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{{Taxobox
you moms tits are mine they are great bang her every night.
| name = Tits and Chickadees
| image = Parus major 4 (Marek Szczepanek).jpg
| image_width = 260px
| image_caption = [[Great Tit]]
| regnum = [[Animal]]ia
| phylum = [[Chordata]]
| classis = [[Aves]]
| ordo = [[passerine|Passeriformes]]
| subordo = [[Passeri]]
| familia = '''Paridae'''
| familia_authority = [[Nicholas Aylward Vigors|Vigors]], 1825
| subdivision_ranks = [[Genus|Genera]]
| subdivision =
5-10, see text.
|
| range_map = Paridae Distribution.PNG
| range_map_width = 200px
| range_map_caption = Global range (In green)
| synonyms =
see text
}}
{{Redirect|Titmouse|Titmouse, Inc.|Chris Prynoski}}
The '''tits''', '''chickadees''', and '''titmice''' comprise '''Paridae''', a large family of small [[passerine]] [[bird]]s which occur in the [[northern hemisphere]] and [[Africa]]. Most were formerly classified in the genus ''Parus''.

These birds are called "chickadees" ([[onomatopoeic]], derived from their distinctive "chick-a dee dee dee" communication or [[alarm call]]) or "titmice" in North America, and just "tits" in the rest of the [[English language|English]] speaking world. The name ''titmouse'' is attested from the 14th century, composed of the [[Old English language|Old English]] name for the bird, ''mase'' ([[Proto-Germanic]] ''*maison'', [[German language|German]] ''Meise'') and ''tit'', denoting something small. The spelling was influenced by ''[[mouse]]'' in the 16th century.

These birds are mainly small stocky woodland [[species]] with short stout [[Beak|bill]]s. Some have crests. They range in length from 10 to 22 centimetres. They are adaptable birds, with a mixed diet including seeds and [[insect]]s.<ref name=EoB>{{cite book |editor=Forshaw, Joseph|author= Perrins, C.|year=1991|title=Encyclopaedia of Animals: Birds|publisher= Merehurst Press|location=London|pages= 202-203|isbn= 1-85391-186-0}}</ref> Many species will live around human habitation and come readily to bird feeders for nuts or seed, and learn to take other foods. In Britain, [[Great Tit]]s and [[Blue Tit]]s famously learned to break open the foil caps sealing bottles of [[milk]] that had been delivered to homes to get at the [[cream]] floating on top.

These are hole-nesting birds, typically using trees, although some species build nests on the ground. They lay anything from three to nineteen speckled white [[Egg (biology)|egg]]s, depending on species.<ref name=EoB/>


==Behaviour==
The tits are active, noisy and social birds. They are [[territory (animal)|territorial]] during the breeding season and often joining [[mixed-species feeding flock]]s during the non-breeding season. The tits are highly adaptable and, after the [[Corvidae|corvids]] (crows and jays) and [[parrot]]s, amongst the most intelligent of all birds.<ref name ="HBW">Gosler, A. & P. Clement (2007) "Family Paridae (Tits and Chickadees)" P.p. 662-709. ''in'' del Hoyo, J.; Elliot, A. & Christie D. (editors). (2007). ''[[Handbook of the Birds of the World]]. Volume 12: Picathartes to Tits and Chickadees''. Lynx Edicions. ISBN 9788496553422</ref>
===Diet and feeding===
The tits are generalist insectivores that consume a wide range of small insects and other invertebrates, particularly small defoliating caterpillars. They also consume [[seed]]s and nuts, particularly in the winter. One characteristic method of foraging in the family is hanging, where they will inspect a branch or twig and leaves from all angles while hanging upside down to feed. In areas where numerous species of tit or titmouse coexist different species will forage in different parts of the tree, their niche determined in no small way by their morphology; larger species forage on the ground, medium sized species foraging on larger branches and the smallest species on the ends of branches. Having obtained larger prey items or seeds tits will engage in hold-hammering, where they will hold the item with one foot and hammer it with the bill until it is open. In this fashion they can even open [[hazelnut]]s in around 20 minutes. A number of genera engage in food caching, hoarding supplies of food during the winter.<ref>Jokinen S & J Suhonen (1995) "Food Caching By Willow and Crested Tits: A Test of Scatterhoarding Models" ''Ecology'' '''76''' (3): 892-898 </ref> These caches are usually of seeds but may be of insects.

