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Laemophloeus germaini Montage.jpg
Laemophloeus germaini
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Polyphaga
Superfamily: Cucujoidea
Family: Laemophloeidae
Ganglbauer, 1899

Acompsophloeus Thomas, 2010
Blubos Lefkovitch, 1962
Brontolaemus Sharp, 1885
Carinophloeus Lefkovitch, 1961
Caulonomus Wollaston, 1862
Charaphloeus Casey, 1916
Cryptolestes Ganglbauer, 1899
Cucujinus Arrow, 1920
Deinophloeus Sharp, 1899
Dysmerus Casey, 1884
Gannes Lefkovitch, 1962
Heterojinus Sengupta & Mukhopadhyay, 1978
Laemophloeus Laporte de Castelnau, 1840
Lathropus Erichson, 1845
Lepidophloeus Thomas, 1984
Leptophloeus Casey, 1916
Magnoleptus Lefkovitch, 1962
Mariolaemus Lefkovitch, 1962
Mestolaemus Lefkovitch, 1962
Metaxyphloeus Thomas, 1984
Microbrontes Reitter, 1874
Microlaemus Lefkovich, 1962
Narthecius J.LeConte, 1861
Nipponophloeus Sasaji, 1983
Notolaemus Lefkovitch, 1959
Odontophloeus Thomas, 1984
Parandrita J.LeConte & Horn, 1880
Paraphloeolaemus Thomas, 2018
Passandrophloeus Kessel, 1921
Phloeipsius Casey, 1916
Phloeolaemus Casey, 1916
Placonotus MacLeay, 1871
Planolestes Lefkovitch, 1958
Pseudophloeus Yablokov-Khnzoryan, 1977
Rhabdophloeus Sharp, 1899
Rhinolaemus Steel, 1954
Rhinomalus Gemm, 1870
Rhinophloeus Sharp, 1899
Sinuatophloeus Kessel, 1921
Xylolestes Lefkovitch, 1962
Xylophloeus Lefkovitch, 1962

Laemophloeidae, "lined flat bark beetles," is a family in the superfamily Cucujoidea characterized by predominantly dorso-ventrally compressed bodies, head and pronotal discs bordered by ridges or grooves, and inverted male genitalia.[1][2] Size range of adults is 1–5 mm (0.04–0.2 in) in length. Currently, it contains 40 genera and about 450 species,[3] and is represented on all continents except Antarctica; species richness is greatest in the tropics.

Historically, Laemophloeidae was treated as a subfamily of Cucujidae, but starting in the middle of the 20th century,[4] most of what had been treated as subfamilies of the Cucujidae were considered to be families.

Most laemophloeids, adults and larvae, are found under bark of dead trees, where they apparently are primarily fungivores,[5] although some genera with adults having subcylindrical bodies (e.g., Leptophloeus, Dysmerus) occur in the galleries of bark beetles (Curculionidae: Scolytinae), upon which they may feed.[6] A few genera, but most particularly Cryptolestes, contain some species that are pests of stored grain products. The most important of these are C. ferrugineus (Stephens), C. pusillus (Schönherr), and C. turcicus (Grouvelle).[7]

Several genera exhibit unusual modifications to male antennae (especially Cryptolestes, Dysmerus, and Microbrontes), with the scape expanded into hook-like or blade-like structures[8][9] Several other genera (Rhinomalus, Rhinophloeus, and Metaxyphloeus) related to Laemophloeus are atypical in that the adults are rostrate to varying degrees[10] Photographs of most world genera are available at,[11] and most North American species are pictured at,[12]


  1. ^ Thomas, M.C. 2002. Family 83. Laemophloeidae Ganglbauer 1899. Pp. 331-334 In: Arnett, R. H., Jr., M. C. Thomas, P. E. Skelley, and J. H. Frank (editors). 2002. American Beetles. Vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press, Boca Raton. xiv + 861pp.
  2. ^ Thomas, M. C., and R.A.B. Leschen. 2010. Laemophloeidae, Ganglbauer, 1899. p. 376-380. In: Leschen, R.A.B., R.G. Beutel, and J.F. Lawrence. Coleoptera, Beetles. Vol. 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). Handbook of Zoology. Walter de Gruyter, Berlin.
  3. ^ Laemophloeidae Species List Archived 2012-10-23 at the Wayback Machine. at Joel Hallan’s Biology Catalog. Texas A&M University. Retrieved on 15 May 2012.
  4. ^ Crowson, R.A. 1955. The natural classification of the families of Coleoptera. Nathaniel Lloyd, London, 187pp.
  5. ^ Lawrence, J.F. 1977. Coleoptera associated with an Hypoxylon species (Ascomycetes: Xylariaceae) on oak. Coleopterists Bulletin 31:309-312.
  6. ^ Thomas, M.C. 1993. The flat bark beetles of Florida (Laemophloeidae, Passandridae, Silvanidae). Arthropods of Florida and Neighboring Land Areas 15: i-viii and 1-93.
  7. ^ Halstead, D.G.H. 1993. Keys for the identification of beetles associated with stored products-II. Laemophloeidae, Passandridae and Silvanidae. Journal of Stored Product Research 29(2): 99-197.
  8. ^ Lefkovitch, L.P. 1958. Unusual antennal characters in some Laemophloeinae (Coleoptera: Cucujidae) and their taxonomic importance. Proceedings of the Royal Entomological Society of London B 27:93-100.
  9. ^ Thomas, M. C. 2009. A review of the genus Dysmerus Casey (Coleoptera: Laemophloeidae). Insecta Mundi 0074: 1-30.
  10. ^ Thomas, M.C. 1984. A new Neotropical genus and species of rostrate Laemophloeinae (Coleoptera: Cucujidae), with discussion of the systematic position of the subfamily. Coleopterists Bulletin 38:67-83.
  11. ^ Photographic Atlas of Laemophloeid Genera Retrieved 31 January 2014
  12. ^ Laemophloeidae of Florida Retrieved 31 January 2014