Bothidae
| Lefteye flounders | |
|---|---|
| Scaldfish, Arnoglossus laterna | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Actinopterygii |
| Order: | Pleuronectiformes |
| Suborder: | Pleuronectoidei |
| Family: | Bothidae |
| Genera | |
|
Arnoglossus |
|
Lefteye flounders are a family, Bothidae, of flounders. They are called "lefteye flounders" because most species lie on the sea bottom on their right side, with both eyes on the left side. A helpful reminder when trying to recall the family name for this fish is that "Bothidae eyes are on the same side o' dey head." [1] The family is also distinguished by the presence of spines on the snout and near the eyes.[2]
Lefteye flounders vary considerably in size between species, ranging from 4.5 centimetres (1.8 in) to 1.5 metres (4.9 ft) in length. They include such economically important species as the Japanese halibut.[2]
[edit] Species
There are 158 species in twenty genera:[1][3]
- Genus Arnoglossus
- Arnoglossus andrewsi (Kurth, 1954).
- Arabian flounder, Arnoglossus arabicus (Norman, 1939).
- Arnoglossus armstrongi (Scott, 1975).
- Spotless lefteye flounder, Arnoglossus aspilos (Bleeker, 1851).
- Bass Strait flounder, Arnoglossus bassensis (Norman, 1926).
- Arnoglossus boops (Hector, 1875).
- Brown lefteye flounder, Arnoglossus brunneus (Fowler, 1934).
- Cape scaldfish, Arnoglossus capensis (Boulenger, 1898).
- East coast flounder, Arnoglossus dalgleishi (von Bonde, 1922).
- Weak lefteye flounder, Arnoglossus debilis (Gilbert, 1905).
- Long lefteye flounder, Arnoglossus elongatus (Weber, 1913).
- Fison's lefteye flounder, Arnoglossus fisoni (Ogilby, 1898).
- Imperial scaldfish, Arnoglossus imperialis (Rafinesque, 1810).
- Japanese lefteye flounder, Arnoglossus japonicus (Hubbs, 1915).
- Scaldback, Arnoglossus kessleri (Schmidt, 1915).
- Scaldfish, Arnoglossus laterna (Walbaum, 1792).
- Large-crested lefteye flounder, Arnoglossus macrolophus (Alcock, 1889).
- Arnoglossus marisrubri (Klausewitz & Schneider, 1986).
- Arnoglossus micrommatus (Amaoka, Arai & Gomon, 1997).
- Mueller's flounder, Arnoglossus muelleri (Klunzinger, 1872).
- Arnoglossus multirastris (Parin, 1983).
- Arnoglossus nigrifrons (Amaoka & Mihara, 2000).
- Sharp-snout lefteye flounder, Arnoglossus oxyrhynchus (Amaoka, 1969).
- Many-spotted lefteye flounder, Arnoglossus polyspilus (Günther, 1880).
- Rueppell's scaldback, Arnoglossus rueppelii (Cocco, 1844).
- Arnoglossus sayaensis (Amaoka & Imamura, 1990).
- Witch, Arnoglossus scapha (Forster, 1801).
- Arnoglossus septemventralis (Amaoka & Mihara, 2000).
- Arnoglossus tapeinosoma (Bleeker, 1865).
- Dwarf lefteye flounder, Arnoglossus tenuis (Günther, 1880).
- Thor's scaldfish, Arnoglossus thori (Kyle, 1913).
- Waite's lefteye flounder, Arnoglossus waitei (Norman, 1926).
- Arnoglossus yamanakai (Fukui, Yamada & Ozawa, 1988).
- Genus Asterorhombus
- Asterorhombus annulatus (Weber, 1913).
- Bleeker's flounder, Asterorhombus bleekeri (Macleay, 1881).
- Angler flatfish, Asterorhombus fijiensis (Norman, 1931).
- Intermediate flounder, Asterorhombus intermedius (Bleeker, 1865).
- Asterorhombus osculus (Amaoka & Arai, 1998).
- Genus Bothus
- Bothus assimilis (Günther, 1862).
- Pacific eyed flounder, Bothus constellatus (Jordan, 1889).
- Bothus ellipticus (Poey, 1860).
- Guinean flounder, Bothus guibei (Stauch, 1966).
- Pacific leopard flounder, Bothus leopardinus (Günther, 1862).
- Plate fish, Bothus lunatus (Linnaeus, 1758).
- Mottled flounder, Bothus maculiferus (Poey, 1860).
- Flowery flounder, Bothus mancus (Broussonet, 1782).
- St. Helena flounder, Bothus mellissi (Norman, 1931).
- Indo-Pacific oval flounder, Bothus myriaster (Temminck & Schlegel, 1846).
