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{{dablink|For the band Camel, see [[Camel (band)]].}}
'''Camels''' are [[even-toed ungulates]] within the [[genus]] '''''Camelus'''''. The [[dromedary]], one-humped or Arabian camel has a single hump, and the [[Bactrian camel]] has two humps. They are native to the dry [[desert]] areas of western [[Asia]], and central and east Asia, respectively.

The life expectancy of a camel is sixty to eighty years. The term ''camel'' is also used more broadly to describe any of the six camel-like creatures in the family [[Camelid]]ae: the two true camels, and the four South American camelids, the [[llama]], [[alpaca]], [[guanaco]], and [[vicuña]].

A fully-grown adult camel stands {{convert|1.85|m|ftin|abbr=on}} at the shoulder and {{convert|2.15|m|ftin|abbr=on}} at the hump. The hump rises about thirty inches out of its body. Camels can run up to {{convert|65|km/h|mi/h|0|abbr=on}} in short bursts and sustain speeds of up to {{convert|40|km/h|mi/h|0|abbr=on}}.
Fossil evidence indicates that the ancestors of modern camels evolved in [[North America]] during the [[Palaeogene]] period, and later spread to Asia. Humans first domesticated camels between 3,500–3,000 years ago. The dromedary and the Bactrian camel are both still used for milk, meat, and as [[working animal|beasts of burden]]—the dromedary in western Asia, and the Bactrian camel further to the north and east in central Asia.

== Distribution and numbers ==
[[Image:2003camel.PNG|thumb|right|Camel headcount in 2003.]]
The almost 14 million [[Dromedary|dromedaries]] alive today are domesticated animals (mostly living in [[Somalia]], [[Sudan]], [[Mauritania]] and nearby countries).

The [[Bactrian camel]] is now reduced to an estimated 1.4 million animals, mostly domesticated. It is thought that there are about 1000 wild Bactrian camels in the [[Gobi Desert]] in China and Mongolia. <ref>[http://www.animalinfo.org/species/artiperi/camebact.htm Wild Bactrian Camel], Animal Info</ref>

There is a substantial [[feral]] population of dromedaries estimated at up to 700,000 in central parts of [[Australia]], descended from individuals introduced as means of transport in the 19th century and early 20th century. This population is growing at approximately 11% per year. The government of [[South Australia]] has decided to cull the animals using aerial marksmen, because the camels use too much of the limited resources needed by [[sheep]] farmers. For more information, see [[Australian feral camel]].

A small population of introduced camels, dromedaries and Bactrians, survived in the [[Southwest United States]] until the 1900s. These animals, imported from Turkey, were part of the [[US Camel Corps]] experiment and used as draft animals in mines and escaped or were released after the project was terminated. A descendant of one of these was seen by a backpacker in [[Los Padres National Forest]] in 1972. Twenty-three Bactrian camels were brought to Canada during the [[Cariboo Gold Rush]].

==Camel hybrids==
{{Unreferencedsection|date=February 2008}}
[[Image:Désert-du-Thar.jpg|left|thumb|Camels being used in [[Thar Desert]] ]]Throughout their domesticated history, Camels have been used as a means of transportation in arid regions like [[Egypt]]. Bactrian camels have two humps and are rugged cold-climate camels, while dromedaries have one hump and are warmer climate dwellers. Dromedary-Bactrian hybrids are called bukhts, are larger than either parent, have a single hump and are good draft camels. The females can be mated back to a Bactrian to produce ¾-bred riding camels. These hybrids are found in Kazakhstan.

The [[Cama (animal)|cama]] is a camel/[[llama]] hybrid bred by scientists who wanted to see how closely related the parent species were. The dromedary is six times the weight of a llama, hence artificial insemination was required to impregnate the llama female (llama male to dromedary female attempts have proven unsuccessful). Though born even smaller than a llama cria, the cama had the short ears and long tail of a camel, no hump and llama-like [[cloven-hoof|cloven hooves]] rather than the dromedary-like pads. At four years old, the cama became sexually mature and interested in llama and guanaco females. A second cama (female) has since been produced using artificial insemination. Because camels and llamas both have 74 [[chromosome]]s, scientists hope that the cama will be fertile. If so, there is potential for increasing size, meat/wool yield and pack/draft ability in South American camels. The cama apparently inherited the poor temperament of both parents as well as demonstrating the relatedness of the New World and Old World camelids.