== Systematics ==
[[Image:Parus palustris02.jpg|thumb|The [[Marsh Tit]] was once placed in the genus ''Parus'' but has now been moved to the genus ''Poecile''.]]
More recently, the large ''Parus'' group has been gradually split into several genera (as indicated below), which has been pioneered by [[North America]]n ornithological authorities and to a more limited degree (as of now) elsewhere. Whereas in the mid-1990s, only ''Pseudopodoces'', ''Baeolophus'', ''Melanochlora'' and ''Sylviparus'' were considered well-supported by the available data as distinct from ''Parus'' (Harrap & Quinn 1996). Today, this arrangement is considered [[paraphyletic]] as indicated by [[mtDNA]] [[cytochrome b|cytochrome ''b'']] [[DNA sequence|sequence]] analysis and ''Parus'' is best restricted to the ''Parus major'' - ''Parus fasciiventer'' [[clade]], and even the latter species' closest relatives might be considered a distinct genus (Gill ''et al.'' 2005).

In the [[Sibley-Ahlquist taxonomy]], the Paridae family is much enlarged to include related groups such as the [[Penduline tit]]s and [[Long-tailed tit]]s, but while the former are quite close to the titmice indeed and could conceivably included in that family together with the [[Stenostiridae|stenostirid "warblers"]], the long-tailed tits are not. Indeed, the [[Yellow-browed Tit]] and the [[Sultan Tit]] are possibly more distant to the titmice than are the penduline tits (Gill ''et al.'' 2005 and Jønsson & Fjeldsa 2006). If the two current families are lumped into the Paridae, the titmice would be a [[subfamily]] '''Parinae'''.

Alternatively, all tits - save the 2 monotypic genera discussed in the preceding section and possibly ''Cyanistes'', but including Hume's Ground Tit - could be lumped in ''Parus''. In any case, 4 major clades of "typical" tits can be recognized: the dark-capped chickadees and their relatives (''Poecile'' including ''Sittiparus''), the long-crested ''Baeolophus'' and ''Lophophanes'' species, the usually tufted, white-cheeked ''Periparus'' (including ''Pardaliparus'') with more subdued coloration and finally ''Parus sensu stricto'' (including ''Melaniparus'' and ''Macholophus''). Still, the interrelationship of these as well as the relationships of many species within the clades are not well resolved at all; analysis of [[Morphology (biology)|morphology]] and [[biogeography]] probably gives more a robust picture than the available molecular data (Gill ''et al.'', 2005).

Titmice have settled North America twice, probably at some time during the Early-Mid [[Pliocene]]. The first were the ancestors of ''Baeolophus''; chickadees arrived somewhat later (Gill ''et al.'', 2005).

==Species in taxonomic order==
* [[Penduline tit]]s and
* [[Stenostiridae|stenostirid "warblers"]]
:might be included here
'''Family: PARIDAE'''
* '''Genus ''[[Poecile]]''''' - formerly included in ''Parus''
** [[Sombre Tit]] ''Poecile lugubris''
** [[Caspian Tit]] ''Poecile hyrcanus'' (often included in ''P. lugubris'')
** [[Marsh Tit]] ''Poecile palustris''
** [[Willow Tit]] ''Poecile montanus''
** [[Songar Tit]] ''Poecile songarus'' (often included in ''P. montanus'')
** [[Carolina Chickadee]] ''Poecile carolinensis''
** [[Black-capped Chickadee]] ''Poecile atricapillus''
** [[Mountain Chickadee]] ''Poecile gambeli''
** [[Mexican Chickadee]] ''Poecile sclateri''
** [[White-browed Tit]] ''Poecile superciliosus''
** [[Père David's Tit]] ''Poecile davidi''
** [[Siberian Tit]] or Gray-headed Chickadee ''Poecile cinctus''
** [[Boreal Chickadee]] ''Poecile hudsonicus''
** [[Chestnut-backed Chickadee]] ''Poecile rufescens''

