Chlamyphoridae

Chlamyphoridae; Temporal range: Late Oligocene to present
	Taxiderm pink fairy armadillo (Chlamyphorus truncatus)
	Illustration of a skeleton of Doedicurus clavicaudatus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Cingulata
Family:	Chlamyphoridae; Bonaparte, 1850
Subfamilies
	Chlamyphorinae; Euphractinae; †Glyptodontinae; Tolypeutinae;

Chlamyphoridae is a family of cingulate mammals. While glyptodonts have traditionally been considered stem-group cingulates outside the group that contains modern armadillos, there had been speculation that extant family Dasypodidae could be paraphyletic based on morphological evidence.^[1]^[2]^[3]^[4] In 2016, an analysis of Doedicurus mtDNA found it was, in fact, nested within the modern armadillos as the sister group of a clade consisting of Chlamyphorinae and Tolypeutinae.^[5]^[6] For this reason all armadillos but Dasypus were relocated to a new family.

Classification

Below is a taxonomy of the extant species of armadillos in this family.

Family Chlamyphoridae

Subfamily Chlamyphorinae
- Genus Calyptophractus
  - Greater fairy armadillo, Calyptophractus retusus
- Genus Chlamyphorus, Chlamyphorus truncatus
  - Pink fairy armadillo, Chlamyphorus truncatus
Subfamily Euphractinae
- Genus Euphractus
  - Six-banded armadillo, Euphractus sexcinctus
- Genus Zaedyus
  - Pichi, Zaedyus picky
- Genus Chaetophractus
  - Screaming hairy armadillo, Chaetophractus vellerosus
  - Big hairy armadillo, Chaetophractus villosus
  - Andean hairy armadillo, Chaetophractus nationi
Subfamily Tolypeutinae
- Genus Cabassous
  - Greater naked-tailed armadillo, Cabassous tatouay
  - Chacoan naked-tailed armadillo, Cabassous chacoensis
  - Northern naked-tailed armadillo, Cabassous centralis
  - Southern naked-tailed armadillo, Cabassous unicinctus
- Genus Priodontes
  - Giant armadillo, Priodontes maximus
- Genus Tolypeutes
  - Southern three-banded armadillo, Tolypeutes matacus
  - Brazilian three-banded armadillo, Tolypeutes tricinctus

Cladogram after the analysis of Delsuc et al., 2016:^[5]

Chlamyphoridae

Euphractinae

Euphractus sexcinctus

Chaetophractus villosus

	Zaedyus pichiy

	Chaetophractus vellerosus

Glyptodontinae

Chlamyphorinae

	Chlamyphorus truncatus

	Calyptophractus retusus

Tolypeutinae

Priodontes maximus

	Tolypeutes tricinctus

	Tolypeutes matacus

Cabassous tatouay

Cabassous chacoensis

	Cabassous centralis

	Cabassous unicinctus

References

^ Simpson, G. G. (1945). The principles of classification and a classification of mammals. Bull. Amer. Museum Nat. History., 85.
^ Grassé, P. P. (1955). Ordre des édentés. Traité de zoologie, 17(2), 1182-1246.
^ Engelmann, G. F. (1985). The phylogeny of the Xenarthra. The evolution and ecology of armadillos, sloths, and vermilinguas. Smithsonian Institution Press, Washington, DC, 51-64.
^ Wible, J. R. (2006). 6 The Phylogeny of Living and Extinct Armadillos (Mammalia, Xenarthra, Cingulata): A Craniodental Analysis. Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles: University of Chicago Press, Chicago, IL, 153-198.
^ ^a ^b Delsuc, F.; Gibb, G.C.; Kuch, M.; Billet, G.; Hautier, L.; Southon, J.; Rouillard, J.-M.; Fernicola, J. C.; Vizcaíno, S. F.; MacPhee, R. D.E.; Poinar, H. N. (2016-02-22). "The phylogenetic affinities of the extinct glyptodonts". Current Biology. 26 (4): R155–R156. doi:10.1016/j.cub.2016.01.039.
^ Gillian C. Gibb, Fabien L. Condamine, Melanie Kuch, Jacob Enk, Nadia Moraes-Barros, Mariella Superina, Hendrik N. Poinar, and Frédéric Delsuc (2016). "Shotgun Mitogenomics Provides a Reference Phylogenetic Framework and Timescale for Living Xenarthrans". Molecular Biology and Evolution. doi:10.1093/molbev/msv250.{{cite journal}}: CS1 maint: multiple names: authors list (link)

[1] Simpson, G. G. (1945). The principles of classification and a classification of mammals. Bull. Amer. Museum Nat. History., 85.

[2] Grassé, P. P. (1955). Ordre des édentés. Traité de zoologie, 17(2), 1182-1246.

[3] Engelmann, G. F. (1985). The phylogeny of the Xenarthra. The evolution and ecology of armadillos, sloths, and vermilinguas. Smithsonian Institution Press, Washington, DC, 51-64.

[4] Wible, J. R. (2006). 6 The Phylogeny of Living and Extinct Armadillos (Mammalia, Xenarthra, Cingulata): A Craniodental Analysis. Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles: University of Chicago Press, Chicago, IL, 153-198.

[Delsuc2016-5] Delsuc, F.; Gibb, G.C.; Kuch, M.; Billet, G.; