Urocyclidae
| Urocyclidae | |
|---|---|
| |
| A live individual of Trochonanina mozambicensis | |
Scientific classification 
| |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Mollusca |
| Class: | Gastropoda |
| Order: | Stylommatophora |
| Suborder: | Helicina |
| Infraorder: | Limacoidei |
| Superfamily: | Helicarionoidea |
| Family: | Urocyclidae Simroth, 1889[1] |
Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea, (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[2]
Some species within this family make and use love darts before mating.[citation needed]
Distribution
The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.[3][4]
Taxonomy
The family Urocyclidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005),[2] that follows classification of Urocyclinae by Van Goethem (1977):[5]
- subfamily Rhysotininae Schileyko, 2002
- subfamily Sheldoniinae Connolly, 1925 (1912)[6] - synonyms: Peltatinae Godwin-Austen, 1912;[7] Trochonanininae Connolly, 1912;[8] Trochozonitinae Iredale, 1914;[9] Ledoulxiinae Pilsbry, 1919;[10] Gymnarionidae Von Mol, 1970;[11] Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002
- subfamily Urocyclinae Simroth, 1889
- tribe Urocyclini Simroth, 1889 - synonyms: Atoxonini Schileyko, 2002; Buettneriini Schileyko, 2002
- tribe Dendrolimacini Van Goethem, 1977[5]
- tribe Leptichnini Van Goethem, 1977[5]
- tribe Upembellini Van Goethem, 1977[5]
Genera
Genera within the family Urocyclidae include:
- unsorted
- † Koruella Pickford, 2009
- Subfamily Rhysotininae Schileyko, 2002
- Rhysotina Ancey, 1887[2]
- Thomeonanina Germain, 1909 : synonym of Rhysotina Ancey, 1887 (objective synonym)
- Subfamily Sheldoniinae
- Acantharion Binder & Tillier, 1985[2]
- Africarion Godwin-Austen, 1883
- Amatarion Van Mol, 1970
- Angustivestis Pilsbry, 1919
- Aspidelus Morelet, 1884
- Aspidotomium Degner, 1932
- Belonarion Pilsbry, 1919
- Bloyetia Bourguignat, 1890
- Camerunarion Van Mol, 1970
- Carinazingis van Bruggen & de Winter, 1990
- Centrafricarion Van Mol, 1970 [11]
- Chlamydarion Van Mol, 1968
- Colparion Laidlaw, 1938
- Degneria Verdcourt, 1956
- Entagaricus Pilsbry, 1919
- Estria Poirier, 1887
- Falloonella Preston, 1914
- Fuellebornia Verdcourt, 1998
- Granularion Germain, 1912
- Gymnarion Pilsbry, 1919[2]
- Hamya Bourguignat, 1885 †
- Kerkophorus Godwin-Austen, 1912
- Lacrimarion Connolly, 1939
- Malagarion Tillier, 1979
- Mesafricarion Pilsbry, 1919
- Microkerkus Godwin-Austen, 1912
- Montanobloyetia Verdcourt, 1961
- Morrumbalia Verdcourt, 1998
- Paragranularion Van Mol, 1970
- Percivalia Preston, 1914
- Plicatonanina Verdcourt, 1961
- Principicochlea D. Holyoak & G. Holyoak, 2020
- Principitrochoidea D. Holyoak & G. Holyoak, 2020
- Ptilototheca Herbert, 2016
- Rhopalogonium Degner, 1932
- Selatodryas Herbert, 2017
- Senegalarion Van Mol, 1970
- Sheldonia Ancey, 1887 - the type genus of the subfamily Sheldoniinae[2][12]
- Sjostedtina Verdcourt, 1961
- Sylvarion Van Mol, 1970
- Thapsia Albers, 1860
- Thielarion Van Mol, 1970
- Thomithapsia D. Holyoak & G. Holyoak, 2020
- Thomitrochoidea D. Holyoak & G. Holyoak, 2020
- Tresia Van Goethem, 1975
- Trochonanina Mousson, 1869[2]
- Trochozonites Pfeffer, 1883[2]
