2021 in reptile paleontology: Difference between revisions
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* A study on the anatomy and replacement pattern of teeth in ''[[Henodus]] chelyops'' is published by Pommery ''et al.'' (2021).<ref>{{Cite journal|last1=Pommery |first1=Y. |last2=Scheyer |first2=T. M. |last3=Neenan |first3=J. M. |last4=Reich |first4=T. |last5=Fernandez |first5=V. |last6=Voeten |first6=D. F. A. E. |last7=Losko |first7=A. S. |last8=Werneburg |first8=I. |year=2021 |title=Dentition and feeding in Placodontia: tooth replacement in ''Henodus chelyops'' |journal=BMC Ecology and Evolution |volume=21 |pages=Article number 136 |doi=10.1186/s12862-021-01835-4 |doi-access=free }}</ref> |
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* Description of a new specimen of ''[[Panzhousaurus]] rotundirostris'' from the [[Guanling Formation]] (Guizhou, China), providing new information on the anatomy of this reptile, and a study on the phylogenetic relationships of this species is published by Lin ''et al.'' (2021).<ref>{{Cite journal|last1=Lin |first1=W.-B. |last2=Jiang |first2=D.-Y. |last3=Rieppel |first3=O. |last4=Motani |first4=R. |last5=Tintori |first5=A. |last6=Sun |first6=Z.-Y. |last7=Zhou |first7=M. |title=''Panzhousaurus rotundirostris'' Jiang et al., 2019 (Diapsida: Sauropterygia) and the Recovery of the Monophyly of Pachypleurosauridae |year=2021 |journal=Journal of Vertebrate Paleontology |volume=in press |pages=e1901730 |doi=10.1080/02724634.2021.1901730 }}</ref> |
* Description of a new specimen of ''[[Panzhousaurus]] rotundirostris'' from the [[Guanling Formation]] (Guizhou, China), providing new information on the anatomy of this reptile, and a study on the phylogenetic relationships of this species is published by Lin ''et al.'' (2021).<ref>{{Cite journal|last1=Lin |first1=W.-B. |last2=Jiang |first2=D.-Y. |last3=Rieppel |first3=O. |last4=Motani |first4=R. |last5=Tintori |first5=A. |last6=Sun |first6=Z.-Y. |last7=Zhou |first7=M. |title=''Panzhousaurus rotundirostris'' Jiang et al., 2019 (Diapsida: Sauropterygia) and the Recovery of the Monophyly of Pachypleurosauridae |year=2021 |journal=Journal of Vertebrate Paleontology |volume=in press |pages=e1901730 |doi=10.1080/02724634.2021.1901730 }}</ref> |
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* Redescription of the anatomy of the skull of ''[[Thalassomedon]] haningtoni'', and a study on the phylogenetic relationships of this species, is published by Sachs ''et al.'' (2021).<ref>{{Cite journal|last1=Sachs |first1=S. |last2=Lindgren |first2=J. |last3=Madzia |first3=D. |last4=Kear |first4=B. P. |year=2021 |title=Cranial osteology of the mid-Cretaceous elasmosaurid ''Thalassomedon haningtoni'' from the Western Interior Seaway of North America |journal=Cretaceous Research |volume=123 |pages=Article 104769 |doi=10.1016/j.cretres.2021.104769 }}</ref> |
* Redescription of the anatomy of the skull of ''[[Thalassomedon]] haningtoni'', and a study on the phylogenetic relationships of this species, is published by Sachs ''et al.'' (2021).<ref>{{Cite journal|last1=Sachs |first1=S. |last2=Lindgren |first2=J. |last3=Madzia |first3=D. |last4=Kear |first4=B. P. |year=2021 |title=Cranial osteology of the mid-Cretaceous elasmosaurid ''Thalassomedon haningtoni'' from the Western Interior Seaway of North America |journal=Cretaceous Research |volume=123 |pages=Article 104769 |doi=10.1016/j.cretres.2021.104769 }}</ref> |
Revision as of 20:03, 5 July 2021
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This list of fossil reptiles described in 2021 is a list of new taxa of fossil reptiles that were described during the year 2021, as well as other significant discoveries and events related to reptile paleontology that occurred in 2021.
