2020 in archosaur paleontology: Difference between revisions

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* A study on the trace elements and isotopic compositions of eggshells of dinosaur eggs from the Cretaceous [[Zhaoying Formation]] ([[Henan]], [[China]]), evaluating their implications for reconstructions of local paleoenvironment, is published by He ''et al.'' (2020).<ref>{{Cite journal|author1=Qing He |author2=Sen Yang |author3=Songhai Jia |author4=Li Xu |author5=Lida Xing |author6=Diansong Gao |author7=Di Liu |author8=Yongli Gao |author9=Yalin Zheng |year=2020 |title=Trace element and isotope geochemistry of macroelongatoolithid eggs as an indicator of palaeoenvironmental reconstruction from the Late Cretaceous Xixia Basin, China |journal=Cretaceous Research |volume=109 |pages=Article 104373 |doi=10.1016/j.cretres.2020.104373 }}</ref>
* A study on the trace elements and isotopic compositions of eggshells of dinosaur eggs from the Cretaceous [[Zhaoying Formation]] ([[Henan]], [[China]]), evaluating their implications for reconstructions of local paleoenvironment, is published by He ''et al.'' (2020).<ref>{{Cite journal|author1=Qing He |author2=Sen Yang |author3=Songhai Jia |author4=Li Xu |author5=Lida Xing |author6=Diansong Gao |author7=Di Liu |author8=Yongli Gao |author9=Yalin Zheng |year=2020 |title=Trace element and isotope geochemistry of macroelongatoolithid eggs as an indicator of palaeoenvironmental reconstruction from the Late Cretaceous Xixia Basin, China |journal=Cretaceous Research |volume=109 |pages=Article 104373 |doi=10.1016/j.cretres.2020.104373 }}</ref>
* A study on the affinities of putative [[gekkota]]n eggshells from the Late Cretaceous of Europe is published by Choi ''et al.'' (2020), who interpret the fossil material of ''[[Pseudogeckoolithus]]'' as theropod eggshells.<ref>{{Cite journal|author1=Seung Choi |author2=Miguel Moreno‐Azanza |author3=Zoltán Csiki‐Sava |author4=Edina Prondvai |author5=Yuong‐Nam Lee |year=2020 |title=Comparative crystallography suggests maniraptoran theropod affinities for latest Cretaceous European 'geckoid' eggshell |journal=Papers in Palaeontology |volume=in press |issue= |pages= |doi=10.1002/spp2.1294 }}</ref>
* A study on the affinities of putative [[gekkota]]n eggshells from the Late Cretaceous of Europe is published by Choi ''et al.'' (2020), who interpret the fossil material of ''[[Pseudogeckoolithus]]'' as theropod eggshells.<ref>{{Cite journal|author1=Seung Choi |author2=Miguel Moreno‐Azanza |author3=Zoltán Csiki‐Sava |author4=Edina Prondvai |author5=Yuong‐Nam Lee |year=2020 |title=Comparative crystallography suggests maniraptoran theropod affinities for latest Cretaceous European 'geckoid' eggshell |journal=Papers in Palaeontology |volume=in press |issue= |pages= |doi=10.1002/spp2.1294 }}</ref>
* Chapelle, Fernandez & Choiniere (2020) evaluate the possibility of estimating the developmental stage of dinosaur embryos, on the basis of a study of skull ossification sequences in embryos of ''[[Massospondylus]] carinatus'' and extant [[sauria]]ns.<ref>{{Cite journal|author1=Kimberley E. J. Chapelle |author2=Vincent Fernandez |author3=Jonah N. Choiniere |year=2020 |title=Conserved in-ovo cranial ossification sequences of extant saurians allow estimation of embryonic dinosaur developmental stages |journal=Scientific Reports |volume=10 |issue=1 |pages=Article number 4224 |doi=10.1038/s41598-020-60292-z }}</ref>
