Triplophysa
Triplophysa | |
---|---|
Triplophysa dalaica | |
Triplophysa stolickai | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Cypriniformes |
Family: | Nemacheilidae |
Genus: | Triplophysa Rendahl (de), 1933 |
Type species | |
Nemacheilus hutjertjuensis Rendahl, 1933
| |
Synonyms | |
see text |
Triplophysa is a genus of fish in the family Nemacheilidae found mainly in and around the Qinghai-Tibet Plateau in China.[1] Currently, the genus is a mixed assemblage of species. Some lineages have been identified and treated as subgenera (Hedinichthys, Indotriplophysa, Labiatophysa, Qinghaichthys and Tarimichthys), but as Wikipedia follows Fishbase for fish species these have been treated as subgenera in Wikipedia, although Kottelat and the Catalog of Fishes treat them as genera.[2][3] FishBase, however, includes these in Triplophysa without specifying subgenera and treats the names given by Kottelat as synonyms.[4]
Ecology
Triplophysa zhaoi holds the record for the lowest altitude for Asian fish: it is found at 50 m (160 ft) below sea level in swamps of the Lükqün oasis, in the Turpan Depression in Xinjiang. In the other end, Triplophysa stolickai holds the record altitude for Asian fish: it is found at 5,200 m (17,100 ft) above sea level in hot springs near the Longmu Lake in western Tibet.[2] Triplophysa dalaica has been used as model species to study adaptation to high-altitude hypoxia and 13 positively selected genes involved in hypoxia response have been identified.[5] Some species are blind cave-dwellers.[1][6]
Species
There are currently 131 recognized species in this genus:[2]
- Triplophysa alexandrae Prokofiev, 2001
- Triplophysa aliensis (Y. F. Wu & S. Q. Zhu, 1979)
- Triplophysa alticeps (Herzenstein, 1888)
- Triplophysa altipinnis Prokofiev, 2003
- Triplophysa aluensis W. X. Li & Z. G. Zhu, 2000
- Triplophysa angeli (P. W. Fang, 1941)
- Triplophysa anshuiensis Wu, Wei, Lan, Du, 2018[7]
- Triplophysa anterodorsalis S. Q. Zhu & W. X. Cao, 1989
- Triplophysa aquaecaeruleae Prokofiev, 2001
- Triplophysa arnoldii Prokofiev, 2006
- Triplophysa bashanensis T. Q. Xu & K. F. Wang, 2009
- Triplophysa bellibarus (T. L. Tchang, T. H. Yueh & H. C. Hwang, 1963)
- Triplophysa bleekeri (Sauvage & Dabry de Thiersant, 1874)
- Triplophysa brachyptera (Herzenstein, 1888)
- Triplophysa brahui (Zugmayer, 1912)
- Triplophysa brevibarba R. H. Ding, 1993
- Triplophysa brevicauda (Herzenstein, 1888)
- Triplophysa cakaensis W. X. Cao & S. Q. Zhu, 1988
- Triplophysa chondrostoma (Herzenstein, 1888)
- Triplophysa choprai (Hora, 1934)
- Triplophysa coniptera (Turdakov, 1954)
- Triplophysa crassicauda (Herzenstein, 1888)
- Triplophysa crassilabris R. H. Ding, 1994
- Triplophysa cuneicephala (T. H. Shaw & T. L. Tchang, 1931)
- Triplophysa dalaica (Kessler, 1876)
- Triplophysa daochengensis Y. Y. Wu, Z. Y. Sun & Y. S. Guo, 2016 [8]
- Triplophysa daqiaoensis R. H. Ding, 1993
- Triplophysa dongganensis J. Yang, 2013 [9]
- Triplophysa dorsalis (Kessler, 1872) (Gray loach)
- Triplophysa dorsonotata (Kessler, 1879)
- Triplophysa eugeniae (Prokofiev, 2002)
- Triplophysa farwelli (Hora, 1935)
- Triplophysa fengshanensis J. H. Lan, 2013 [9]
- Triplophysa flavicorpus J. X. Yang, X. Y. Chen & J. H. Lan, 2004