Squamates
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Rage et al. |
Eocene |
|||||
Gen. et sp. nov |
Valid |
Scarpetta |
A member of Iguania. Genus includes new species C. lovei. |
|||||
Sp. nov |
Valid |
Bochaton, Charles & Lenoble |
Quaternary |
France |
||||
Sp. nov |
Valid |
Bolet et al. |
A lizard of uncertain phylogenetic placement. |
|||||
Sp. nov |
Valid |
Georgalis, Čerňanský & Klembara |
Probably late Eocene |
A member of Anguimorpha belonging to the family Palaeovaranidae. |
||||
Gen. et comb. nov |
Valid |
Smith & Habersetzer |
Eocene |
Messel Formation |
A member of Anguimorpha belonging to the family Palaeovaranidae. The type species is "Saniwa" feisti Stritzke (1983). |
|||
Sp. nov |
In press |
Longrich et al. |
A mosasaur. |
|||||
Sp. nov |
Valid |
Thorn et al. |
Late Oligocene |
An egerniine skink. |
||||
Gen. et sp. nov |
In press |
Wagner et al. |
A member of the family Agamidae. Genus includes new species P. monocoli. |
|||||
Sp. nov |
Valid |
Georgalis, Čerňanský & Klembara |
Probably Oligocene |
Quercy Phosphorites Formation |
A member of the family Lacertidae belonging to the subfamily Gallotiinae. |
|||
Gen. et sp. nov |
Valid |
Smith & Scanferla |
||||||
Gen. et sp. nov |
In press |
Longrich et al. |
Ouled Abdoun Basin |
A mosasaurine mosasaur. Genus includes new species X. calminechari. |
Research
- A study on the diversity of jaw sizes, lower jaw shape and morphology of teeth in Cretaceous squamates is published by Herrera-Flores, Stubbs & Benton (2021), who interpret their findings as indicating that a substantial expansion of ecomorphological diversity of squamates occurred in the mid-Cretaceous, 110–90 Ma, before the first rise in taxonomic diversity of this group in the Campanian.[12]
- A study on the dietary preferences of lizards from the Upper Cretaceous Iharkút vertebrate locality (Hungary) is published by Gere et al. (2021).[13]
- A study on the locomotion of five Cretaceous lizards, and on its implications for the knowledge of the ancestral locomotion type in lizards, is published by Villaseñor-Amador, Suárez & Cruz (2021), who interpret their findings as indicating that Huehuecuetzpalli mixtecus was bipedal while Tijubina pontei was facultatively bipedal.[14]
- A study on the fossil record of lizards and snakes from the Guadeloupe Islands, assessing their evolutionary history and diversity over the past 40,000 years, is published by Bochaton et al. (2021), who interpret their findings as indicative of a massive extinction of Guadeloupe’s snakes and lizards following European colonization, preceded by thousands of years of coexistence with earlier Indigenous populations.[15]
- A study the shape and size variation in the maxillae of extant New Zealand diplodactylids, and on its implications for the knowledge of the affinities of subfossil diplodactylid remains, is published by Scarsbrook et al. (2021).[16]
- Fossils of Gallotia goliath are described for the first time from the El Hierro island (Canary Islands) by Palacios-García et al. (2021), providing the first evidence of the possible coexistence of two giant fossil species of Gallotia on the same island.[17]
- Redescription and a study on the geographic provenance of the fossil material of "Liodon" asiaticum is published by Bardet et al. (2021).[18]
- One to seven month-long life histories of specimens of Platecarpus tympaniticus and Clidastes propython collected from chalk deposits of the Western Interior Seaway and Mississippi Embayment in Kansas and Alabama are reconstruted by Travis Taylor et al. (2021), who interpret their findings as indicative of semi-regular travels of these mosasaurs from marine to freshwater coastal environments and consumption of freshwater.[19]
Ichthyosauromorphs
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Fernández et al. |
An ichthyosaur belonging to the family Ophthalmosauridae. Genus includes new species C. gaspariniae. |
|||||
Gen. et sp. nov |
Valid |
Barrientos-Lara & Alvarado-Ortega |
An ichthyosaur belonging to the family Ophthalmosauridae. The type species is P. yacahuitztli |
Research
- A study on the anatomy of the skull of the holotype specimen of Besanosaurus leptorhynchus, description of additional specimens from the Middle Triassic Besano Formation (Italy/Switzerland) and a study on the phylogenetic relationships of this species is published by Bindellini et al. (2021), who interpret Mikadocephalus gracilirostris as a junior synonym of B. leptorhynchus.[22]