* '''Genus ''[[Periparus]]''''' - formerly included in ''Parus''<!-- ZoolAbhStaatlMusTierkdDresden55 p103 -->
** [[Black-breasted Tit]] or Rufous-naped Tit ''Periparus rufonuchalis''
** [[Rufous-vented Tit]] ''Periparus rubidiventris''<!-- Evolution8:19 -->
** [[Spot-winged Tit]] or Black-crested Tit ''Periparus melanolophus''<!-- Evolution8:19 -->
** [[Coal Tit]] ''Periparus ater'' - possibly paraphyletic
** [[Yellow-bellied Tit]] ''Periparus venustulus'' - sometimes separated in ''Pardaliparus''<!-- Evolution8:19 -->
** [[Elegant Tit]] ''Periparus elegans'' - probably paraphyletic; sometimes separated in ''Pardaliparus''<!-- Evolution8:19 -->
** [[Palawan Tit]] ''Periparus amabilis'' - sometimes separated in ''Pardaliparus''<!-- Evolution8:19 -->

* '''Genus ''[[Lophophanes]]''''' - formerly included in ''Parus''
** [[Crested Tit]] ''Lophophanes cristatus''
** [[Grey-crested Tit]] ''Lophophanes dichrous''<!-- Evolution8:19 -->

* '''Genus ''[[Baeolophus]]''''' - formerly included in ''Parus''
** [[Bridled Titmouse]] ''Baeolophus wollweberi''
** [[Oak Titmouse]] ''Baeolophus inornatus''
** [[Juniper Titmouse]] ''Baeolophus ridgwayi''
** [[Tufted Titmouse]] ''Baeolophus bicolor''
** [[Black-crested Titmouse]] ''Baeolophus atricristatus''

* '''Genus ''[[Parus]]''
** [[White-shouldered Tit]] ''Parus guineensis'' - sometimes separated in ''Melaniparus''
** [[White-winged Black Tit]] ''Parus leucomelas'' - sometimes separated in ''Melaniparus''
** [[Southern Black Tit]] ''Parus niger'' - sometimes separated in ''Melaniparus''
** [[Carp's Tit]] ''Parus carpi'' - sometimes separated in ''Melaniparus''
** [[White-bellied Tit]] ''Parus albiventris'' - sometimes separated in ''Melaniparus''
** [[White-backed Tit]] ''Parus leuconotus'' - sometimes separated in ''Melaniparus''
** [[Dusky Tit]] ''Parus funereus'' - sometimes separated in ''Melaniparus''
** [[Rufous-bellied Tit]] ''Parus rufiventris'' - sometimes separated in ''Melaniparus''
** [[Cinnamon-breasted Tit]] ''Parus pallidiventris'' - sometimes separated in ''Melaniparus''
** [[Red-throated Tit]] ''Parus fringillinus'' - sometimes separated in ''Melaniparus''
** [[Stripe-breasted Tit]] ''Parus fasciiventer'' - sometimes separated in ''Melaniparus''
** [[Somali Tit]] or Acacia Tit ''Parus thruppi'' - sometimes separated in ''Melaniparus''
** [[Miombo Tit]] ''Parus griseiventris'' - sometimes separated in ''Melaniparus''
** [[Ashy Tit]] ''Parus cinerascens'' - sometimes separated in ''Melaniparus''
** [[Southern Grey Tit]] ''Parus afer'' - sometimes separated in ''Melaniparus''
** [[Great Tit]] ''Parus major''
** [[Japanese Tit]] ''Parus minor'' (often included in ''P. major'')
** [[Turkestan Tit]] ''Parus bokharensis''
** [[Green-backed Tit]] ''Parus monticolus''<!-- Evolution8:19 -->
** [[White-winged Tit]] ''Parus nuchalis''
** [[Black-lored Tit]] ''Parus xanthogenys'' - sometimes separated in ''Macholophus''
** [[Yellow-cheeked Tit]] ''Parus spilonotus''
** [[Yellow Tit]] ''Parus holsti'' - sometimes separated in ''Macholophus''<!-- Evolution8:19 -->
** [[White-fronted Tit]] ''Parus semilarvatus'' - sometimes separated in ''Sittiparus''<!-- Evolution8:19 -->

