Cochlops

Cochlops; Temporal range: Early-Middle Miocene ; ~18–12 Ma PreꞒ Ꞓ O S D C P T J K Pg N
	Carapace of Cochlops
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Cingulata
Family:	Chlamyphoridae
Subfamily:	†Glyptodontinae
Genus:	†Cochlops; Ameghino, 1889
Type species
	Cochlops muricatus; Ameghino, 1889
Species
	C. debilis Ameghino, 1891; C. muricatus Ameghino, 1889;
Synonyms
	Metopotoxus Ameghino, 1895;

Cochlops is an extinct genus of glyptodont. It lived from the Early to Middle Miocene, and its fossilized remains have been found in South America.

Description

This animal, like all glyptodonts, had an armor formed by numerous osteoderms fused together, protecting most of its body. Its skull was characterized by a shortened rostrum, a facial profile strongly inclined towards the front, and the occipital plane was oblique. The shape of its skull prefigured that of later genera such as Panochthus. Its carapace was characterized by particularly wrinkled osteoderms, especially compared to other forms of archaic glyptodonts, such as Asterostemma and Propalaehoplophorus ; some osteoderms, especially in the pelvic area, had a particular ornamentation, with a large central conical tubercle surrounded by smaller wrinkled tubercles.

Classification

The genus Cochlops was first described in 1889 by Florentino Ameghino, based on fossil remains found in Early Miocene terrains of Argentina. The type species is Cochlops muricatus, and the species C. debilis, described in 1891 by Ameghin, is also ascribed to the genus. The genus Metopotoxus was later synonymized with the genus Cochlops.

Cochlops was a basal glyptodont, belonging to the tribe Propalaehoplophorini, and related to the genera Propalaehoplophorus, Asterostemma and Eucinepeltus.

Modern cladistic analysis suggests that Propalaehoplophorini is a paraphyletic group. Cladogram after Barasoain et al. 2022:^[1]

Glyptodonts

Traditional Glyptodontinae

Boreostemma

	Glyptotherium

	Glyptodon

"Austral clade"

Propalaehoplophorus

Cochlops

	Plohophorus

	Pseudohoplophorus

Doedicurinae

	Doedicurus

	Eleutherocercus

Neosclerocalyptus

Hoplophorini

Hoplophorus

	Propanochthus

	Panochthus

Bibliography

F. Ameghino. 1889. Contribución al conocimiento de los mamíferos fósiles de la República Argentina [Contribution to the knowledge of the fossil mammals of the Argentine Republic]. Actas de la Academia Nacional de Ciencias de la República Argentina en Córdoba 6:xxxii-1027
F. Ameghino. 1891. Nuevos restos de mamíferos fósiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. – Especies nuevas, adiciones y correcciones [New remains of fossil mammals discovered by Carlos Ameghino in the lower Eocene of southern Patagonia. – New species, additions, and corrections]. Revista Argentina de Historia Natural 1:289-328
S. F. Vizcaíno, J. C. Fernicola, and M. S. Bargo. 2012. Paleobiology of Santacrucian glyptodonts and armadillos (Xenarthra, Cingulata). In S. F. Vizcaíno, R. F. Kay, M. S. Bargo (eds.), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation 194–215

References

^ Barasoain, Daniel; Zurita, Alfredo E.; Croft, Darin A.; Montalvo, Claudia I.; Contreras, Víctor H.; Miño-Boilini, Ángel R.; Tomassini, Rodrigo L. (June 2022). "A New Glyptodont (Xenarthra: Cingulata) from the Late Miocene of Argentina: New Clues About the Oldest Extra-Patagonian Radiation in Southern South America". Journal of Mammalian Evolution. 29 (2): 263–282. doi:10.1007/s10914-021-09599-w. ISSN 1064-7554. S2CID 245945029.

[:2-1] Barasoain, Daniel; Zurita, Alfredo E.; Croft, Darin A.; Montalvo, Claudia I.; Contreras, Víctor H.; Miño-Boilini, Ángel R.; Tomassini, Rodrigo L. (June 2022). "A New Glyptodont (Xenarthra: Cingulata) from the Late Miocene of Argentina: New Clues About the Oldest Extra-Patagonian Radiation in Southern South America". Journal of Mammalian Evolution. 29 (2): 263–282. doi:10.1007/s10914-021-09599-w. ISSN 1064-7554. S2CID 245945029.

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