== Eco-behavioural adaptations ==
[[Image:Cameljordaniandesert.jpg|right|thumb|''Camelus dromedarius'', [[Wadi Rum]], [[Jordan]].]]
[[Image:Camels in Dubai 2.jpg|right|thumb|Domesticated camel calves in [[Dubai]].]]
[[Image:GntCamelsOnRide.jpg|right|thumb|Exhibition camels munching [[neem]] leaves on a street at [[Guntur City]], [[India]].]]
[[Image:Camelus dromedarius in Singapore Zoo.JPG|right|thumb|''Camelus dromedarius'' in the Singapore Zoo.]]
Camels do not store water in their humps as is commonly believed; they are actually a reservoir of fatty tissue. When this tissue is metabolized, it acts as a source of energy, and would yield more than 1&nbsp;g of water for each 1&nbsp;g of fat converted through reaction with oxygen from air. This process of fat metabolization generates a net loss of water through respiration for the oxygen required to convert the fat.<ref>[http://www.biol.lu.se/zoofysiol/Djurartiklar/Kamel.html What secrets lie within the camel's hump?], [[Lund University]], [[Sweden]], Accessed 7 January 2008</ref>

Their ability to withstand long periods without water is due to a series of physiological adaptations. Their [[red blood cell]]s have an oval shape, unlike those of other [[mammals]], which are circular. This is to facilitate their flow in a [[dehydration|dehydrated]] state. These cells are also more stable<ref>[http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=Retrieve&db=PubMed&list_uids=816376&dopt=Abstract Unique properties of the camel erythrocyte membrane], [[National Center for Biotechnology Information|NCBI]], 5 April 1975</ref> in order to withstand high [[osmosis|osmotic]] variation without rupturing when drinking large amounts of water ({{convert|100|l|0|USgal}} to {{convert|150|l|0|USgal}} in one drink).<ref>[http://www.zoo-hannover.de/zoo-hannover/en/zoo_v3/tiere_attraktionen/tiere_az/tiere_detail_726.html Dromedary], Hannover Zoo, Accessed 8 January 2008</ref>

Camels are able to withstand changes in [[body temperature]] and water content that would kill most other animals. Their temperature ranges from {{convert|34|°C|°F|0|abbr=on}} at night up to {{convert|41|°C|°F|0|abbr=on}} during the day, and only above this threshold will they begin to sweat. The upper body temperature range is often not reached during the day in milder climatic conditions, and therefore, the camel may not sweat at all during the day. Evaporation of their sweat takes place at the skin level, not at the surface of their coat, thereby being very efficient at cooling the body compared to the amount of water lost through sweating. This ability to fluctuate body temperature and the efficiency of their sweating allows them to preserve about five litres of water a day. {{Fact|date=December 2007}}

A feature of their nostrils is that a large amount of water vapor in their exhalations is trapped and returned to their body fluids, thereby reducing the amount of water lost through respiration.

They can withstand at least 20-25% weight loss due to sweating (most mammals can only withstand about 3-4% dehydration before cardiac failure results from the thickened blood). A camel's blood remains hydrated, even though the body fluids are lost, until this 25% limit is reached.

Camels eating green herbage can ingest sufficient moisture in milder conditions to maintain their body's hydrated state without the need for drinking.

A camel's thick coat reflects sunlight. A shorn camel has to sweat 50% more to avoid overheating. It also insulates them from the intense heat that radiates from the desert sand. Their long legs help by keeping them further from the hot ground. Camels have been known to swim.

Their mouth is very sturdy, able to chew thorny desert plants. Long eyelashes and ear hairs, together with sealable nostrils, form a barrier against sand. Their gait (moving both legs on one side at the same time) and their widened feet help them move without sinking into the sand.

The [[kidney]]s and [[intestines]] of a camel are very efficient at retaining water. Urine comes out as a thick syrup, and their faeces are so dry that they can fuel fires.