* The discovery of [[sternal plates]] of ''[[Tawa hallae]]'' from the [[Late Triassic]] of [[New Mexico]] and [[Arizona]], representing the oldest known dinosaur sternal plates described so far, is reported by Bradley ''et al.'' (2020), who note the presence of morphological features similar to sternal traits in [[Avialae|avialans]].<ref>{{Cite journal|author1=Alexander B. Bradley |author2=Sara H. Burch |author3=Alan H. Turner |author4=Nathan D. Smith |author5=Randall B. Irmis |author6=Sterling J. Nesbitt |year=2020 |title=Sternal elements of early dinosaurs fill a critical gap in the evolution of the sternum in Avemetatarsalia (Reptilia: Archosauria) |journal=Journal of Vertebrate Paleontology |volume=39 |issue=5 |pages=e1700992 |doi=10.1080/02724634.2019.1700992 }}</ref>
* The discovery of [[sternal plates]] of ''[[Tawa hallae]]'' from the [[Late Triassic]] of [[New Mexico]] and [[Arizona]], representing the oldest known dinosaur sternal plates described so far, is reported by Bradley ''et al.'' (2020), who note the presence of morphological features similar to sternal traits in [[Avialae|avialans]].<ref>{{Cite journal|author1=Alexander B. Bradley |author2=Sara H. Burch |author3=Alan H. Turner |author4=Nathan D. Smith |author5=Randall B. Irmis |author6=Sterling J. Nesbitt |year=2020 |title=Sternal elements of early dinosaurs fill a critical gap in the evolution of the sternum in Avemetatarsalia (Reptilia: Archosauria) |journal=Journal of Vertebrate Paleontology |volume=39 |issue=5 |pages=e1700992 |doi=10.1080/02724634.2019.1700992 }}</ref>
* New fossil material of theropod dinosaurs representing a wide taxonomic range is reported from the [[Late Jurassic]] of the Langenberg Quarry ([[Lower Saxony]], [[Germany]]) by Evers & Wings (2020), who interpret these fossils as evidence of the presence of several taxa of theropods in the Late Jurassic archipelago in the area of Central Europe.<ref>{{Cite journal|author1=Serjoscha W. Evers |author2=Oliver Wings |year=2020 |title=Late Jurassic theropod dinosaur bones from the Langenberg Quarry (Lower Saxony, Germany) provide evidence for several theropod lineages in the central European archipelago |journal=PeerJ |volume=8 |pages=e8437 |doi=10.7717/peerj.8437 |pmid=32071804 |pmc=7007975 }}</ref>
* New fossil material of theropod dinosaurs representing a wide taxonomic range is reported from the [[Late Jurassic]] of the Langenberg Quarry ([[Lower Saxony]], [[Germany]]) by Evers & Wings (2020), who interpret these fossils as evidence of the presence of several taxa of theropods in the Late Jurassic archipelago in the area of Central Europe.<ref>{{Cite journal|author1=Serjoscha W. Evers |author2=Oliver Wings |year=2020 |title=Late Jurassic theropod dinosaur bones from the Langenberg Quarry (Lower Saxony, Germany) provide evidence for several theropod lineages in the central European archipelago |journal=PeerJ |volume=8 |pages=e8437 |doi=10.7717/peerj.8437 |pmid=32071804 |pmc=7007975 }}</ref>