- Tropidocochlion Verdcourt, 1998
- Verdcourtia Van Mol, 1970
- Verrucarion Van Mol, 1970 [13]
- Zingis E. von Martens, 1878
- Zonitarion Pfeffer, 1883[2]
- Ledoulxia Bourguignat, 1885:[2] synonym of Trochonanina Mousson, 1869
- Peltatus Godwin-Austen, 1908:[2] synonym of Sheldonia Ancey, 1887
- Subfamily Urocyclinae
Tribe Dendrolimacini
- Dendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini[2]
Tribe Leptichnini
- Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini[2]
Tribe Upembellini
- Leptichnoides Van Goethem, 1975
- Upembella Van Goethem, 1969 - the type genus of the tribe Upembellini[2]
tribe Urocyclini
- Anisotoxon Van Goethem, 1975
- Atoxon Simroth, 1888[2]
- Atoxonoides Van Goethem, 1973
- Atrichotoxon Simroth, 1910
- Buettneria Simroth, 1910[14]
- Bukobia Simroth, 1896
- Elisolimax Cockerell, 1893
- Emphysetes Verdcourt, 2003
- Microcyclus Simroth, 1896
- Nupnus Van Goethem, 1975
- Pembatoxon Van Goethem, 1975
- Phaneroporus Simroth, 1888
- Polytoxon Simroth, 1897
- Pseudatoxon Van Goethem, 1975
- Tanzalimax Rowson, Paustian & Van Goethem, 2017
- Trichotoxon Simroth, 1888
- Udzungwalimax Rowson, Paustian & Van Goethem, 2017
- Urocyclus Gray, 1864 - the type genus of the family Urocyclidae[2]
Cladogram
The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade:[3]
References
- ^ Simroth H. (1899). "Beiträge zur Kenntniss der Nacktschnecken". Nova Acta, Kaiserlich Leopoldinish-Carolinische Deutsche Akademie der Naturforscher (= Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum) 54(1): 1-91, plates 1-4. p. 62.
- ^ a b c d e f g h i j k l m n o p Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
- ^ a b Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.
- ^ Herbert, Dai; Kilburn, Dick (2004). Field guide to the land snails and slugs of eastern south Africa (1st ed.). Pietermaritzburg :Natal Museum: Natal Museum. ISBN 0-620-32415-5.
- ^ a b c d Van Goethem J. (1977). "Révision systématique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)". Musée Royal de l'Afrique Centrale , Annales, Sciences zoologiques 218: 355 pp., 4 plates.
- ^ Connolly (1925). Annals and Magazine of Natural History (9)15: 467.
- ^ Godwin-Austen H. H. (1912). "A review of South-African Land-Mollusca belonging to the Family Zonitidae". Annals and Magazine of Natural History (8)9: 122-139. Plate 1-7. p. 124.
- ^ Connolly (1912). Annals of the South African Museum 11(3): 101.
- ^ Iredale (1914). Proceedings of the Malacological Society of London 11(2): 122.
- ^ Pilsbry H. A. (1919). Bulletin of the American Museum of Natural History 40: 245.
- ^ a b Van Mol J. J. (1970). "Révision des Urocyclidae (Mollusca, Gastropoda, Pulmonata). Anatomie - systématique - zoogéographie". Annales Musée Royal de l’Afrique Centrale, Sciences Zoologiques 180: 1-231.
- ^ Herbert D. G. (2004). Sheldonia puzeyi. 2006 IUCN Red List of Threatened Species. Downloaded on 7 August 2007.
- ^ de Winter A. J. (1987). "Verrucarion demeryi, a new semi-slug from Sierra Leone (Mollusca: Pulmonata: Urocyclidae)". Basteria 51(4-6): 87-93. abstract.
- ^ Simroth H. (1910). In: Voeltzkow Reise O.-Afr., Wiss. Ergebn. 2(5): 611.
External links
Media related to Urocyclidae at Wikimedia Commons