All camelids have an unusual immune system. In other species, the Y-shaped [[antibody]] molecules consist of two heavy (or long) chains along the length of the Y, and two light (or short) chains at each tip of the Y. In the camel, the antibody molecules have only two heavy chains, which makes them smaller and more durable. These antibodies, which were discovered in 1993, probably developed 50 million years ago, after camelids split from ruminants and pigs, according to biochemist Serge Muyldermans.<ref>'Camelized' antibodies make waves, news story, Robert Koenig, Science, 318:1373, 30 November 2007.</ref>

The camel is the only animal to have replaced the wheel (mainly in North Africa) where the wheel had already been established. The camel did not lose that distinction until the wheel was combined with the internal combustion engine in the 20th century.

==Camel farming==
{{main|Camel farming in Sudan}}

==Military uses of camels==
[[Image:Camel corps at Magdhaba.jpg|left|thumb|English Imperial Camel Corps Brigade in Egypt]]
{{sect-stub}}
{{main|Camel cavalry}}
Attempts have been made to employ camels as [[cavalry]] and [[dragoon]] mounts and as freight animals in lieu of horses and mules. In some places, such as [[Australia]], some of the camels have become [[feral]] and are considered to be dangerous to travelers on camels. The camels were mostly used in combat because of their ability to scare off horses in close ranges, a quality famously employed by the [[Achaemenid]] Persians when fighting [[Lydia]], although the Persians usually used camels as baggage trains for arrows and equipment. The horses detest the smell of camels, and therefore, the horses in the vicinity become harder to control. The United States Army had an active [[U.S. Camel Corps|camel corps]] stationed in California in the 19th century, and the brick stables may still be seen at the [[Benicia Arsenal]] in [[Benicia, California]], now converted to artists' and artisans' studio spaces. Camels have been used in wars throughout Africa, and also in the East Roman Empire as auxiliary forces known as [[Dromedarii]] recruited in desert provinces.
{{clear}}

== Cuisine ==
===Dairy===
[[Image:Camel portrait.jpg|thumb|Fur on a camel]]
Camel milk is a staple food of desert nomad tribes and is richer in fat and protein than cow milk. Camel milk cannot be made into [[butter]] in the traditional churning method. It can be made if it is soured first, churned, and then a clarifying agent is added, or if it is churned at 24&ndash;25&nbsp;°C (75&ndash;76&nbsp;°F), but times will vary greatly in achieving results. The milk can readily be made into [[yogurt]]. Butter or yogurt made from camel milk is said to have a very faint greenish tinge. Camel milk is said to have many healthful properties and is used as a medicinal product in [[India]]; [[Bedouin]] tribes believe that camel milk has great curative powers if the camel's diet consists of certain plants. In [[Ethiopia]], the milk is considered an [[aphrodisiac]].

Camel milk, until recently, was impossible to make into traditional cheese since [[rennet]] was unable to coagulate the milk proteins to allow the collection of [[curd|curds]]. Under the commission of the [[Food and Agriculture Organization|FAO]], Professor J.P. Ramet of the [[École Nationale Supérieure d'Agronomie et des Industries Alimentaires]] (ENSAIA) was able to produce curdling by the addition of [[calcium phosphate]] and vegetable rennet.<ref>[http://www.fao.org/news/2001/010701-e.htm Fresh from your local drome'dairy'?] [[Food and Agriculture Organization]], July 6, 2001</ref> The cheese produced from this process has low levels of cholesterol and lactose. The sale of camel cheese is limited due to the low yield of cheese from milk and the uncertainty of pasteurization levels for camel milk which makes adherence to dairy import regulations difficult.

===Meat===
A camel carcass can provide a substantial amount of meat. The male dromedary carcass can weigh {{convert|400|kg|lb|0|abbr=on|sigfig=1}} or more, while the carcass of a male Bactrian can weigh up to {{convert|650|kg|lb|0|abbr=on|sigfig=2}}. The carcass of a female camel weighs less than the male, ranging between 250 and 350&nbsp;kg (550&ndash;770&nbsp;lb), but can provide a substantial amount of meat. The brisket, ribs and loin are among the preferred parts, but the hump is considered a delicacy and is most favored. It is reported that camel meat tastes like coarse beef, but older camels can prove to be tough and less flavorful.