- Triplophysa furva S. Q. Zhu, 1992
- Triplophysa fuxianensis J. X. Yang & X. L. Chu, 1990
- Triplophysa gejiuensis (X. L. Chu & Y. R. Chen, 1979)
- Triplophysa gerzeensis W. X. Cao & S. Q. Zhu, 1988
- Triplophysa gracilis (F. Day, 1877)
- Triplophysa grahami (Regan, 1906)
- Triplophysa griffithii (Günther, 1868)
- Triplophysa grummorum Prokofiev, 2010
- Triplophysa gundriseri Prokofiev, 2002
- Triplophysa herzensteini (Berg, 1909)
- Triplophysa hexiensis (T. Q. Zhao & X. T. Wang, 1988)
- Triplophysa heyangensis S. Q. Zhu, 1992
- Triplophysa hialmari Prokofiev, 2001
- Triplophysa hsutschouensis (Rendahl (de), 1933)
- Triplophysa huanglongensis Gao, 1992
- Triplophysa huanjiangensis J. Yang, T. J. Wu & J. H. Lan, 2011
- Triplophysa huapingensis L. P. Zheng, J. X. Yang & X. Y. Chen, 2012 [10]
- Triplophysa hutjertjuensis (Rendahl (de), 1933)
- Triplophysa incipiens (Herzenstein, 1888)
- Triplophysa intermedia (Kessler, 1876)
- Triplophysa jianchuanensis L. P. Zheng, L. N. Du, X. Y. Chen & J. X. Yang, 2010 (Jianchuan stone loach)
- Triplophysa jiarongensis Y. Lin, C. Li & J. K. Song, 2012 [11]
- Triplophysa kafirnigani (Turdakov, 1948)
- Triplophysa kashmirensis (Hora, 1922) [12]
- Triplophysa kaznakowi Prokofiev, 2004
- Triplophysa kullmanni Bănărescu, Nalbant & Ladiges, 1975
- Triplophysa kungessana (Kessler, 1879)
- Triplophysa labiata (Kessler, 1874) (Plain thick-lip loach)
- Triplophysa lacusnigri (L. S. Berg, 1928)
- Triplophysa lacustris J. X. Yang & X. L. Chu, 1990
- Triplophysa langpingensis J. Yang, 2013 [9]
- Triplophysa laterimaculata J. L. Li, N. F. Liu & J. X. Yang, 2007
- Triplophysa laticeps W. Zhou & G. H. Cui, 1997
- Triplophysa leptosoma (Herzenstein, 1888)
- Triplophysa lihuensis T. J. Wu, J. Yang & J. H. Lan, 2012 [13]
- Triplophysa lixianensis C. L. He, Z. B. Song & E. Zhang, 2008
- Triplophysa longianguis Y. F. Wu & C. Z. Wu, 1984
- Triplophysa longibarbata (Y. R. Chen, J. X. Yang, Sket & Aljančič, 1998)
- Triplophysa longipectoralis L. P. Zheng, L. N. Du, X. Y. Chen & J. X. Yang, 2009
- Triplophysa longliensis Q. Ren, J. X. Yang & X. Y. Chen, 2012 [14]
- Triplophysa macrocephala J. Yang, T. J. Wu & J. X. Yang, 2012 [15]
- Triplophysa macromaculata J. X. Yang, 1990
- Triplophysa macrophthalma S. Q. Zhu & Q. Z. Guo, 1985
- Triplophysa macropterus (Herzenstein, 1888)
- Triplophysa markehenensis (S. Q. Zhu & Y. F. Wu, 1981)
- Triplophysa marmorata (Heckel, 1838) [12]
- Triplophysa microphysa (P. W. Fang, 1935)
- Triplophysa microps (Steindachner, 1866)
- Triplophysa microphthalma (Kessler, 1879)
- Triplophysa minuta (Li, 1966)
- Triplophysa moquensis R. H. Ding, 1994
- Triplophysa nandanensis J. H. Lan, J. X. Yang & Y. R. Chen, 1995
- Triplophysa nasalis (Kessler, 1876) (incertae sedis but probably belongs in this genus)
- Triplophysa nanpanjiangensis (S. Q. Zhu & W. X. Cao, 1988)
- Triplophysa nasobarbatula D. Z. Wang & D. J. Li, 2001
- Triplophysa ninglangensis Y. F. Wu & C. Z. Wu, 1988
- Triplophysa nujiangensa X. Y. Chen, G. H. Cui & J. X. Yang, 2004
- Triplophysa obscura X. T. Wang, 1987
- Triplophysa obtusirostra Y. F. Wu & C. Z. Wu, 1988
- Triplophysa orientalis (Herzenstein, 1888)
