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Temporal range: Late Cretaceous - Holocene, 65–0 Ma
Talpa europaea MHNT.jpg
European mole (Talpa europaea)
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Clade: Eutheria
Infraclass: Placentalia
Subcohort: Exafroplacentalia
Magnorder: Boreoeutheria

Boreoeutheria (synonymous with Boreotheria) (Greek: βόρειο "north" + ευ "good" + θεριό "beast") is a clade (magnorder) of placental mammals that is composed of the sister taxa Laurasiatheria (most hoofed mammals, most pawed carnivores, and several other groups) and Euarchontoglires (Supraprimates). It is now well supported by DNA sequence analyses, as well as retrotransposon presence or absence data.

The earliest known fossils belonging to this group date to about 65 million years ago, shortly after the K-Pg extinction event, though molecular data suggests they may have originated earlier, during the Cretaceous period.[1]

With the exception of rhinoceroses and cetaceans, male members of the clade share the distinction of external testicles.

Boreoeutherian ancestor[edit]

The common ancestor of Boreoeutheria lived between 100 and 80 million years ago. The boreoeutherian ancestor gave rise to species as diverse as cat, dog, mouse, rabbit, whale and humans. The concept of boreoeutherian ancestor was first proposed in 2004 in the journal Genome Research.[2][3] The genome sequence of the boreoeutherian ancestor can be computationally predicted with high accuracy. It is estimated to contain three billion base pairs.[2]


Class Mammalia











While it is agreed that the cetaceans evolved within artiodactyls much of the branching order within Laurasiatheria is not yet well resolved. In particular the most difficult order to place definitively has been and still is the Perissodactyla. Their placement within the Zooamata is controversial.

One study has suggested that the carnivores, cetaceans, chiroptera and ungulates form an ancient clade.[4] This is supported by another study that suggests that Eulipotyphla are the earliest diverging clade within the Laurasiatheria.[5]


  1. ^ O'Leary, M. A.; Bloch, J. I.; Flynn, J. J.; Gaudin, T. J.; Giallombardo, A.; Giannini, N. P.; Cirranello, A. L. (2013). "The placental mammal ancestor and the post–K-Pg radiation of placentals". Science 339 (6120): 662–667. 
  2. ^ a b John Roach (January 25, 2005). "Scientists Recreate Genome of Ancient Human Ancestor". National Geographic. Retrieved 14 Feb 2015. 
  3. ^ Mathieu Blanchette, Eric D. Green, Webb Miller, David Haussler (2004). "Reconstructing large regions of an ancestral mammalian genome in silico". Genome Research 14: 2412–2423. doi:10.1101/gr.2800104. Retrieved 14 Feb 2015. 
  4. ^ Tsagkogeorga, G; Parker, J; Stupka, E; Cotton, JA; Rossiter, SJ (2013). "Phylogenomic analyses elucidate the evolutionary relationships of bats". Curr Biol 23: 2262–2267. doi:10.1016/j.cub.2013.09.014. 
  5. ^ Morgan, CC; Foster, PG; Webb, AE; Pisani, D; McInerney, JO; O'Connell, MJ (2013). "Heterogeneous models place the root of the placental mammal phylogeny". Mol Biol Evol 30 (9): 2145–256. doi:10.1093/molbev/mst117. 

Additional references[edit]

  • Waddell, PJ; Kishino, H; Ota, R (2001). "A phylogenetic foundation for comparative mammalian genomics". Genome Inform Ser Workshop Genome Inform 12: 141–154. 
  • Murphy, William J.; Eizirik, Eduardo; Springer, Mark S.; et al. (2001). "Resolution of the Early Placental Mammal Radiation Using Bayesian Phylogenetics". Science 294 (5550): 2348–2351. 
  • Kriegs; Ole, Jan; Churakov, Gennady; Kiefmann, Martin; Jordan, Ursula; Brosius, Juergen; Schmitz, Juergen (2006). "Retroposed Elements as Archives for the Evolutionary History of Placental Mammals". PLoS Biol 4 (4): e91. doi:10.1371/journal.pbio.0040091. 
  • Blanchette, M; Green, ED; Miller, W; Haussler, D (Dec 2004). "Reconstructing large regions of an ancestral mammalian genome in silico". Genome Res 14 (12): 2412–23. doi:10.1101/gr.2800104. 
  • Ma, J; Zhang, L; Suh, BB; Raney, BJ; Burhans, RC; Kent, WJ; Blanchette, M; Haussler, D; Miller, W (Dec 2006). "Reconstructing contiguous regions of an ancestral genome". Genome Res 16 (12): 1557–65. doi:10.1101/gr.5383506. 

External links[edit]