* '''Genus ''Pseudopodoces'''''
** [[Hume's Ground Tit]] ''Pseudopodoces humilis'' (previously "Hume's Ground Jay") - this species has only recently been removed from the crow family [[Corvidae]] and placed here.<ref>James, H. F. et al. (2003). ''Pseudopodoces humilis'', a misclassified terrestrial tit (Paridae) of the Tibetan Plateau: evolutionary consequences of shifting adaptive zones. ''Ibis'' 145: 185–202.[http://www.nmnh.si.edu/vert/birds/pdf/hfj4.pdf pdf file]</ref>

* '''Genus ''[[Cyanistes]]''''' - formerly included in ''Parus''
** [[Blue Tit]] ''Cyanistes caeruleus''
** [[Canary Blue Tit]] ''Cyanistes teneriffae''
** [[Azure Tit]] ''Cyanistes cyanus''
** [[Yellow-breasted Tit]](Gold) ''Cyanistes flavipectus'' (often included in ''C. cyanus'')
** [[Varied Tit]] ''Cyanistes varius'' - sometimes separated in ''Sittiparus'' or ''Poecile''

* '''Genus ''Sylviparus'''''
** [[Yellow-browed Tit]] ''Sylviparus modestus''

* '''Genus ''Melanochlora'''''
** [[Sultan Tit]] ''Melanochlora sultanea''

:These two [[monotypic]] genera are possibly less close to titmice than are the penduline tits.

==References==<!-- Syst. Biol. 45(4):47 -->
<References/>
* '''Del Hoyo''', J.; Elliot, A. & Christie D. (editors). (2006). [[Handbook of the Birds of the World]]. Volume 12: Picathartes to Tits and Chickadees. Lynx Edicions. ISBN 9788496553422

* '''Gill''', Frank B.; Slikas, Beth & Sheldon, Frederick H. (2005): Phylogeny of titmice (Paridae): II. Species relationships based on sequences of the mitochondrial cytochrome-b gene. ''[[Auk (journal)|Auk]]'' '''122''': 121-143. <small>[[Digital Object Identifier|DOI]]: 10.1642/0004-8038(2005)122[0121:POTPIS]2.0.CO;2</small> [http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1642%2F0004-8038%282005%29122%5B0121%3APOTPIS%5D2.0.CO%3B2 HTML abstract]

* '''Harrap''', Simon & '''Quinn''', David (1996): ''Tits, Nuthatches & Treecreepers''. [[Helm Identification Guides|Christopher Helm]], London. <small>ISBN 0-7136-3964-4</small>

* '''Jønsson''', Knud A. & '''Fjeldså''', Jon (2006): Determining biogeographical patterns of dispersal and diversification in oscine passerine birds in Australia, Southeast Asia and Africa. ''[[Journal of Biogeography|J. Biogeogr.]]'' '''33'''(7): 1155–1165. <small>{{DOI|10.1111/j.1365-2699.2006.01507.x}}</small> (HTML abstract)

* '''Slikas''', Beth; Sheldon, Frederick H.; Gill, Frank B. (1996): Phylogeny of titmice (Paridae): I. Estimate of relationships among subgenera based on DNA-DNA hybridization. ''[[Journal of Avian Biology (journal)|Journal of Avian Biology]]'' '''27''': 70-82.

== External links ==
* Titmouse videos and photos: [http://ibc.lynxeds.com/family/chickadees-tits-paridae Internet Bird Collection]
* Photos of Eastern Blackcapped Chicadee: [http://www.pennsneckneighborhoodpark.biz/PNNP/Wildlife/index_files/wildlife_birds_one.htm]

[[Category:Bird families]]
[[Category:Paridae|*]]