- Triplophysa papillosolabiata (Kessler, 1879)
- Triplophysa pappenheimi (P. W. Fang, 1935)
- Triplophysa paradoxa (Turdakov, 1955)
- Triplophysa parvus Z. M. Chen, W. X. Li & J. X. Yang, 2009
- Triplophysa polyfasciata R. H. Ding, 1996
- Triplophysa pseudoscleroptera (S. Q. Zhu & Y. F. Wu, 1981)
- Triplophysa pseudostenura C. L. He, E. Zhang & Z. B. Song, 2012 [16]
- Triplophysa qilianensis W. J. Li, X. C. Chen & Y. P. Hu, 2015 [17]
- Triplophysa qiubeiensis W. X. Li & H. F. Yang, 2008
- Triplophysa robusta (Kessler, 1876)
- Triplophysa rosa X. Y. Chen & J. X. Yang, 2005
- Triplophysa rotundiventris (Y. F. Wu & Yuan Chen, 1979)
- Triplophysa scapanognatha Prokofiev, 2007
- Triplophysa scleroptera (Herzenstein, 1888)
- Triplophysa sellaefer (Nichols, 1925)
- Triplophysa sewerzowi (G. V. Nikolskii, 1938) (Severtsov’s loach)
- Triplophysa shaanxiensis J. X. Chen, 1987
- Triplophysa shilinensis Y. R. Chen & J. X. Yang, 1992
- Triplophysa shiyangensis (T. Q. Zhao & X. T. Wang, 1983)
- Triplophysa siluroides (Herzenstein, 1888)
- Triplophysa stenura (Herzenstein, 1888)
- Triplophysa stewarti (Hora, 1922)
- Triplophysa stolickai (Steindachner, 1866) (Tibetan stone loach)
- Triplophysa strauchii (Kessler, 1874) (Spotted thick-lip loach)
- Triplophysa tanggulaensis (S. Q. Zhu, 1982)
- Triplophysa tenuicauda (Steindachner, 1866)
- Triplophysa tenuis (F. Day, 1877)
- Triplophysa tianeensis X. Y. Chen, G. H. Cui & J. X. Yang, 2004
- Triplophysa tianxingensis H. F. Yang, W. X. Li & Z. M. Chen, 2016 [18]
- Triplophysa tibetana (Regan, 1905)
- Triplophysa turpanensis Y. F. Wu & C. Z. Wu, 1992
- Triplophysa ulacholica (V. P. Anikin, 1905) (Issyk-kul naked loach)
- Triplophysa uranoscopus (Kessler, 1872)
- Triplophysa venusta S. Q. Zhu & W. X. Cao, 1988
- Triplophysa waisihani L. Cao & E. Zhang, 2008
- Triplophysa wuweiensis (S. C. Li & S. Y. Chang, 1974)
- Triplophysa xiangshuingensis W. X. Li, 2004
- Triplophysa xiangxiensis (G. Y. Yang, F. X. Yuan & Y. M. Liao, 1986)
- Triplophysa xichouensis Liu, Pan, Yang & Chen, 2017[19]
- Triplophysa xichangensis S. Q. Zhu & W. X. Cao, 1989
- Triplophysa xingshanensis (G. R. Yang & C. X. Xie, 1983)
- Triplophysa xiqiensis R. H. Ding & Q. Lai, 1996
- Triplophysa yajiangensis S. L. Yan, Z. Y. Sun & Y. S. Guo, 2015 [20]
- Triplophysa yaopeizhii T. Q. Xu, C. G. Zhang & B. Cai, 1995
- Triplophysa yarkandensis (F. Day, 1877) (Kashgarian loach)
- Triplophysa yasinensis (Alcock, 1898)
- Triplophysa yunnanensis J. X. Yang, 1990
- Triplophysas zaidamensis (Kessler, 1874)
- Triplophysa zamegacephala (T. Q. Zhao, 1985)
- Triplophysa zhaoi Prokofiev, 2006
- Triplophysa zhenfengensis D. Z. Wang & D. J. Li, 2001
References
- ^ a b Chen, X.-Y.; Yang, J.-X. (2005). "Triplophysa rosa sp. nov.: a new blind loach from China". Journal of Fish Biology. 66 (3): 599–608. doi:10.1111/j.0022-1112.2005.00622.x.
- ^ a b c Kottelat, M. (2012). "Conspectus_cobitidum.pdf Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei)" (PDF). The Raffles Bulletin of Zoology. Supplement No. 26: 1–199.
- ^ Eschmeyer, W.N. (30 Aug 2016). "Catalog of Fishes". California Academy of Sciences.
- ^ Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Triplophysa". FishBase. June 2016 version.
