|Fossil specimen of Darwinopterus modularis|
Wang et al., 2009
Wang et al., 2009
The Wukongopteridae were first named by Wang et al. in 2009, not yet giving an exact definition. The clade Wukongopteridae was first defined by Wang et al. in 2010 as "the most recent common ancestor of Wukongopterus lii and Kunpengopterus sinensis, and all of its descendants".
Wukongopterids are characterized by a unique combination of "primitive" and advanced pterosaurian features. While they had long tails and other features characteristic of other "rhamphorhynchoids", they also had distinct pterodactyloid features, such as long vertebrae in the neck and a single skull opening in front of the eyes, the nasoantorbital fenestra (in most "rhamphorhynchoids", the antorbital fenestra and the nasal opening are separate). This feature lead to Darwinopterus modularis being placed by Lü e.a. in a new clade of pterosaurs, the Monofenestrata, or "with a single opening", forming this group together with the true Pterodactyloidea, to the exclusion of the Rhamphorhynchidae and other more primitive pterosaurs which had separate nasal and antorbital fenestrae. However, according to Wang e.a. it is equally possible that the Wukongopteridae formed a more basal group, below the Rhamphorhynchidae.
- Lü J., Unwin D.M., Jin X., Liu Y., Ji Q. (2010). "Evidence for modular evolution in a long-tailed pterosaur with a pterodactyloid skull". Proceedings of the Royal Society B. 277 (1680): 383–389. doi:10.1098/rspb.2009.1603. PMC 2842655. PMID 19828548.CS1 maint: multiple names: authors list (link)
- *Wang X., Kellner A.W.A., Jiang S., Meng X. (2009). "An unusual long-tailed pterosaur with elongated neck from western Liaoning of China". Anais da Academia Brasileira de Ciências. 81 (4): 793–812. doi:10.1590/s0001-37652009000400016.CS1 maint: multiple names: authors list (link)
- Wang, Xiaolin; Kellner, Alexander W.A.; Jiang, Shunxing, Cheng, Xin, Meng, Xi & Rodrigues, Taissa (2010). "New long-tailed pterosaurs (Wukongopteridae) from western Liaoning, China" (PDF). Anais da Academia Brasileira de Ciências. 82 (4): 1045–1062. doi:10.1590/S0001-37652010000400024.CS1 maint: multiple names: authors list (link)