- New stenopterygiid ichthyosaur fossils, representing some of the best preserved Toarcian specimens from Europe (including a specimen with possible soft tissue preservation), are described from south-east France by Martin et al. (2021), who also attempt to determine the causes of the state of preservation of the studied specimens, and evaluate the implications of the study site for the knowledge of the environmental perturbations associated with the Toarcian Oceanic Anoxic Event.[23]
- A study on the anatomy of narial structures in Early Jurassic ichthyosaurs, and on their implications for the knowledge of the evolution of the bony subdivision of the external naris in more derived ichthyosaurs, is published by Massare, Wahl & Lomax (2021).[24]
- The first unambiguous ichthyosaur remains from Antarctica reported to date are described from the Upper Jurassic Ameghino (=Nordenskjöld) Formation by Campos et al. (2021), who also revise remains of two ichthyosaur specimens from the Upper Jurassic of Madagascar, and describe a third specimen which is the most complete ichthyosaur from this region of Gondwanaland.[25]
- A study aiming to determine whether Cretaceous ichthyosaur remains from Australia can be attributed to the species Platypterygius australis on the basis of vertebral data alone is published by Vakil, Webb & Cook (2021).[26]
Sauropterygians
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Campbell et al. |
An elasmosaurid plesiosaur. The type species is F. sloanae. |
|||||
Gen. et sp. nov |
Valid |
Puértolas-Pascual et al. |
An early member of Plesiosauroidea. The type species is P. moelensis. |
Research
- A study on the anatomy and replacement pattern of teeth in Henodus chelyops is published by Pommery et al. (2021).[29]
- Description of a new specimen of Panzhousaurus rotundirostris from the Guanling Formation (Guizhou, China), providing new information on the anatomy of this reptile, and a study on the phylogenetic relationships of this species is published by Lin et al. (2021).[30]
- Redescription of the anatomy of the skull of Thalassomedon haningtoni, and a study on the phylogenetic relationships of this species, is published by Sachs et al. (2021).[31]
- New information of the skeletal anatomy of Alexandronectes zealandiensis is provided by O'Gorman et al. (2021).[32]
- The most complete specimen of Kawanectes lafquenianum reported to date, providing new information on the anatomy of this plesiosaur, is described by O’Gorman (2021).[33]
Turtles
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Hirayama et al. |
||||||
Akoranemys[35] | Gen. et sp. nov | In press | Pérez-García | Late Cretaceous (Cenomanian) | Madagascar | A member of the family Bothremydidae belonging to the tribe Bothremydini and subtribe Bothremydina. Genus includes new species A. madagasika. | ||
Gen. et sp. nov |
In press |
Maniel, de la Fuente & Canale |
Late Cretaceous (Cenomanian) |
A member of the family Bothremydidae belonging to the tribe Cearachelyini. Genus includes new species E. pritchardi. |
||||
Gen. et sp. nov |
Valid |
De Lapparent de Broin, Breton & Rioult |
A member of the family Plesiochelyidae. Genus includes new species G. lennieri. |
|||||
Sp. nov |
Valid |
Lyson et al. |
A member of the family Baenidae. |
|||||
Sp. nov |
Valid |
Lyson, Petermann & Miller |
A soft-shelled turtle. |
|||||
Gen. et sp. nov |
Valid |
Adrian et al. |
Late Cretaceous (Cenomanian) |
A member of the family Bothremydidae. The type species is P. appalachius. |
||||
Sp. nov |
In press |
Pérez-García et al. |
Jurassic-Cretaceous transition |
|||||
Gen. et sp. nov |
Valid |
Joyce et al. |
Late Cretaceous (Maastrichtian) |
A member of Pleurodira belonging to the group Pelomedusoides. The type species is S. mailakavava. |
Research
- A study on the morphology of the skeleton of Chinlechelys tenertesta, and on its implications for the knowledge of the origin of turtles, is published by Lichtig & Lucas (2021), who reject the interpretation of Eunotosaurus africanus and Pappochelys rosinae as stem-turtles.[43]
- Reconstruction of the skull and neuroanatomical structures of Tartaruscola teodorii is presented by Martín-Jiménez & Pérez-García (2021).[44]
- Redescription of the anatomy of the skull of Arundelemys dardeni is published by Evers, Rollot & Joyce (2021).[45]
- A study on the morphological variability in the shell of Pleurosternon bullockii is published by Guerrero & Pérez-García (2021).[46]