Camel meat has been eaten for centuries. It has been recorded by [[Ancient Greece|ancient Greek]] writers as an available dish in ancient [[Persia]] at banquets, usually roasted whole. The ancient [[Ancient Rome|Roman]] emperor [[Heliogabalus]] enjoyed camel's heel. Camel meat is still eaten in certain regions including [[Somalia]], where it is called ''Hilib geyl'', [[Saudi Arabia]], [[Egypt]], [[Libya]], [[Sudan]], [[Kazakhstan]] and other arid regions where alternative forms of protein may be limited or where camel meat has had a long cultural history. In the Middle East, camel meat is the rarest and most prized source of [[pastırma]]. Not just the meat, but also blood is a consumable item as is the case in northern [[Kenya]], where camel blood is a source of [[iron]], [[vitamin D]], salts and minerals (although [[Halal|Muslims do not drink or consume blood products]]). According to [[Jewish]] tradition, [[Taboo food and drink#Camels|camel meat]] and milk are taboo. Camels possess only one of the two [[kosher foods|Kosher criteria]]; although they [[Ruminant|chew their cuds]], they do not possess split hooves. (See: [[Taboo food and drink]])

===Health issues===
A 2005 report issued jointly by the Saudi Ministry of Health and the United States Center for Disease Controls details cases of human [[bubonic plague]] resulting from the ingestion of raw camel liver.<ref>[http://www.cdc.gov/ncidod/EID/vol11no09/05-0081.htm Plague from Eating Raw Camel Liver], [[Centers for Disease Control and Prevention]], Vol 11, No. 9, September 2005</ref>

Revision as of 21:06, 16 March 2008

Camels are even-toed ungulates within the genus Camelus. The dromedary, one-humped or Arabian camel has a single hump, and the Bactrian camel has two humps. They are native to the dry desert areas of western Asia, and central and east Asia, respectively.

The life expectancy of a camel is sixty to eighty years. The term camel is also used more broadly to describe any of the six camel-like creatures in the family Camelidae: the two true camels, and the four South American camelids, the llama, alpaca, guanaco, and vicuña.

A fully-grown adult camel stands 1.85 m (6 ft 1 in) at the shoulder and 2.15 m (7 ft 1 in) at the hump. The hump rises about thirty inches out of its body. Camels can run up to 65 km/h (40 mph) in short bursts and sustain speeds of up to 40 km/h (25 mph).

Fossil evidence indicates that the ancestors of modern camels evolved in North America during the Palaeogene period, and later spread to Asia. Humans first domesticated camels between 3,500–3,000 years ago. The dromedary and the Bactrian camel are both still used for milk, meat, and as beasts of burden—the dromedary in western Asia, and the Bactrian camel further to the north and east in central Asia.

Distribution and numbers

File:2003camel.PNG
Camel headcount in 2003.

The almost 14 million dromedaries alive today are domesticated animals (mostly living in Somalia, Sudan, Mauritania and nearby countries).

The Bactrian camel is now reduced to an estimated 1.4 million animals, mostly domesticated. It is thought that there are about 1000 wild Bactrian camels in the Gobi Desert in China and Mongolia. [1]

There is a substantial feral population of dromedaries estimated at up to 700,000 in central parts of Australia, descended from individuals introduced as means of transport in the 19th century and early 20th century. This population is growing at approximately 11% per year. The government of South Australia has decided to cull the animals using aerial marksmen, because the camels use too much of the limited resources needed by sheep farmers. For more information, see Australian feral camel.

A small population of introduced camels, dromedaries and Bactrians, survived in the Southwest United States until the 1900s. These animals, imported from Turkey, were part of the US Camel Corps experiment and used as draft animals in mines and escaped or were released after the project was terminated. A descendant of one of these was seen by a backpacker in Los Padres National Forest in 1972. Twenty-three Bactrian camels were brought to Canada during the Cariboo Gold Rush.

Camel hybrids

Camels being used in Thar Desert

Throughout their domesticated history, Camels have been used as a means of transportation in arid regions like Egypt. Bactrian camels have two humps and are rugged cold-climate camels, while dromedaries have one hump and are warmer climate dwellers. Dromedary-Bactrian hybrids are called bukhts, are larger than either parent, have a single hump and are good draft camels. The females can be mated back to a Bactrian to produce ¾-bred riding camels. These hybrids are found in Kazakhstan.