Revision as of 10:14, 9 April 2020

List of years in archosaur paleontology
In paleontology
2017
2018
2019
2020
2021
2022
2023
In science
2017
2018
2019
2020
2021
2022
2023
+...

This article records new taxa of fossil archosaurs of every kind that are scheduled described during the year 2020, as well as other significant discoveries and events related to paleontology of archosaurs that are scheduled to occur in the year 2020.

General research

  • A study on the evolution of metabolic rates along the bird stem lineage is published by Rezende et al. (2020).[1]
  • A review of the anatomy of the respiratory systems and mechanics of breathing in living and fossil archosaurs, evaluating their physiological implications, is published by Brocklehurst et al. (2020).[2]
  • A study aiming to determine the relationship between atmospheric O2 and CO2 levels during the Late Triassic and the evolution of skeletal pneumaticity and respiratory systems in theropod dinosaurs and in paracrocodylomorphs is published by Hudgins, Uhen & Hinnov (2020).[3]
  • A study on the phylogenetic distribution and structural diversity of medullary bone in extant birds, reevaluating the criteria proposed to allow the identification of medullary bone in fossils of avemetatarsalians, is published by Canoville, Schweitzer & Zanno (2020).[4]
  • An archosaur egg of uncertain affinities, with eggshell containing several parallel dark bands, is reported from the Upper Cretaceous of South Korea by Choi et al. (2020), who investigate the origin of the dark bands, and name a new ootaxon Aenigmaoolithus vesicularis.[5]
  • A study on the relationship between the curvatures of ungual bones and behaviour in extant birds and squamates, evaluating its implications for the knowledge of the lifestyle of Mesozoic birds and non-avian theropods, is published by Cobb & Sellers (2020).[6]
  • Xing, Cockx & McKellar (2020) describe a large sample set of 150 specimens of the Cretaceous Burmese amber containing feathers most likely belonging to non-avian dinosaurs and enantiornithean birds.[7]

Pseudosuchians

Research

  • Redescription of the anatomy of the postcranial skeleton of Riojasuchus tenuisceps, and a study on the phylogenetic affinities of ornithosuchids, is published by von Baczko, Desojo & Ponce (2020).[8]
  • A three-dimensional reconstruction of the armour plates around the tail of Stagonolepis robertsoni is presented by Keeble & Benton (2020).[9]
  • Taxonomic revision, anatomical description, and a study on the phylogenetic relationships of the type and referred materials of Prestosuchus from the original collections of Friedrich von Huene is published by Desojo, von Baczko & Rauhut (2020), who transfer the species Stagonosuchus nyassicus to the genus Prestosuchus.[10]
  • A study on the anatomy of the braincase of Almadasuchus figarii, and on early evolution of cranial pneumaticity in Crocodylomorpha, is published by Leardi, Pol & Clark (2020).[11]
  • A study on the impact of the habitat on the evolution of body size in Crocodyliformes, based on data from extant and fossil taxa, is published by Gearty & Payne (2020).[12]
  • New fossil material of crocodylomorphs from the Birket Qarun Formation in the Fayum Depression (Egypt), including the first record of a sebecosuchian from the late Eocene of Africa, is described by Stefanic et al. (2020).[13]
  • A study on the thermophysiology of metriorhynchids, as indicated by the oxygen isotope composition of the tooth enamel phosphate, is published by Séon et al. (2020).[14]
  • Fossil material of two large-bodied metriorhynchids is reported from lower Kimmeridgian sediments in Bavaria and Baden-Württemberg (Germany) by Abel, Sachs & Young (2020), who interpret these fossils as evidence of a new lineage of large-bodied geosaurines from the Kimmeridgian and Tithonian of Europe.[15]
  • A study on the anatomy and biomechanics of baurusuchid skulls, evaluating their implications for the knowledge of likely predatory behaviors of baurusuchids, is published by Montefeltro et al. (2020).[16]
  • New information on the anatomy of the endocranial cavities of Campinasuchus dinizi is presented by Fonseca et al. (2020).[17]
  • New specimen of Susisuchus anatoceps, displaying a non-eusuchian type palate (i.e. choana not entirely bounded by the pterygoids), is described by Montefeltro et al. (2020), who evaluate the implications of this finding for the knowledge of the anatomy of this taxon and the phylogenetic position of susisuchids.[18]
  • A study on skull anatomy and phylogenetic relationships of Bernissartia fagesii is published by Martin et al. (2020).[19]
  • Reconstruction of the internal cavities of the skull of Agaresuchus fontisensis, including the cavities that contained the brain, nerves and blood vessels, is presented by Serrano‐Martínez et al. (2020).[20]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Bottosaurus fustidens [21]

Sp. nov

Valid

Cossette

Paleocene (Tiffanian)

Black Peaks Formation

 United States
( Texas)

A caiman.

Dynamosuchus [22]

Gen. et sp. nov

Valid

Müller et al.

Late Triassic (Carnian)

Santa Maria

 Brazil

A member of the family Ornithosuchidae. The type species is D. collisensis.