- New fossil material of Plesiochelys is described from the Upper Jurassic (Tithonian) of the Lusitanian Basin (Portugal) by Pérez-García & Ortega (2021), who also revise the Oxfordian species "Hispaniachelys" prebetica and transfer it to the genus Plesiochelys, making it the oldest representative of this genus reported to date.[47]
- Two specimens of thalassochelydian turtles, including a partial hindlimb with well-preserved remains of skin and a large, articulated skeleton of Thalassemys bruntrutana, documenting the presence of particularly elongate forelimbs in this turtle, are described from the Upper Jurassic of Germany by Joyce, Mäuser & Evers (2021), who interpret these specimens as providing evidence of presence of highly keratinized and partially stiffened flippers in some Late Jurassic turtles, structurally similar to those of extant sea turtles.[48]
- Georgalis (2021) describes a large costal of a soft-shelled turtle from the Eocene of Mali, representing the first record of pan-trionychid turtles from the Paleogene of Africa reported to date.[49]
- New specimens of the tortoises Hadrianus majusculus and H. corsoni and geoemydids Echmatemys haydeni and E. naomi, including skull and juvenile material, and providing new information on the morphology, intraspecific variation and ontogeny of these turtles, are described from the Eocene Willwood Formation and Green River Formation (Wyoming, United States) by Lichtig, Lucas & Jasinski (2021).[50]
- A study on the phylogenetic relationships and evolutionary history of the tortoise Chelonoidis alburyorum from The Bahamas, based on data from nearly complete mitochondrial genomes, is published by Kehlmaier et al. (2021).[51]
- A well-preserved humerus a of giant turtle, representing the first known record of gigantic Mesozoic sea turtles in Africa, is described from the Maastrichtian Dakhla Formation (Egypt) by Abu El-Kheir, AbdelGawad & Kassab (2021).[52]
- New specimen of Euclastes wielandi is described from the Paleocene (Danian) Hornerstown Formation (New Jersey, United States) by Ullmann & Carr (2021), who interpret this specimen as proving that E. wielandi is a senior synonym of Catapleura repanda, and study the phylogenetic relationships of E. wielandi.[53]
Archosauriformes
Archosaurs
Other archosauriforms
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Ezcurra, Bandyopadhyay & Gower |
A member of the family Erythrosuchidae. The type species is B. tapani. |
|||||
Gen. et sp. nov |
Valid |
Nesbitt et al. |
A dome-headed archosauriform closely related to Triopticus, with which it forms the new clade Protopyknosia. The type species is K. kuttyi. |
|||||
Gen. et sp. nov |
Valid |
Heckert et al. |
A short-faced archosauriform of uncertain affinity, possibly an early-diverging crocodylomorph. The type species is S. sucherorum. |
Research
- A large phytosaur specimen likely belonging to the species Smilosuchus gregorii, preserved with evidence of pathologies evoking aspects of both osteomyelitis and hypertrophic osteopathy, is described from Norian strata near St. Johns (Arizona, United States) by Heckert, Viner & Carrano (2021).[57]
Other reptiles
New taxa
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Ezcurra et al. |
A rhynchosaur. Genus includes new species E. carrolli. |
|||||
Gen. et sp. nov |
Valid |
Cisneros et al. |
A member of the family Acleistorhinidae. Genus includes new species K. fortunata. |
|||||
Mengshanosaurus[60] | Gen. et sp. nov | Valid | Meng et al. | Early Cretaceous (Berriasian-Valanginian) | Mengyin Formation | China | A choristodere belonging to Neochoristodera, the type species is M. minimus. | |
Gen. et sp. nov |
Valid |
Pinheiro, Silva-Neves & Da-Rosa |
A procolophonid parareptile. Genus includes new species O. insolitus. |
|||||
Gen. et sp. nov |
Valid |
Villa et al. |
Late Jurassic (Kimmeridgian) |
A rhynchocephalian belonging to the family Sphenodontidae. The type species is S. velserae. |
||||
Gen. et sp. nov |
Valid |
Kligman et al. |
Late Triassic (Norian) |
A rhynchocephalian belonging to the group Opisthodontia. Genus includes new species T. purgatorii. |
||||
Gen. et sp. nov |
Valid |
Sues & Kligman |
Late Triassic (Carnian) |
Vinita Formation |
A member of Lepidosauromorpha. Genus includes new species V. lizae. |
Research
- Revision of putative Carboniferous reptile tracks, evaluating their implications for the knowledge of the evolution of locomotor capabilities of putative trackmakers and of the biogeography of the earliest reptiles, is published by Marchetti et al. (2021), who interpret Hylopus hardingi as tracks probably produced by anamniote reptiliomorphs.[65]