[[ang:Māse]]
[[ca:Pàrid]]
[[cs:Sýkorovití]]
[[cy:Titw]]
[[de:Meisen]]
[[et:Tihaslased]]
[[es:Paridae]]
[[eo:Parida familio]]
[[fa:چرخ‌ریسک]]
[[fr:Paridae]]
[[ko:박새과]]
[[hr:Sjenice]]
[[io:Paruo]]
[[it:Paridae]]
[[ka:წივწივასებრნი]]
[[kv:Пыста]]
[[lb:Meesen]]
[[lt:Zyliniai]]
[[hu:Cinegefélék]]
[[nl:Mezen (vogels)]]
[[ja:シジュウカラ科]]
[[no:Meisefamilien]]
[[nn:Meisefamilien]]
[[pl:Sikorowate]]
[[pt:Paridae]]
[[ru:Синицевые]]
[[sah:Татыйык]]
[[sk:Sýkorkovité]]
[[fi:Tiaiset]]
[[sv:Mesfåglar]]
[[tr:Baştankara]]
[[uk:Синиця]]
[[zh:山雀科]]

Revision as of 14:48, 25 March 2009

Tits and Chickadees
Great Tit
Scientific classification
Kingdom:
Animalia
Phylum:
Chordata
Class:
Aves
Order:
Passeriformes
Suborder:
Passeri
Family:
Paridae

Vigors, 1825
Genera

5-10, see text.

Global range (In green)
Synonyms

see text

"Titmouse" redirects here. For Titmouse, Inc., see Chris Prynoski.

The tits, chickadees, and titmice comprise Paridae, a large family of small passerine birds which occur in the northern hemisphere and Africa. Most were formerly classified in the genus Parus.

These birds are called "chickadees" (onomatopoeic, derived from their distinctive "chick-a dee dee dee" communication or alarm call) or "titmice" in North America, and just "tits" in the rest of the English speaking world. The name titmouse is attested from the 14th century, composed of the Old English name for the bird, mase (Proto-Germanic *maison, German Meise) and tit, denoting something small. The spelling was influenced by mouse in the 16th century.

These birds are mainly small stocky woodland species with short stout bills. Some have crests. They range in length from 10 to 22 centimetres. They are adaptable birds, with a mixed diet including seeds and insects.[1] Many species will live around human habitation and come readily to bird feeders for nuts or seed, and learn to take other foods. In Britain, Great Tits and Blue Tits famously learned to break open the foil caps sealing bottles of milk that had been delivered to homes to get at the cream floating on top.

These are hole-nesting birds, typically using trees, although some species build nests on the ground. They lay anything from three to nineteen speckled white eggs, depending on species.[1]


Behaviour

The tits are active, noisy and social birds. They are territorial during the breeding season and often joining mixed-species feeding flocks during the non-breeding season. The tits are highly adaptable and, after the corvids (crows and jays) and parrots, amongst the most intelligent of all birds.[2]

Diet and feeding

The tits are generalist insectivores that consume a wide range of small insects and other invertebrates, particularly small defoliating caterpillars. They also consume seeds and nuts, particularly in the winter. One characteristic method of foraging in the family is hanging, where they will inspect a branch or twig and leaves from all angles while hanging upside down to feed. In areas where numerous species of tit or titmouse coexist different species will forage in different parts of the tree, their niche determined in no small way by their morphology; larger species forage on the ground, medium sized species foraging on larger branches and the smallest species on the ends of branches. Having obtained larger prey items or seeds tits will engage in hold-hammering, where they will hold the item with one foot and hammer it with the bill until it is open. In this fashion they can even open hazelnuts in around 20 minutes. A number of genera engage in food caching, hoarding supplies of food during the winter.[3] These caches are usually of seeds but may be of insects.

Systematics

The Marsh Tit was once placed in the genus Parus but has now been moved to the genus Poecile.

More recently, the large Parus group has been gradually split into several genera (as indicated below), which has been pioneered by North American ornithological authorities and to a more limited degree (as of now) elsewhere. Whereas in the mid-1990s, only Pseudopodoces, Baeolophus, Melanochlora and Sylviparus were considered well-supported by the available data as distinct from Parus (Harrap & Quinn 1996). Today, this arrangement is considered paraphyletic as indicated by mtDNA cytochrome b sequence analysis and Parus is best restricted to the Parus major - Parus fasciiventer clade, and even the latter species' closest relatives might be considered a distinct genus (Gill et al. 2005).