Non-avialan dinosaurs

Research

  • A study on the biogeography of the Cretaceous Australian dinosaur fauna is published by Kubo (2020).[23]
  • A study on the evolutionary history of dinosaur integument, aiming to determine the most likely ancestral integumentary condition in dinosaurs, is published by Campione, Barrett & Evans (2020).[24]
  • A study aiming to determine dinosaur body temperatures on the basis of data from fossil eggshells, comparing them with paleoenvironmental temperatures, and evaluating their implications for the knowledge of dinosaur thermoregulation, is published by Dawson et al. (2020).[25]
  • A study on the trace elements and isotopic compositions of eggshells of dinosaur eggs from the Cretaceous Zhaoying Formation (Henan, China), evaluating their implications for reconstructions of local paleoenvironment, is published by He et al. (2020).[26]
  • A study on the affinities of putative gekkotan eggshells from the Late Cretaceous of Europe is published by Choi et al. (2020), who interpret the fossil material of Pseudogeckoolithus as theropod eggshells.[27]
  • Chapelle, Fernandez & Choiniere (2020) evaluate the possibility of estimating the developmental stage of dinosaur embryos, on the basis of a study of skull ossification sequences in embryos of Massospondylus carinatus and extant saurians.[28]
  • The discovery of sternal plates of Tawa hallae from the Late Triassic of New Mexico and Arizona, representing the oldest known dinosaur sternal plates described so far, is reported by Bradley et al. (2020), who note the presence of morphological features similar to sternal traits in avialans.[29]
  • New fossil material of theropod dinosaurs representing a wide taxonomic range is reported from the Late Jurassic of the Langenberg Quarry (Lower Saxony, Germany) by Evers & Wings (2020), who interpret these fossils as evidence of the presence of several taxa of theropods in the Late Jurassic archipelago in the area of Central Europe.[30]
  • New theropod fossil material is reported from the Griman Creek Formation by Brougham, Smith & Bell (2020), who interpret it as evidence of the presence of noasaurids in Australia during the Cretaceous.[31]
  • A study on the anatomy of teeth of Sinraptor dongi, comparing it with dentition of other theropods and evaluating its implications for the knowledge of the feeding ecology of S. dongi, is published by Hendrickx et al. (2020).[32]
  • A study on an indeterminate megaraptoran specimen from the Winton Formation (Australia) is published by White et al. (2020), who interpret this finding as evidence of either ontogenetic or intraspecific variation in Australovenator, or the presence of a second megaraptorid taxon in the Winton Formation.[33]
  • A study on the endocranial anatomy of Bistahieversor sealeyi, evaluating its implications for the knowledge of the evolution of the brains and sinuses of tyrannosauroids, is published by McKeown et al. (2020).[34]
  • A study on the bone microstructure of two half-grown specimens of Tyrannosaurus rex, evaluating its implications for the knowledge of the early life history of members of this species and the taxonomic validity of Nanotyrannus lancensis, is published by Woodward et al. (2020).[35]
  • A study on the anatomy of the integumentary structures of Juravenator starki and Sciurumimus albersdoerferi is published by Foth et al. (2020).[36]
  • New fossil material of Chirostenotes pergracilis, representing the first associated mandibular and postcranial material of a caenagnathid from the Dinosaur Park Formation (Alberta, Canada), is described by Funston & Currie (2020), who evaluate the implications of these fossils for the knowledge of taxonomy and diversity of caenagnathids from the Dinosaur Park Formation and the growth patterns of Chirostenotes pergracilis.[37]
