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Samolus valerandi ssp parviflorus.jpg
S. valerandi
Scientific classification e
Kingdom: Plantae
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Ericales
Family: Primulaceae
Subfamily: Theophrastoideae
Tribe: Samoleae
Genus: Samolus

Samolus (known as brookweed, or water pimpernel) is a widely distributed genus of about a dozen species of water-loving herbs. According to the APG III classification, this genus belongs to the Primulaceae family in the order Ericales. It was considered as closely related to a clade comprising the Theophrastaceae,[1] and was treated as part of that family or in its own monogeneric family, the Samolaceae. The APG III system does not recognize these families and instead includes all species formerly belonging to Theophrastaceae in the Primulaceae family.


The last complete taxonomic treatment of this genus recognizes the following species:[2][3]

Although some local floras recognize the North-American populations of S. valerandi as a separate species (S. parviflorus) or subspecies (S. valerandi ssp. parviflorus),[4] molecular and morphological data indicate that S. vagans and S. parviflorus should not be regarded as separate species but as part of a widespread "S. valerandi species complex.[5]


  1. ^ Caris PL, Smets EF (2004). "A floral ontogenetic study on the sister group relationship between the genus Samolus (Primulaceae) and the Theophrastaceae". American Journal of Botany. 91 (5): 627–643. doi:10.3732/ajb.91.5.627. PMID 21653418.
  2. ^ Crusio WE (20 February 1982). "Het geslacht Samolus L". Communications of the Dutch Waterplant Society (in Dutch). 2: 13–25.
  3. ^ Crusio WE (26 May 1984). "Notes on the genus Samolus L. (Primulaceae)". Communications of the Dutch Waterplant Society. 6: 13–16.
  4. ^ "Samolus". Flora of North America. Retrieved 2018-02-01.
  5. ^ Jones, K.; Anderberg, A. A.; Ronse De Craene, L. P.; Wanntorp, L. (23 June 2012). "Origin, diversification, and evolution of Samolus valerandi (Samolaceae, Ericales)". Plant Systematics and Evolution. 298 (8): 1523–1531. doi:10.1007/s00606-012-0655-z.