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Temporal range: Early Paleocene - Holocene, 65–0 Ma
Talpa europaea MHNT.jpg
European mole (Talpa europaea)
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Clade: Exafroplacentalia
Magnorder: Boreoeutheria

Boreoeutheria (synonymous with Boreotheria) (Greek: βόρειο "north" + ευ "good" + θεριό "beast") is a clade (magnorder) of placental mammals that is composed of the sister taxa Laurasiatheria (most hoofed mammals, most pawed carnivores, and several other groups) and Euarchontoglires (Supraprimates). It is now well supported by DNA sequence analyses, as well as retrotransposon presence or absence data.

The earliest known fossils belonging to this group date to about 65 million years ago, shortly after the K-Pg extinction event, though molecular data suggest they may have originated earlier, during the Cretaceous period.[1]

With the exception of rhinoceroses and cetaceans, male members of the clade share the distinction of external testicles.[2]

Boreoeutherian ancestor[edit]

The common ancestor of Boreoeutheria lived between 100 and 80 million years ago. The boreoeutherian ancestor gave rise to species as diverse as giraffe, dog, mouse, bat, whale and humans. The concept of boreoeutherian ancestor was first proposed in 2004 in the journal Genome Research.[3][4] The genome sequence of the boreoeutherian ancestor can be computationally predicted with high accuracy. It is estimated to contain three billion base pairs.[3]


Class Mammalia


The weakly favoured cladogram favours Boreoeuthearia as a basal Eutherian clade as sister to the Atlantogenata.[5][6]









While it is agreed that the cetaceans evolved within artiodactyls, much of the branching order within Laurasiatheria is not yet well resolved. In particular, the most difficult order to place definitively has been and still is Perissodactyla: Their placement within Zooamata is controversial.

One study has suggested that the carnivores, cetaceans, chiroptera and ungulates form an ancient clade.[7] This is supported by another study that suggests that Eulipotyphla are the earliest diverging clade within the Laurasiatheria.[8]


  1. ^ O'Leary, M. A.; Bloch, J. I.; Flynn, J. J.; Gaudin, T. J.; Giallombardo, A.; Giannini, N. P.; Cirranello, A. L. (2013). "The placental mammal ancestor and the post–K-Pg radiation of placentals". Science. 339 (6120): 662–667. PMID 23393258. doi:10.1126/science.1229237. 
  2. ^ D. S. Mills; Jeremy N. Marchant-Forde (2010). The Encyclopedia of Applied Animal Behaviour and Welfare. CABI. pp. 293–. ISBN 978-0-85199-724-7. 
  3. ^ a b John Roach (January 25, 2005). "Scientists Recreate Genome of Ancient Human Ancestor". National Geographic. Retrieved 14 Feb 2015. 
  4. ^ Mathieu Blanchette; Eric D. Green; Webb Miller; David Haussler (2004). "Reconstructing large regions of an ancestral mammalian genome in silico". Genome Research. 14: 2412–2423. PMC 534665Freely accessible. PMID 15574820. doi:10.1101/gr.2800104. Retrieved 14 Feb 2015. 
  5. ^ Foley, Nicole M.; Springer, Mark S.; Teeling, Emma C. (2016-07-19). "Mammal madness: is the mammal tree of life not yet resolved?". Phil. Trans. R. Soc. B. 371 (1699): 20150140. ISSN 0962-8436. PMC 4920340Freely accessible. PMID 27325836. doi:10.1098/rstb.2015.0140. 
  6. ^ Tarver, James E.; Reis, Mario dos; Mirarab, Siavash; Moran, Raymond J.; Parker, Sean; O’Reilly, Joseph E.; King, Benjamin L.; O’Connell, Mary J.; Asher, Robert J. (2016-02-01). "The Interrelationships of Placental Mammals and the Limits of Phylogenetic Inference". Genome Biology and Evolution. 8 (2): 330–344. ISSN 1759-6653. PMC 4779606Freely accessible. PMID 26733575. doi:10.1093/gbe/evv261. 
  7. ^ Tsagkogeorga, G; Parker, J; Stupka, E; Cotton, JA; Rossiter, SJ (2013). "Phylogenomic analyses elucidate the evolutionary relationships of bats". Curr Biol. 23: 2262–2267. PMID 24184098. doi:10.1016/j.cub.2013.09.014. 
  8. ^ Morgan, CC; Foster, PG; Webb, AE; Pisani, D; McInerney, JO; O'Connell, MJ (2013). "Heterogeneous models place the root of the placental mammal phylogeny". Mol Biol Evol. 30 (9): 2145–256. doi:10.1093/molbev/mst117. 

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