  • Description of a partial skeleton of a caenagnathid theropod from the Upper Cretaceous Hell Creek Formation (Montana, United States) and study on the bone histology of this specimen is published by Cullen et al. (2020), who evaluate the implications of their findings for the knowledge of the utility of size as a determinant for referral of incomplete or fragmentary skeletal remains to specific or new coelurosaur taxa.[38]
  • New theropod teeth, possibly belonging to members of the family Dromaeosauridae and representing the first record of that group from the southern Junggar Basin, are reported from the Upper Jurassic Qigu Formation (China) by Maisch & Matzke (2020).[39]
  • A study on the differences in the locomotor and predatory specializations of eudromaeosaurs and unenlagiines, as indicated by the anatomy of their hindlimbs, is published by Gianechini, Ercoli & Díaz‐Martínez (2020).[40]
  • A study on eudromaeosaurian maxillae, aiming to determine the extent to which maxillae can be used to draw ecological and phylogenetic inferences about dromaeosaurids, is published by Powers, Sullivan & Currie (2020).[41]
  • A study on the histology of the humeri of two basal sauropod specimens from the Jurassic of Niger and Thailand, reporting evidence of a layer of the radial fibrolamellar bone buried in the outer cortex of these bones, is published by Jentgen-Ceschino, Stein & Fischer (2020), who interpret their findings as evidence of these sauropods being affected by pathologies similar to Ewing's sarcoma and avian osteopetrosis or haemangioma.[42]
  • A study comparing articulation and range of motion of necks of extant giraffes and Spinophorosaurus nigerensis is published by Vidal et al. (2020).[43]
  • A review of the distribution of the Cretaceous fossils of rebbachisaurid sauropods is published by Pereira et al. (2020), who report the first occurrence of a rebbachisaurid from the Açu Formation (Potiguar Basin, Brazil), and discuss its paleobiogeographic implications.[44]
  • A large sauropod humerus, probably belonging to a member of the species Fusuisaurus zhaoi, is described from the Lower Cretaceous Xinlong Formation (Guangxi, China) by Mo et al. (2020).[45]
  • A study on histology and affinities of two bone fragments from the Upper Cretaceous (lower Santonian to/or lower Campanian) of the Western Srednogorie (Bulgaria) is published by Nikolov et al. (2020), who interpret these fossils as bones of a titanosaur sauropod, coming from a time interval when sauropods are rare in the fossil record of Europe.[46]
  • Voegele et al. (2020) reconstruct the forelimb and shoulder girdle musculature of Dreadnoughtus schrani.[47]
  • A study on the microstructure of the tooth enamel of Manidens condorensis, evaluating its implications for the knowledge of the evolution of tooth enamel in Ornithischia, is published by Becerra & Pol (2020).[48]
  • A study on the structure and development of the dermal skeleton of Scelidosaurus harrisonii is published by Norman (2020).[49]
  • A study on the bone microstructure of Mongolian hadrosauroid dinosaurs, evaluating its implications for the knowledge of growth strategies and evolution of gigantism in hadrosauroids, is published by Słowiak et al. (2020).[50]
  • A study on pathologies affecting two hadrosaurid vertebrae from the Dinosaur Provincial Park (Alberta, Canada) is published by Rothschild et al. (2020), who consider Langerhans cell histiocytosis to be the most likely diagnosis, making it the first case of LCH recognized in a dinosaur so far.[51]
  • A study on the migratory behaviours of hadrosaurs, as indicated by strontium isotope data from hadrosaur teeth from the Late Cretaceous of Alberta (Canada), is published by Terrill, Henderson & Anderson (2020).[52]
  • A study on the interior structure of the nasal spine of Tsintaosaurus spinorhinus is published by Zhang et al. (2020).[53]
  • Evidence of preservation of proteins, chromosomes and chemical markers of DNA in the cartilage of a nestling of Hypacrosaurus stebingeri from the Campanian Two Medicine Formation (Montana, United States) is presented by Bailleul et al. (2020).[54]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Abdarainurus[55]

