Doswelliidae

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Doswelliidae
Temporal range: Middle - Late Triassic
Doswellia BW.jpg
Life restoration Doswellia kaltenbachi, the most well known dosweliid
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Proterochampsia
Family: Doswelliidae
Weems, 1980
Genera
Life restoration of Archeopelta arborensis

Doswelliidae is a family of carnivore archosauriforms. Doswelliids existed in North and South America during the Middle–Late Triassic period and were among the most derived non-archosaurian archosauriforms. The family was named by R. E. Weems in 1980 and it has been placed in its own suborder, Doswelliina.[1]

The Doswelliidae has long been considered a monospecific family of basal archosauriforms represented by Doswellia kaltenbachi from the Late Triassic of North America. However, a 2011 cladistic analysis recovered Archeopelta, Doswellia and Tarjadia within a monophyletic Doswelliidae. The phylogenetic analysis also indicates that Doswellidae is the closest large monophyletic clade to Archosauria (only the Chinese archosauriform Yonghesuchus nested closer to archosaurs).[2] In 2013, the fourth and fifth genera and species of dosweliids were named, Jaxtasuchus salomoni based on several skeletons found in the Ladinian-age Lower Keuper of Germany,[3] and Ankylosuchus chinlegroupensis based on fragments of four vertebrae, parts of the skull and of a limb bone from the early Carnian Colorado City Formation.[4] Both Ankylosuchus and Jaxtasuchus are more closely related to Doswellia than to more basal doswelliids.[3][4] A phylogenetic analysis by Ezcurra (2016) recovered Doswelliidae within Proterochampsia, which was found to be the sister taxon of Archosauria. The aquatic Vancleavea was also referred to Doswelliidae.[5]

 Eucrocopoda 

Dorosuchus neoetus




Euparkeria capensis (Euparkeriidae)




Dongusuchus efremovi



Yarasuchus deccanensis




 Archosauria 

Avemetatarsalia


 Pseudosuchia 

Phytosauria



The rest of Pseudosuchia (incl. Koilamasuchus)




 Proterochampsia 
 Doswelliidae 

Vancleavea campi





Archeopelta arborensis



Tarjadia ruthae





Doswellia kaltenbachi



Jaxtasuchus salomoni





 Proterochampsidae 


Proterochampsa barrionuevoi



Proterochampsa nodosa





Cerritosaurus binsfeldi




Tropidosuchus romeri


 Rhadinosuchinae 

Pseudochampsa ischigualastensis




Gualosuchus reigi




Chanaresuchus bonapartei



Rhadinosuchus gracilis












References[edit]

  1. ^ R. E. Weems (1980). "An unusual newly discovered archosaur from the Upper Triassic of Virginia, U.S.A.". Transactions of the American Philosophical Society, New Series. 70 (7): 1–53. doi:10.2307/1006472. 
  2. ^ Julia B. Desojo; Martin D. Ezcurra; Cesar L. Schultz (2011). "An unusual new archosauriform from the Middle–Late Triassic of southern Brazil and the monophyly of Doswelliidae". Zoological Journal of the Linnean Society. 161 (4): 839–871. doi:10.1111/j.1096-3642.2010.00655.x. 
  3. ^ a b Schoch, R. R.; Sues, H. D. (2013). "A new archosauriform reptile from the Middle Triassic (Ladinian) of Germany". Journal of Systematic Palaeontology: 1. doi:10.1080/14772019.2013.781066. 
  4. ^ a b Lucas, S.G.; Spielmann, J.A.; Hunt, A.P. (2013). "A new doswelliid archosauromorph from the Upper Triassic of West Texas" (PDF). In Tanner, L.H.; Spielmann, J.A.; and Lucas, S.G. (eds.). The Triassic System. New Mexico Museum of Natural History and Science Bulletin. 61. pp. 382–388. 
  5. ^ Ezcurra MD. (2016) The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms. PeerJ, 4:e1778 [1]