Cetotheriidae
| Cetotheriidae Temporal range:
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|---|---|
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| Heterocetus guiscardii (nomen dubium) vertebrae | |
Scientific classification 
| |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Artiodactyla |
| Infraorder: | Cetacea |
| Parvorder: | Mysticeti |
| Family: | Cetotheriidae Brandt 1872 |
| Subfamilies and genera | |
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See text | |
Cetotheriidae is a family of baleen whales (suborder Mysticeti).[1] The family was generally thought to have existed from the Late Oligocene to the Early Pleistocene before going extinct. However, the phylogenetic studies conducted by Fordyce & Marx 2013 recovered the living pygmy right whale as a member of Cetotheriidae; if confirmed these results would resurrect Cetotheriidae from extinction.
Taxonomy
After its description by Brant in 1872, Cetotheriidae was used as a wastebasket taxon for baleen whales which were not assignable to extant whale families.[2]
Comparing the cranial and mandibular morphology of 23 taxa (including late archaeocetes and both fossil and extant mysticetes),[3] Bouetel & Muizon 2006 found Cetotheriidae in this traditional sense to be polyphyletic. Based on ten cranial characters, they also concluded that of the twelve included fossil baleen-bearing mysticetes, six formed a monophyletic group, Cetotheriidae sensu stricto.[4]
Several phylogenetic studies since Bouetel & Muizon 2006 support the monophyly of a small group of core Cetotheriidae sensu stricto, archaic mysticetes with a cranium that have "a long ascending process of the maxilla with anteriorly diverging lateral border that interdigitates with the frontal" and some other characters.[5] This group is limited to Cetotherium rathkii, Metopocetus durinasus, Mixocetus elysius, Herpetocetus scaldiensis, H. transatlanticus, H. bramblei, Nannocetus eremus, and Piscobalaena nana.[6] The remaining genera placed in the family are considered Cetotheriidae sensu lato and are often referred to as the 'cetotheres'.[5]
Bisconti, Lambert & Bosselaers 2013 considered the primitive 'cetothere' Joumocetus the most basal named taxon of their new superfamily Thalassotherii (Cetotheriidae s.l., Cetotheriidae s.s., Eschrichtiidae (gray whales) and Balaenopteridae (rorquals)) and suggested that the term "Cetotheriidae s.l." should be renamed "basal" or "stem thalassotherians".[7] Fordyce & Marx 2013 found that the pygmy right whale formed a well-supported clade with Eschrichtiidae and Balaenopteridae based on molecular data, and that, within 'cetotheres', it was most closely related to the herpetocetines (Herpetocetus and Nannocetus)[8] Bisconti et al. 2013, however, found, based on morphological data, it to be more closely related to Balaenidae (the bowhead and right whales), but added that additional specimens is expected to resolve these conflicting results within a few years.[9]
Classification of Cetotheriidae according to Eli Adli et al. (2014)[10] and the Fossilworks online database:[1]
- †Cephalotropis Cope 1896[11]
- †Joumocetus Kimura & Hasegawa 2010[11]
- Subfamily: †Cetotheriinae Whitmore and Barnes, 2008
- †Brandtocetus Gol'din and Startsev 2014[12]
- †Cetotherium Brandt 1843[11][13]
- †Eucetotherium Brandt 1873[11]
- †Imerocetus Mchedlidze 1964
- †Kurdalagonus Tarasenko & Lopatin 2012a[11]
- †Otradnocetus Mchedlidze 1984
- †Vampalus Tarasenko & Lopatin 2012b
- †Zygiocetus Tarasenko 2014[14]
- Subfamily †Herpetocetinae Steeman, 2007
- †Herentalia Bisconti 2015[15]
- †Herpetocetus Van Beneden 1872[11][13]
- †Metopocetus Cope 1896[11][13]
- †Nannocetus Kellogg 1929[11][13]
- †Piscobalaena Pilleri and Siber 1989[11][13]
- Subfamily Neobalaeninae Miller 1923
- Caperea (pygmy right whale) Gray 1873
- †Miocaperea Bisconti 2012
Incertae sedis:
- †Hibacetus Otsuka and Ota 2008
References
Notes
- ^ a b "Cetotheriidae". Fossilworks. Retrieved December 2013.
{{cite web}}: Check date values in:|accessdate=(help) - ^ Steeman 2010
- ^ Bouetel & Muizon 2006, p. 373
- ^ Bouetel & Muizon 2006, Abstract
- ^ a b Bisconti, Lambert & Bosselaers 2013, p. 96
- ^ Bisconti, Lambert & Bosselaers 2013, p. 121
- ^ Bisconti, Lambert & Bosselaers 2013, p. 98
- ^ Fordyce & Marx 2013, p. 3
- ^ Fordyce & Marx 2013, pp. 122–123
- ^ Joseph J. El Adli, Thomas A. Deméré and Robert W. Boessenecker (2014). "Herpetocetus morrowi (Cetacea: Mysticeti), a new species of diminutive baleen whale from the Upper Pliocene (Piacenzian) of California, USA, with observations on the evolution and relationships of the Cetotheriidae". Zoological Journal of the Linnean Society 170 (2): 400–466. doi:10.1111/zoj.12108.