The cama is a camel/llama hybrid bred by scientists who wanted to see how closely related the parent species were. The dromedary is six times the weight of a llama, hence artificial insemination was required to impregnate the llama female (llama male to dromedary female attempts have proven unsuccessful). Though born even smaller than a llama cria, the cama had the short ears and long tail of a camel, no hump and llama-like cloven hooves rather than the dromedary-like pads. At four years old, the cama became sexually mature and interested in llama and guanaco females. A second cama (female) has since been produced using artificial insemination. Because camels and llamas both have 74 chromosomes, scientists hope that the cama will be fertile. If so, there is potential for increasing size, meat/wool yield and pack/draft ability in South American camels. The cama apparently inherited the poor temperament of both parents as well as demonstrating the relatedness of the New World and Old World camelids.

Eco-behavioural adaptations

Camelus dromedarius, Wadi Rum, Jordan.
Domesticated camel calves in Dubai.
Exhibition camels munching neem leaves on a street at Guntur City, India.
Camelus dromedarius in the Singapore Zoo.

Camels do not store water in their humps as is commonly believed; they are actually a reservoir of fatty tissue. When this tissue is metabolized, it acts as a source of energy, and would yield more than 1 g of water for each 1 g of fat converted through reaction with oxygen from air. This process of fat metabolization generates a net loss of water through respiration for the oxygen required to convert the fat.[2]

Their ability to withstand long periods without water is due to a series of physiological adaptations. Their red blood cells have an oval shape, unlike those of other mammals, which are circular. This is to facilitate their flow in a dehydrated state. These cells are also more stable[3] in order to withstand high osmotic variation without rupturing when drinking large amounts of water (100 litres (22 imp gal; 26 US gal)* to 150 litres (33 imp gal; 40 US gal)* in one drink).[4]

Camels are able to withstand changes in body temperature and water content that would kill most other animals. Their temperature ranges from 34 °C (93 °F) at night up to 41 °C (106 °F) during the day, and only above this threshold will they begin to sweat. The upper body temperature range is often not reached during the day in milder climatic conditions, and therefore, the camel may not sweat at all during the day. Evaporation of their sweat takes place at the skin level, not at the surface of their coat, thereby being very efficient at cooling the body compared to the amount of water lost through sweating. This ability to fluctuate body temperature and the efficiency of their sweating allows them to preserve about five litres of water a day. [citation needed]

A feature of their nostrils is that a large amount of water vapor in their exhalations is trapped and returned to their body fluids, thereby reducing the amount of water lost through respiration.

They can withstand at least 20-25% weight loss due to sweating (most mammals can only withstand about 3-4% dehydration before cardiac failure results from the thickened blood). A camel's blood remains hydrated, even though the body fluids are lost, until this 25% limit is reached.

Camels eating green herbage can ingest sufficient moisture in milder conditions to maintain their body's hydrated state without the need for drinking.

A camel's thick coat reflects sunlight. A shorn camel has to sweat 50% more to avoid overheating. It also insulates them from the intense heat that radiates from the desert sand. Their long legs help by keeping them further from the hot ground. Camels have been known to swim.

Their mouth is very sturdy, able to chew thorny desert plants. Long eyelashes and ear hairs, together with sealable nostrils, form a barrier against sand. Their gait (moving both legs on one side at the same time) and their widened feet help them move without sinking into the sand.

The kidneys and intestines of a camel are very efficient at retaining water. Urine comes out as a thick syrup, and their faeces are so dry that they can fuel fires.

All camelids have an unusual immune system. In other species, the Y-shaped antibody molecules consist of two heavy (or long) chains along the length of the Y, and two light (or short) chains at each tip of the Y. In the camel, the antibody molecules have only two heavy chains, which makes them smaller and more durable. These antibodies, which were discovered in 1993, probably developed 50 million years ago, after camelids split from ruminants and pigs, according to biochemist Serge Muyldermans.[5]

The camel is the only animal to have replaced the wheel (mainly in North Africa) where the wheel had already been established. The camel did not lose that distinction until the wheel was combined with the internal combustion engine in the 20th century.