- A study on the anatomy of the skull of Nochelesaurus alexanderi is published by Van den Brandt et al. (2021).[66]
- A study on the anatomy of the postcranial skeletons of Bradysaurus baini, Embrithosaurus schwarzi and Nochelesaurus alexanderi is published by van den Brandt et al. (2021).[67]
- Romano et al. (2021) provide a body mass estimate of Scutosaurus karpinskii.[68]
- Tetrapod footprints assigned to the ichnogenus Hyloidichnus, produced by captorhinids or similar reptiles and providing new information on the locomotion of these reptiles, are described from the Permian Pelitic Formation of Gonfaron (Le Luc Basin, Var, France) by Logghe et al. (2021).[69]
- Description of a new specimen of Anthracodromeus longipes from the Carboniferous Allegheny Group (Ohio, United States), preserving an ungual-bearing manus and pedes, and a study on the locomotor ecology of this reptile is published by Mann et al. (2021).[70]
- A study on the skeletal anatomy, gliding apparatus and phylogenetic relationships of Weigeltisaurus jaekeli, based on data from a nearly complete skeleton from the Upper Permian Kupferschiefer (Germany), is published by Pritchard et al. (2021).[71]
- Putative choristoderan maxilla from the Bathonian Peski locality (Moskvoretskaya Formation; Moscow Oblast, Russia) is reinterpreted as a fossil a member of Lepidosauromorpha similar to Fraxinisaura and Marmoretta by Skutschas et al. (2021), who interpret this specimens as the first known record of basal lepidosauromorphs in the Middle Jurassic of European Russia.[72]
- New fossil material of neochoristoderes, representing the first known record of this group from the Atlantic coastal plain, is described from the latest Cretaceous of the Navesink Formation and the Ellisdale Fossil Site (New Jersey, United States) by Dudgeon et al. (2021), who also attempt to determine possible causes of the apparent rarity of neochoristoderes in Appalachia.[73]
- A study on the phylogenetic relationships of archosauromorph reptiles, aiming to determine the relationships of "protorosaurs", is published by Spiekman, Fraser & Scheyer (2021), who name a new clade Dinocephalosauridae.[74]
- A study on patterns of neurocentral suture closure in the vertebrae of hyperodapedontine rhynchosaurs during their ontogeny, evaluating their implications for the knowledge of the evolution of patterns of neurocentral suture closure in archosauromorph reptiles and most likely ancestral condition in archosaurs, is published by Heinrich et al. (2021).[75]
Reptiles in general
- A study comparing the morphology of the maxillary canal of Heleosaurus scholtzi, Varanosaurus acutrostris, Orovenator mayorum and Prolacerta broomi, and evaluating the implications of the morphology of the maxillary canal for the knowledge of the phylogenetic placement of varanopids, is published by Benoit et al. (2021).[76]
- Fischer, Weis & Thuy (2021) describe new ichthyosaur and plesiosaur fossils from successive geological formations in Belgium and Luxembourg spanning the Lower–Middle Jurassic transition, and evaluate the implications of these fossils for the knowledge of the evolution of marine reptile assemblages across the Early–Middle Jurassic transition.[77]
References
- ^ Rage, J.-C.; Adaci, M.; Bensalah, M.; Mahboubi, M.; Marivaux, L.; Mebrouk, F.; Tabuce, R. (2021). "Latest Early-early Middle Eocene deposits of Algeria (Glib Zegdou, HGL50), yield the richest and most diverse fauna of amphibians and squamate reptiles from the Palaeogene of Africa" (PDF). Palæovertebrata. 44 (1): e1. doi:10.18563/pv.44.1.e1.
- ^ Scarpetta, S. G. (2021). "Iguanian lizards from the Split Rock Formation, Wyoming: exploring the modernization of the North American lizard fauna". Journal of Systematic Palaeontology. 19 (3): 221–251. doi:10.1080/14772019.2021.1894612. S2CID 233402435.
- ^ Bochaton, C.; Charles, L.; Lenoble, A. (2021). "Historical and fossil evidence of an extinct endemic species of Leiocephalus (Squamata: Leiocephalidae) from the Guadeloupe Islands". Zootaxa. 4927 (3): 383–409. doi:10.11646/zootaxa.4927.3.4. PMID 33756701.
- ^ Bolet, A.; Stanley, E. L.; Daza, J. D.; Arias, J. S.; Čerňanský, A.; Vidal-García, M.; Bauer, A. M.; Bevitt, J. J.; Peretti, A.; Evans, S. E. (2021). "Unusual morphology in the mid-Cretaceous lizard Oculudentavis". Current Biology. Online edition. doi:10.1016/j.cub.2021.05.040. PMID 34129826.