- Eyed flounder, Bothus ocellatus (Günther, 1880).
- Leopard flounder, Bothus pantherinus (Rüppell, 1830).
- Wide-eyed flounder, Bothus podas (Delaroche, 1809).
Wide-eyed flounder, Bothus podas - Twospot flounder, Bothus robinsi (Topp & Hoff, 1972).
- Bothus swio (Hensley, 1997).
- Bothus tricirrhitus (Kotthaus, 1977).
- Genus Chascanopsetta
- Chascanopsetta crumenalis (Gilbert & Cramer, 1897).
- Chascanopsetta elski (Foroshchuk, 1991).
- Chascanopsetta kenyaensis (Hensley & Smale, 1997).
- Pelican flounder, Chascanopsetta lugubris (Alcock, 1894).
- Chascanopsetta megagnatha (Amaoka & Parin, 1990).
- Chascanopsetta micrognatha (Amaoka & Yamamoto, 1984).
- Chascanopsetta prognatha (Norman, 1939).
- Chascanopsetta prorigera (Gilbert, 1905).
- Genus Crossorhombus
- Blue flounder, Crossorhombus azureus (Alcock, 1889).
- Lord Howe Island flounder, Crossorhombus howensis (Hensley & Randall, 1993).
- Crossorhombus kanekonis (Tanaka, 1918).
- Kobe flounder, Crossorhombus kobensis (Jordan & Starks, 1906).
- Broadbrow flounder, Crossorhombus valderostratus (Alcock, 1890).
- Genus Engyophrys
- Speckled-tail flounder, Engyophrys sanctilaurentii (Jordan & Bollman, 1890).
- American spiny flounder, Engyophrys senta (Ginsburg, 1933).
- Genus Engyprosopon
- Engyprosopon arenicola (Jordan & Evermann, 1903).
- Bellona flounder, Engyprosopon bellonaensis (Amaoka, Mihara & Rivaton, 1993).
- Cocos Island flounder, Engyprosopon cocosensis (Bleeker, 1855).
- Engyprosopon filimanus (Li & Wang, 1982).
- Engyprosopon filipennis (Wu & Tang, 1935).
- Largescale flounder, Engyprosopon grandisquama (Temminck & Schlegel, 1846).
- Engyprosopon hawaiiensis (Jordan & Evermann, 1903).
- Engyprosopon hensleyi (Amaoka & Imamura, 1990).
- Hureau's flounder, Engyprosopon hureaui (Quéro & Golani, 1990).
- Engyprosopon latifrons (Regan, 1908).
- Engyprosopon longipelvis (Amaoka, 1969).
- Long pectoral fin flounder, Engyprosopon longipterum (Amaoka, Mihara & Rivaton, 1993).
- Engyprosopon macrolepis (Regan, 1908).
- Olive wide-eyed flounder, Engyprosopon maldivensis (Regan, 1908).
- Engyprosopon marquisensis (Amaoka & Séret, 2005).[4]
- Engyprosopon mogkii (Bleeker, 1854).
- Engyprosopon multisquama (Amaoka, 1963).
- Natal flounder, Engyprosopon natalensis (Regan, 1920).
- Engyprosopon obliquioculatum (Fowler, 1934).
- Engyprosopon raoulensis (Amaoka & Mihara, 1995).
- Engyprosopon regani (Hensley & Suzumoto, 1990).
- Long snout flounder, Engyprosopon rostratum (Amaoka, Mihara & Rivaton, 1993).
- Engyprosopon sechellensis (Regan, 1908).
- Seven pelvic ray flounder, Engyprosopon septempes (Amaoka, Mihara & Rivaton, 1993).
- Engyprosopon vanuatuensis (Amaoka & Séret, 2005).
- Engyprosopon xenandrus (Gilbert, 1905).
- Engyprosopon xystrias (Hubbs, 1915).
- Genus Grammatobothus
- Krempf's flounder, Grammatobothus krempfi (Chabanaud, 1929).
- Pennant flounder, Grammatobothus pennatus (Ogilby, 1913).
- Threespot flounder, Grammatobothus polyophthalmus (Bleeker, 1865).
- Genus Japonolaeops
- Japonolaeops dentatus (Amaoka, 1969).
- Genus Kamoharaia
- Wide-mouthed flounder, Kamoharaia megastoma (Kamohara, 1936).
- Genus Laeops
- Clear fin-base flounder, Laeops clarus (Fowler, 1934).
- Laeops cypho (Fowler, 1934).
- Philippine slender flounder, Laeops gracilis (Fowler, 1934).
- Günther's flounder, Laeops guentheri (Alcock, 1890).
- Laeops kitaharae (Smith & Pope, 1906).
- Laeops macrophthalmus (Alcock, 1889).