Gen. et sp. nov

Valid

Averianov & Lopatin

Late Cretaceous

Alagteeg

 Mongolia

A sauropod dinosaur, probably a basal member of Titanosauria. Genus includes new species A. barsboldi.

Adratiklit[56]

Gen. et sp. nov

Valid

Maidment et al.

Middle Jurassic (Bathonian)

El Mers II

 Morocco

A member of Stegosauria. Genus includes new species A. boulahfa. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Allosaurus jimmadseni[57]

Sp. nov

Valid

Chure & Loewen

Late Jurassic (Kimmeridgian)

Morrison

 United States
( Colorado
 Utah
 Wyoming)

Amanzia[58]

Gen. et comb. nov

Schwarz et al.

Late Jurassic (Kimmeridgian)

Reuchenette

  Switzerland

A non-neosauropod eusauropod of uncertain phylogenetic placement. The type species is "Ornithopsis" greppini Huene (1922).

Analong[59]

Gen. et sp. nov

In press

Ren et al.

Middle Jurassic

Chuanjie Formation

 China

A mamenchisaurid sauropod. Genus includes new species A. chuanjieensis.

Dineobellator[60]

Gen. et sp. nov

Valid

Jasinski, Sullivan & Dodson

Late Cretaceous (Maastrichtian)

Ojo Alamo

 United States
( New Mexico)

A dromaeosaurid theropod. The type species is D. notohesperus.

Huinculsaurus[61]

Gen. et sp. nov

Valid

Baiano, Coria & Cau

Late Cretaceous (late Cenomanian-Turonian)

Huincul

 Argentina

A theropod related to Elaphrosaurus. Genus includes new species H. montesi.

Jinbeisaurus[62]

Gen. et sp. nov

Valid

Wu et al.

Late Cretaceous

Huiquanpu

 China

A tyrannosauroid theropod. Genus includes new species J. wangi. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Lajasvenator[63]

Gen. et sp. nov

Valid

Coria et al.

Early Cretaceous (Valanginian)

Mulichinco Formation

 Argentina

A carcharodontosaurid theropod. Genus includes new species L. ascheriae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Omeisaurus puxiani[64]

Sp. nov

In press

Tan et al.

Middle Jurassic

Shaximiao Formation

 China

A mamenchisaurid sauropod.

Thanatotheristes[65]

Gen. et sp. nov

Valid

Voris et al.

Late Cretaceous (Campanian)

Foremost

 Canada
( Alberta)

A tyrannosaurid theropod. Genus includes new species T. degrootorum.

Tralkasaurus[66]

Gen. et sp. nov

Valid

Cerroni et al.

Late Cretaceous (Cenomanian-Turonian)

Huincul

 Argentina

An abelisaurid theropod. Genus includes new species T. cuyi. Announced in 2019; the final version of the article naming was published in 2020.

Vallibonavenatrix[67]

Gen. et sp. nov

Valid

Malafaia et al.

Early Cretaceous (Barremian)

Arcillas de Morella

 Spain

A spinosaurid theropod. Genus includes new species V. cani. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Wulong[68]

Gen. et sp. nov

Valid

Poust et al.

Early Cretaceous (Aptian)

Jiufotang

 China

A microraptorine dromaeosaurid theropod. Genus includes new species W. bohaiensis.

Xunmenglong[69]

Gen. et sp. nov

Valid

Xing et al.

Early Cretaceous

Huajiying

 China

A compsognathid theropod. Genus includes new species X. yinliangis. Announced in 2019; the final version of the article naming it was published in 2020.

Yamanasaurus[70]

Gen. et sp. nov

Valid

Apesteguía et al.

Late Cretaceous

Río Playas

 Ecuador

A saltasaurine titanosaur. Genus includes new species Y. lojaensis. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Yunyangosaurus[71]

Gen. et sp. nov

Dai et al.

Middle Jurassic

Xintiangou

 China

A tetanuran theropod, possibly a member of Megalosauroidea. The type species is Y. puanensis.