- ^ a b Georgalis, G. L.; Čerňanský, A.; Klembara, J. (2021). "Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material". Geodiversitas. 43 (9): 219–293. doi:10.5252/geodiversitas2021v43a9.
- ^ Smith, K. T.; Habersetzer, J. (2021). "The anatomy, phylogenetic relationships, and autecology of the carnivorous lizard "Saniwa" feisti Stritzke, 1983 from the Eocene of Messel, Germany". Comptes Rendus Palevol. 20 (23): 441–506. doi:10.5852/cr-palevol2021v20a23.
- ^ Longrich, N. R.; Bardet, N.; Khaldoune, F.; Khadiri Yazami, O.; Jalil, N.-E. (2021). "Pluridens serpentis, a new mosasaurid (Mosasauridae: Halisaurinae) from the Maastrichtian of Morocco and implications for mosasaur diversity". Cretaceous Research. 126: Article 104882. doi:10.1016/j.cretres.2021.104882.
- ^ Thorn, K. M.; Hutchinson, M. N.; Lee, M. S. Y.; Brown, N. J.; Camens, A. B.; Worthy, T. H. (2021). "A new species of Proegernia from the Namba Formation in South Australia and the early evolution and environment of Australian egerniine skinks". Royal Society Open Science. 8 (2): 201686. Bibcode:2021RSOS....801686T. doi:10.1098/rsos.201686. ISSN 2054-5703. PMC 8074667. PMID 33972861.
- ^ Wagner, P.; Stanley, E. L.; Daza, J. D.; Bauer, A. M. (2021). "A new agamid lizard in mid-Cretaceous amber from northern Myanmar". Cretaceous Research. 124: Article 104813. doi:10.1016/j.cretres.2021.104813.
- ^ Smith, K. T.; Scanferla, A. (2021). "A nearly complete skeleton of the oldest definitive erycine boid (Messel, Germany)". Geodiversitas. 43 (1): 1–24. doi:10.5252/geodiversitas2021v43a1.
- ^ Longrich, N. R.; Bardet, N.; Schulp, A. S.; Jalil, N.-E. (2021). "Xenodens calminechari gen. et sp. nov., a bizarre mosasaurid (Mosasauridae, Squamata) with shark-like cutting teeth from the upper Maastrichtian of Morocco, North Africa". Cretaceous Research. 123: Article 104764. doi:10.1016/j.cretres.2021.104764.
- ^ Herrera-Flores, J. A.; Stubbs, T. L.; Benton, M. J. (2021). "Ecomorphological diversification of squamates in the Cretaceous". Royal Society Open Science. 8 (3): Article ID 201961. Bibcode:2021RSOS....801961H. doi:10.1098/rsos.201961. PMC 8074880. PMID 33959350.
- ^ Gere, K.; Bodor, E. R.; Makádi, L.; Ősi, A. (2021). "Complex food preference analysis of the Late Cretaceous (Santonian) lizards from Iharkút (Bakony Mountains, Hungary)". Historical Biology: An International Journal of Paleobiology. in press: 1–17. doi:10.1080/08912963.2021.1887862.
- ^ Villaseñor-Amador, D.; Suárez, N. X.; Cruz, J. A. (2021). "Bipedalism in Mexican Albian lizard (Squamata) and the locomotion type in other Cretaceous lizards". Journal of South American Earth Sciences. 109: Article 103299. Bibcode:2021JSAES.10903299V. doi:10.1016/j.jsames.2021.103299.
- ^ Bochaton, C.; Paradis, E.; Bailon, S.; Grouard, S.; Ineich, I.; Lenoble, A.; Lorvelec, O.; Tresset, A.; Boivin, N. (2021). "Large-scale reptile extinctions following European colonization of the Guadeloupe Islands". Science Advance. 7 (21): eabg2111. Bibcode:2021SciA....7G2111B. doi:10.1126/sciadv.abg2111. PMC 8133755. PMID 34138736.
- ^ Scarsbrook, L.; Sherratt, E.; Hitchmough, R. A.; Rawlence, N. J. (2021). "Skeletal variation in extant species enables systematic identification of New Zealand's large, subfossil diplodactylids". BMC Ecology and Evolution. 21 (1): Article number 67. doi:10.1186/s12862-021-01808-7. PMC 8080345. PMID 33906608.
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