In the Sibley-Ahlquist taxonomy, the Paridae family is much enlarged to include related groups such as the Penduline tits and Long-tailed tits, but while the former are quite close to the titmice indeed and could conceivably included in that family together with the stenostirid "warblers", the long-tailed tits are not. Indeed, the Yellow-browed Tit and the Sultan Tit are possibly more distant to the titmice than are the penduline tits (Gill et al. 2005 and Jønsson & Fjeldsa 2006). If the two current families are lumped into the Paridae, the titmice would be a subfamily Parinae.

Alternatively, all tits - save the 2 monotypic genera discussed in the preceding section and possibly Cyanistes, but including Hume's Ground Tit - could be lumped in Parus. In any case, 4 major clades of "typical" tits can be recognized: the dark-capped chickadees and their relatives (Poecile including Sittiparus), the long-crested Baeolophus and Lophophanes species, the usually tufted, white-cheeked Periparus (including Pardaliparus) with more subdued coloration and finally Parus sensu stricto (including Melaniparus and Macholophus). Still, the interrelationship of these as well as the relationships of many species within the clades are not well resolved at all; analysis of morphology and biogeography probably gives more a robust picture than the available molecular data (Gill et al., 2005).

Titmice have settled North America twice, probably at some time during the Early-Mid Pliocene. The first were the ancestors of Baeolophus; chickadees arrived somewhat later (Gill et al., 2005).

Species in taxonomic order

might be included here

Family: PARIDAE

  • Genus Pseudopodoces
    • Hume's Ground Tit Pseudopodoces humilis (previously "Hume's Ground Jay") - this species has only recently been removed from the crow family Corvidae and placed here.[4]
  • Genus Melanochlora
These two monotypic genera are possibly less close to titmice than are the penduline tits.

References

  1. ^ a b Perrins, C. (1991). Forshaw, Joseph (ed.). Encyclopaedia of Animals: Birds. London: Merehurst Press. pp. 202–203. ISBN 1-85391-186-0.
  2. ^ Gosler, A. & P. Clement (2007) "Family Paridae (Tits and Chickadees)" P.p. 662-709. in del Hoyo, J.; Elliot, A. & Christie D. (editors). (2007). Handbook of the Birds of the World. Volume 12: Picathartes to Tits and Chickadees. Lynx Edicions. ISBN 9788496553422
  3. ^ Jokinen S & J Suhonen (1995) "Food Caching By Willow and Crested Tits: A Test of Scatterhoarding Models" Ecology 76 (3): 892-898
  4. ^ James, H. F. et al. (2003). Pseudopodoces humilis, a misclassified terrestrial tit (Paridae) of the Tibetan Plateau: evolutionary consequences of shifting adaptive zones. Ibis 145: 185–202.pdf file
  • Del Hoyo, J.; Elliot, A. & Christie D. (editors). (2006). Handbook of the Birds of the World. Volume 12: Picathartes to Tits and Chickadees. Lynx Edicions. ISBN 9788496553422
  • Gill, Frank B.; Slikas, Beth & Sheldon, Frederick H. (2005): Phylogeny of titmice (Paridae): II. Species relationships based on sequences of the mitochondrial cytochrome-b gene. Auk 122: 121-143. DOI: 10.1642/0004-8038(2005)122[0121:POTPIS]2.0.CO;2 HTML abstract
  • Harrap, Simon & Quinn, David (1996): Tits, Nuthatches & Treecreepers. Christopher Helm, London. ISBN 0-7136-3964-4
  • Jønsson, Knud A. & Fjeldså, Jon (2006): Determining biogeographical patterns of dispersal and diversification in oscine passerine birds in Australia, Southeast Asia and Africa. J. Biogeogr. 33(7): 1155–1165. doi:10.1111/j.1365-2699.2006.01507.x (HTML abstract)
  • Slikas, Beth; Sheldon, Frederick H.; Gill, Frank B. (1996): Phylogeny of titmice (Paridae): I. Estimate of relationships among subgenera based on DNA-DNA hybridization. Journal of Avian Biology 27: 70-82.
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