Birds

Research

  • A study aiming to determine the volumes of the brain structures used to infer behavior or functional capabilities in Archaeopteryx lithographica, Lithornis plebius, Dinornis robustus, Paraptenodytes antarcticus, Psilopterus lemoinei, Llallawavis scagliai and an unnamed Miocene galliform is published by Early, Ridgely & Witmer (2020).[72]
  • A feather fragment from an aquatic bird is reported from amber recovered from the Pipestone Creek bonebed from the Campanian Wapiti Formation (Alberta, Canada) by Cockx et al. (2020).[73]
  • A study on the life history of the elephant birds, as indicated by bone histology, is published by Chinsamy et al. (2020).[74]
  • Volkova & Zelenkov (2020) describe new fossil material of geese from the late Miocene locality Khyargas Nuur 2 in western Mongolia, and evaluate the implications of these fossils for the knowledge of the late Miocene evolution and paleogeography of geese.[75]
  • A coracoid of a small-bodied paraortygid is reported from the Uinta Formation (Utah, United States) by Stidham, Townsend & Holroyd (2020), representing the only known pangalliform from the middle Eocene of North America, occurring in a temporal gap in their history between the early Eocene Gallinuloides wyomingensis and late Eocene Nanortyx inexpectus.[76]
  • Barton et al. (2020) reinterpret purported chicken specimens from the Neolithic site at Dadiwan as remains of pheasants, and argue that these remains provide evidence of exploitation of grain-fed pheasants by early farmers in arid northwest China.[77]
  • A study on the morphological diversity of bills of extant and fossil penguins, and its relationship with feeding habits, is published by Chávez-Hoffmeister (2020).[78]
  • Partial skeleton of an early penguin (possibly belonging to the species Muriwaimanu tuatahi), preserving the first complete wing of a Paleocene penguin reported so far and providing new information on the skeletal anatomy of this taxon, is described from the Waipara Greensand (New Zealand) by Mayr et al. (2020).[79]
  • An articulated wing of Palaeeudyptes gunnari, preserving mineralized skin, is described from the Eocene (Lutetian) of Seymour Island (Antarctica) by Acosta Hospitaleche et al. (2020).[80]
  • New fossil material of penguins and a member of Gruiformes is reported from the Eocene of the Seymour Island by Davis et al. (2020), supporting previously controversial reports of Gruiformes from Antarctica.[81]
  • New fossil material of larks is reported from the late Pliocene localities in Transbaikalia (Russia) and Mongolia by Palastrova & Zelenkov (2020), who transfer the species Pliocalcarius orkhonensis to the genus Eremophila, and evaluate the implications of their findings for the knowledge of the evolutionary history of larks.[82]
  • An exceptionally well-preserved bird carcass found in the Siberian permafrost and dated to approximately 44–49 ka BP is described by Dussex et al. (2020), who identify this specimen as a female horned lark, and evaluate the implications of this specimen for the knowledge of the evolution and biogeography of its species during the Pleistocene.[83]
  • Flamingo-like and anatid-like fossil bird footprints will be described from the Vinchina Formation (Argentina) by Farina et al. (2020), who name new ichnotaxa Phoenicopterichnum lucioi and P. vinchinaensis.[84]
  • A study on the impact of the climate changes of the last 35,000 years on small birds from the La Brea Tar Pits is published by Long, Prothero & Syverson (2020).[85]
  • A study comparing predicted breeding and wintering distributions for landbird species identified from the La Brea Tar Pits during the Last Glacial Maximum, aiming to determine if niche models successfully predict species’ presence, to estimate the degree of species turnover, to evaluate the fluidity of life history strategies of birds from La Brea, and to compare niche breadths of bark-foraging birds from La Brea between the Last Glacial Maximum and the present, is published by Zink et al. (2020).[86]
  • New fossil material of seabirds, including remains of the little auk or a related species, is reported from the Pleistocene Kazusa and Shimosa groups (Japan) by Watanabe et al. (2020), who interpret this finding as possible evidence that the little auk more widespread in the North Pacific in the middle Pleistocene than it is today.[87]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Antarcticavis[88]

Gen. et sp. nov

In press

Cordes-Person et al.

Late Cretaceous (Maastrichtian)

Snow Hill Island

Antarctica

A bird of uncertain phylogenetic placement, possibly a member of Ornithuromorpha belonging to the group Ornithurae. The type species is A. capelambensis. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Asio ecuadoriensis[89]

Sp. nov

In press

Lo Coco, Agnolín & Carrión

Late Pleistocene

 Ecuador

An owl, a species of Asio.

Asteriornis[90]

Gen. et sp. nov

Valid

Field et al.