- ^ a b c d e f g h i Gol'din, Startsev & Krakhmalnaya 2013, pp. 4–5
- ^ Pavel Gol’din and Dmitry Startsev (2014). "Brandtocetus, a new genus of baleen whales (Cetacea, Cetotheriidae) from the late Miocene of Crimea, Ukraine". Journal of Vertebrate Paleontology 34 (2): 419–433. doi:10.1080/02724634.2013.799482.
- ^ a b c d e Bouetel & Muizon 2006, p. 376
- ^ K. K. Tarasenko. 2014. New Genera of Baleen Whales (Cetacea, Mammalia) from the Miocene of the Northern Caucasus and Ciscaucasia: 3. Zygiocetus gen. nov. (Middle Sarmatian, Adygea). Paleontological Journal 48(5):551-562
- ^ Michelangelo Bisconti (2015). Anatomy of a new cetotheriid genus and species from the Miocene of Herentals, Belgium, and the phylogenetic and palaeobiogeographical relationships of Cetotheriidae s.s. (Mammalia, Cetacea, Mysticeti). Journal of Systematic Palaeontology 13 (5): 377-395. doi:10.1080/14772019.2014.890136.
Sources
- Bisconti, M.; Lambert, O.; Bosselaers, M. (2013). "Taxonomic revision of Isocetus depauwi (Mammalia, Cetacea, Mysticeti) and the phylogenetic relationships of archaic 'cetothere' mysticetes". Palaeontology. 56 (1): 95–127. doi:10.1111/j.1475-4983.2012.01168.x. ISSN 0031-0239. OCLC 826744606.
{{cite journal}}:|access-date=requires|url=(help); Check date values in:|accessdate=(help); Invalid|ref=harv(help); Unknown parameter|layurl=ignored (help) - Bouetel, V.; Muizon, C., de (2006). "The anatomy and relationships of Piscobalaena nana (Cetacea, Mysticeti), a Cetotheriidae s.s. from the early Pliocene of Peru" (PDF). Geodiversitas. 28 (2): 319–395. Retrieved November 2013.
{{cite journal}}: Check date values in:|accessdate=(help); Invalid|ref=harv(help)CS1 maint: multiple names: authors list (link) - Brandt, J. F. (1872). "Über eine neue Classification der Bartenwhale (Balaenoidea) mit Berücksichtigung der untergegangenen Gattungen derselben". Bulletin de L'Académie imperiale des sciences de St.-Pétersbourg. 3 (in German). 17. Retrieved November 2013.
{{cite journal}}: Check date values in:|accessdate=(help); Invalid|ref=harv(help); Unknown parameter|laydate=ignored (help); Unknown parameter|layurl=ignored (help) - Fordyce, R. Ewan; Marx, Felix G. (2013). "The pygmy right whale Caperea marginata: the last of the cetotheres". Proceedings of the Royal Society B: Biological Sciences. 280 (1753). doi:10.1098/rspb.2012.2645.
{{cite journal}}: Invalid|ref=harv(help) - Gol'din, P.; Startsev, D.; Krakhmalnaya, T. (2013). "The anatomy of Cetotherium riabinini Hofstein, 1948, a baleen whale from the late Miocene of Ukraine". Acta Palaeontologica Polonica. In press. doi:10.4202/app.2012.0107.
{{cite journal}}: Invalid|ref=harv(help) - Kimura, T.; Hasegawa, Y. (2010). "A New Baleen Whale (Mysticeti: Cetotheriidae) from the Earliest Late Miocene of Japan and a Reconsideration of the Phylogeny of Cetotheres". Journal of Vertebrate Paleontology. 30 (2): 577–591. doi:10.1080/02724631003621912.
{{cite journal}}: Invalid|ref=harv(help) - Steeman, M. E. (2010). "The extinct baleen whale fauna from the Miocene-Pliocene of Belgium and the diagnostic cetacean ear bones". Journal of Systematic Palaeontology. 8 (1): 63–80. doi:10.1080/14772011003594961.
{{cite journal}}: Invalid|ref=harv(help) - Tarasenko, K. K.; Lopatin, A. V. (2012a). "New Baleen Whale Genera (Cetacea, Mammalia) from the Miocene of the Northern Caucasus and Ciscaucasia: 1. Kurdalagonus gen. nov. from the Middle–Late Sarmatian of Adygea". Paleontological Journal. 46 (5): 531–542. doi:10.1134/s0031030112050115.
{{cite journal}}: Invalid|ref=harv(help); Unknown parameter|laydate=ignored (help); Unknown parameter|layurl=ignored (help) - Tarasenko, K. K.; Lopatin, A. V. (2012b). "New Baleen Whale Genera (Cetacea, Mammalia) from the Miocene of the Northern Caucasus and Ciscaucasia: 2. Vampalus gen. nov. from the Middle-Late Miocene of Chechnya and Krasnodar Region". Paleontological Journal. 46 (6): 620–629. doi:10.1134/s003103011206010x.
{{cite journal}}: Invalid|ref=harv(help); Unknown parameter|laydate=ignored (help); Unknown parameter|layurl=ignored (help)
