monkeys". American Journal of Biological Anthropology. 88 (4): 525–562. doi:10.1002/ajpa.1330880408. Retrieved 9 July 2023.
  29. ^ Brandon-Jones, Douglas & Rowell, Thelma E. (1984). Macdonald, D. (ed.). The Encyclopedia of Mammals. New York: Facts on File. pp. 370–405. ISBN 0-87196-871-1.
  30. ^ Skinner, J.D. & Smithers, R.H.N. (1990). The mammals of the southern African subregion (2nd ed.). Pretoria (South Africa): University of Pretoria. p. 771. ISBN 0-86979-802-2.
  31. ^ Napier, P.H., ed. (1981). "Part II: Family Cercopithecidae, Subfamily Cercopithecinae". Catalogue of primates in the British Museum (Natural History) and elsewhere in the British Isles. London: British Museum (Natural History). p. 203.
  32. ^ Tibetan macaque videos, photos and facts – Macaca thibetana Archived 2012-08-23 at the Wayback Machine. ARKive (2006-12-22). Retrieved on 2012-08-21.
  33. ^ Hanya, Goro; Ménard, Nelly (22 February 2011). "Dietary adaptations of temperate primates: comparisons of Japanese and Barbary macaques". Primates. 52: 192. doi:10.1007/s10329-011-0239-5. Retrieved 9 July 2023.
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