Late Cretaceous (Maastrichtian)

Maastricht

 Belgium

An early member of Neornithes, occupying a position close to the last common ancestor of Galloanserae. Genus includes new species A. maastrichtensis.

?Crossvallia waiparensis[91]

Sp. nov

Valid

Mayr et al.

Paleocene

Waipara

 New Zealand

A large-sized penguin. Announced in 2019; the final version of the article naming it was published in 2020.

Khinganornis[92]

Gen. et sp. nov

In press

Wang et al.

Early Cretaceous (Aptian)

Longjiang

 China

A derived member of Ornithuromorpha. Genus includes the new species K. hulunbuirensis.

Linxiavis[93] Gen. et sp. nov Li et al. Late Miocene Liushu Formation  China A sandgrouse. The type species is L. inaquosus.

Oculudentavis[94]

Gen. et sp. nov

Valid

Xing et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of Avialae of uncertain phylogenetic placement. Genus includes new species O. khaungraae.

Pterosaurs

Research

  • Mazin & Pouech (2020) describe non-pterodactyloid pterosaur tracks from the ichnological site known as "the Pterosaur Beach of Crayssac" (Tithonian; south-western France), evaluate the implications of these tracks for the knowledge of the terrestrial capabilities of non-pterodactyloid pterosaurs, and name a new ichnogenus Rhamphichnus.[95]
  • A coleoid cephalopod specimen preserved with an associated tooth of a pterosaur (probably Rhamphorhynchus) is reported from the Upper Jurassic Altmühltal Formation (Germany) by Hoffmann et al. (2020), who evaluate the implications of this finding for the knowledge of feeding behaviours of Rhamphorhynchus.[96]
  • A well-preserved basihyal is reported for the first time in a pterosaur specimen (possibly belonging to the species Gladocephaloideus jingangshanensis) from the Lower Cretaceous Yixian Formation (China) by Jiang et al. (2020).[97]
  • Jacobs et al. (2020) describe new fossil material of pterosaurs from the Kem Kem Beds (Morocco), bringing the Kem Kem pterosaur fauna up to at least nine species (of which three are ornithocheirids), and confirming that toothed pterosaurs remained diverse during the mid-Cretaceous.[98]
  • Averianov (2020) reassesses the taxonomy of the Lonchodectidae, transferring the species "Lonchodraco" machaerorhynchus (including L. microdon and Pterodactylus oweni) to the genus Ikrandraco. [99]
  • New information on the anatomy of Dsungaripterus weii (especially on the palatal region), based on the study of new and previously collected specimens, is published by Chen et al. (2020).[100]

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Afrotapejara [101]

Gen. et sp. nov

In press

Martill et al.

Cretaceous

Kem Kem

 Morocco

A tapejarid pterosaur. Genus includes new species A. zouhri.

Albadraco [102]

Gen. et sp. nov

In press

Solomon et al.

Late Cretaceous (Maastrichtian)

Sard

 Romania

An azhdarchid pterosaur. Genus includes new species A. tharmisensis. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Apatorhamphus[103]

Gen. et sp. nov

In press

McPhee et al.

Middle Cretaceous (Albian/Cenomanian)

Kem Kem

 Morocco

A possible chaoyangopterid azhdarchoid pterosaur. Genus includes new species A. gyrostega.[103]

Luchibang[104]

Gen. et sp. nov

Valid

Hone et al.

Early Cretaceous

Yixian

 China

A member of the family Istiodactylidae. The type species is L. xinzhe.

Ordosipterus[105]

Gen. et sp. nov

Valid

Ji

Early Cretaceous

Luohandong Formation

 China

A member of the family Dsungaripteridae. The type species is O. planignathus.

Other archosaurs

  • A study on the anatomy, locomotion and phylogenetic relationships of Scleromochlus taylori is published by Bennett (2020).[106]
  • A study on the musculoskeletal apparatus and posture of Silesaurus opolensis, evaluating its implications for the knowledge of the evolution of the fully erect limb posture in archosaurs, is published by Piechowski & Tałanda (2020).[107]

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