Camel farming

Military uses of camels

English Imperial Camel Corps Brigade in Egypt

Attempts have been made to employ camels as cavalry and dragoon mounts and as freight animals in lieu of horses and mules. In some places, such as Australia, some of the camels have become feral and are considered to be dangerous to travelers on camels. The camels were mostly used in combat because of their ability to scare off horses in close ranges, a quality famously employed by the Achaemenid Persians when fighting Lydia, although the Persians usually used camels as baggage trains for arrows and equipment. The horses detest the smell of camels, and therefore, the horses in the vicinity become harder to control. The United States Army had an active camel corps stationed in California in the 19th century, and the brick stables may still be seen at the Benicia Arsenal in Benicia, California, now converted to artists' and artisans' studio spaces. Camels have been used in wars throughout Africa, and also in the East Roman Empire as auxiliary forces known as Dromedarii recruited in desert provinces.

Cuisine

Dairy

Fur on a camel

Camel milk is a staple food of desert nomad tribes and is richer in fat and protein than cow milk. Camel milk cannot be made into butter in the traditional churning method. It can be made if it is soured first, churned, and then a clarifying agent is added, or if it is churned at 24–25 °C (75–76 °F), but times will vary greatly in achieving results. The milk can readily be made into yogurt. Butter or yogurt made from camel milk is said to have a very faint greenish tinge. Camel milk is said to have many healthful properties and is used as a medicinal product in India; Bedouin tribes believe that camel milk has great curative powers if the camel's diet consists of certain plants. In Ethiopia, the milk is considered an aphrodisiac.

Camel milk, until recently, was impossible to make into traditional cheese since rennet was unable to coagulate the milk proteins to allow the collection of curds. Under the commission of the FAO, Professor J.P. Ramet of the École Nationale Supérieure d'Agronomie et des Industries Alimentaires (ENSAIA) was able to produce curdling by the addition of calcium phosphate and vegetable rennet.[6] The cheese produced from this process has low levels of cholesterol and lactose. The sale of camel cheese is limited due to the low yield of cheese from milk and the uncertainty of pasteurization levels for camel milk which makes adherence to dairy import regulations difficult.

Meat

A camel carcass can provide a substantial amount of meat. The male dromedary carcass can weigh 400 kg (882 lb) or more, while the carcass of a male Bactrian can weigh up to 650 kg (1,433 lb). The carcass of a female camel weighs less than the male, ranging between 250 and 350 kg (550–770 lb), but can provide a substantial amount of meat. The brisket, ribs and loin are among the preferred parts, but the hump is considered a delicacy and is most favored. It is reported that camel meat tastes like coarse beef, but older camels can prove to be tough and less flavorful.

Camel meat has been eaten for centuries. It has been recorded by ancient Greek writers as an available dish in ancient Persia at banquets, usually roasted whole. The ancient Roman emperor Heliogabalus enjoyed camel's heel. Camel meat is still eaten in certain regions including Somalia, where it is called Hilib geyl, Saudi Arabia, Egypt, Libya, Sudan, Kazakhstan and other arid regions where alternative forms of protein may be limited or where camel meat has had a long cultural history. In the Middle East, camel meat is the rarest and most prized source of pastırma. Not just the meat, but also blood is a consumable item as is the case in northern Kenya, where camel blood is a source of iron, vitamin D, salts and minerals (although Muslims do not drink or consume blood products). According to Jewish tradition, camel meat and milk are taboo. Camels possess only one of the two Kosher criteria; although they chew their cuds, they do not possess split hooves. (See: Taboo food and drink)

Health issues

A 2005 report issued jointly by the Saudi Ministry of Health and the United States Center for Disease Controls details cases of human bubonic plague resulting from the ingestion of raw camel liver.[7]

  1. ^ Wild Bactrian Camel, Animal Info
  2. ^ What secrets lie within the camel's hump?, Lund University, Sweden, Accessed 7 January 2008
  3. ^ Unique properties of the camel erythrocyte membrane, NCBI, 5 April 1975
  4. ^ Dromedary, Hannover Zoo, Accessed 8 January 2008
  5. ^ 'Camelized' antibodies make waves, news story, Robert Koenig, Science, 318:1373, 30 November 2007.
  6. ^ Fresh from your local drome'dairy'? Food and Agriculture Organization, July 6, 2001
  7. ^ Plague from Eating Raw Camel Liver, Centers for Disease Control and Prevention, Vol 11, No. 9, September 2005