- ^ Wang, Y.; Yang, L.; Wu, B.; Song, Z.; He, S. (2015). "Transcriptome analysis of the plateau fish (Triplophysa dalaica): Implications for adaptation to hypoxia in fishes". Gene. 565 (2): 211–220. doi:10.1016/j.gene.2015.04.023. PMID 25869933.
- ^ Romero, A.; Zhao, Y.; Chen, X. (2009). "The Hypogean fishes of China". Environmental Biology of Fishes. 86 (1): 211–278. doi:10.1007/s10641-009-9441-3.
- ^ "Triplophysa anshuiensis, a new species of blind loach from the Xijiang River, China (Teleostei, Nemacheilidae)". Zookeys. Retrieved 3 April 2018.
- ^ Wu, Y.-Y.; Sun, Z.-Y.; Guo, Y.-S. (2016). "A new species of the genus Triplophysa (Cypriniformes: Nemacheilidae), Triplophysa daochengensis, from Sichuan Province, China". Zoological Research. 37 (5): 290–296.
- ^ a b c Cave Fishes of Guangxi, China. Science Press. 2013. p. 284. ISBN 9787030376589.
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ignored (help) - ^ Zheng, L.-P.; Yang, J.-X.; Chen, X.-Y. (2012). "A new species of Triplophysa (Nemacheilidae: Cypriniformes), from Guangxi, southern China". Journal of Fish Biology. 80 (4): 831–841. doi:10.1111/j.1095-8649.2012.03227.x. PMID 22471802.
- ^ Lin, Y.; Li, C.; Song, J.-K. (2012). "A new species of troglobitic loach (Cypriniformes, Balitoridae), Triplophysa jiarongensis, from Guizhou, China". Acta Zootaxonomica Sinica. 37 (3): 640–647.
- ^ a b Bashir, A.; Bisht, B.S.; Mir, J.I.; Kumar, R.; Patiyal, R.S. (2016). "Morphological, molecular characterization and taxonomic status of Triplophysa marmorata and Triplophysa kashmirensis (Cypriniformes: Nemacheilidae) from Kashmir valley, India". Revista de Biología Tropical. 64 (2): 473–482. doi:10.15517/rbt.v64i2.19591. PMID 29451748.
- ^ Wu, T.-J.; Yang, J.; Lan, J.-H. (2012). "A New Blind Loach Triplophysa lihuensis sp. nov. (Teleostei: Balitoridae) from Guangxi, China" (PDF). Zoological Studies. 51 (6): 874–880.
- ^ Ren, Q.; Yang, J.-X.; Chen, X.-Y. (2012). "A new species of the genus Triplophysa (Cypriniformes: Nemacheilidae), Triplophysa longliensis sp. nov., from Guizhou, China" (PDF). Zootaxa. 3586: 187–194. doi:10.11646/zootaxa.3586.1.17.
- ^ Yang, J.; Wu, T.-J.; Yang, J.-X. (2012). "A new cave-dwelling loach, Triplophysa macrocephala (Teleostei: Cypriniformes: Balitoridae), from Guangxi, China". Environmental Biology of Fishes. 93 (2): 169–175. doi:10.1007/s10641-011-9901-4.
- ^ He, C.; Zhang, E.; Song, Z. (2012). "Triplophysa pseudostenura, a new nemacheiline loach (Cypriniformes: Balitoridae) from the Yalong River of China" (PDF). Zootaxa. 3586: 272–280. doi:10.11646/zootaxa.3586.1.26.
- ^ Li, W.-J.; Chen, X.-C.; Hu, Y.-P. (2015). "A new species of the genus Triplophysa (Nemacheilinae), Triplophysa qilianensis sp. nov., from Qinghai, China" (PDF). Zootaxa. 3905 (3): 418–424. doi:10.11646/zootaxa.3905.3.7.
- ^ Yang, H.-F.; Li, W.-X.; Chen, Z.-M. (2016). "A new cave species of the genus Triplophysa from Yunnan, China". Zoological Research. 37 (5): 297–300. doi:10.13918/j.issn.2095-8137.2016.5.296. PMC 5071343. PMID 27686789.
- ^ Liu, S. W.; Pan, X. F.; Yang, J. X.; Chen, X. Y. (2017). "A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China". Journal of Fish Biology. 90 (3): 834–846. doi:10.1111/jfb.13201. ISSN 0022-1112. PMID 28155227.
- ^ Yan, S.-L.; Sun, Z.-Y.; Guo, Y.-S. (2015). "A new species of Triplophysa Rendahl (Cypriniformes, Nemacheilidae) from Sichuan Province, China". Zoological Research. 36 (5): 299–304. PMC 4771960. PMID 26452694.