- Khaki flounder, Laeops natalensis (Regan, 1920).
- Laeops nigrescens (Lloyd, 1907).
- Blackspotted flounder, Laeops nigromaculatus (von Bonde, 1922).
- Small headed flounder, Laeops parviceps (Günther, 1880).
- Longarm flounder, Laeops pectoralis (von Bonde, 1922).
- Laeops tungkongensis (Chen & Weng, 1965).
- Genus Lophonectes
- Crested flounder, Lophonectes gallus (Günther, 1880).
- Lophonectes mongonuiensis (Regan, 1914).
- Genus Monolene
- Slim flounder, Monolene antillarum (Norman, 1933).
- Asaedae flounder, Monolene asaedai (Clark, 1936).
- Monolene atrimana (Goode & Bean, 1886).
- Monolene danae (Bruun, 1937).
- Acapulco flounder, Monolene dubiosa (Garman, 1899).
- Monolene helenensis (Amaoka & Imamura, 2000).
- Pacific deepwater flounder, Monolene maculipinna (Garman, 1899).
- Monolene megalepis Woods, 1961.
- Merten's moonflounder, Monolene mertensi (Poll, 1959).
- Smallmouth moonflounder, Monolene microstoma (Kuronuma, 1940).
- Deepwater flounder, Monolene sessilicauda (Goode, 1880).
Deepwater flounder, Monolene sessilicauda
- Genus Neolaeops
- Crosseyed flounder, Neolaeops microphthalmus (von Bonde, 1922).
- Genus Parabothus
- Parabothus amaokai (Parin, 1983).
- Parabothus budkeri (Chabanaud, 1943).
- Parabothus chlorospilus (Gilbert, 1905).
- Parabothus coarctatus (Gilbert, 1905).
- Parabothus filipes (Amaoka, Mihara & Rivaton, 1997).
- Parabothus kiensis (Tanaka, 1918).
- Parabothus malhensis (Regan, 1908).
- Manyscaled flounder, Parabothus polylepis (Alcock, 1889).
- Parabothus taiwanensis (Amaoka & Shen, 1993).
- Genus Perissias
- Striped-fin flounder, Perissias taeniopterus (Gilbert, 1890).
- Genus Psettina
- Psettina brevirictis (Alcock, 1890).
- Psettina filimana (Li & Wang, 1982).
- Rough-scaled flounder, Psettina gigantea (Amaoka, 1963).
- Psettina hainanensis (Wu & Tang, 1935).
- Psettina iijimae (Jordan & Starks, 1904).
- Psettina multisquamea (Fedorov & Foroshchuk, 1988).
- Psettina profunda (Weber, 1913).
- Psettina senta (Ginsburg, 1933).
- Psettina tosana (Amaoka, 1963).
- Psettina variegata (Fowler, 1934).
- Genus Taeniopsetta
- Indo-Pacific ocellated flounder, Taeniopsetta ocellata (Günther, 1880).
- Taeniopsetta radula (Gilbert, 1905).
- Genus Tosarhombus
- Tosarhombus brevis (Amaoka, Mihara & Rivaton, 1997).
- Tosarhombus longimanus (Amaoka, Mihara & Rivaton, 1997).
- Tosarhombus neocaledonicus (Amaoka & Rivaton, 1991).
- Tosarhombus nielseni (Amaoka & Rivaton, 1991).
Sash flounder, Trichopsetta ventralis - Tosarhombus octoculatus (Amaoka, 1969).
- Tosarhombus smithi (Nielsen, 1964).
- Genus Trichopsetta
- Caribbean flounder, Trichopsetta caribbaea (Anderson & Gutherz, 1967).
- Trichopsetta melasma (Anderson & Gutherz, 1967).
- Trichopsetta orbisulcus (Anderson & Gutherz, 1967).
- Sash flounder, Trichopsetta ventralis (Goode & Bean, 1885).
[edit] References
- ^ a b Froese, Rainer, and Daniel Pauly, eds. (2006). "Bothidae" in FishBase. February 2006 version.
- ^ a b Chapleau, Francois & Amaoka, Kunio (1998). Paxton, J.R. & Eschmeyer, W.N.. ed. Encyclopedia of Fishes. San Diego: Academic Press. pp. 224–225. ISBN 0-12-547665-5.
- ^ "Bothidae". Integrated Taxonomic Information System. http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=172714. Retrieved 2 April 2006.
- ^ Kunio Amaoka and Bernard Séret (2005). "Engyprosopon marquisensis, a new species of bothid flounder (Pleuronectiformes: Bothidae) from the Marquesas Islands (French Polynesia)". Ichthyological Research 52 (4): 373–378. doi:10.1007/s10228-005-0299-x.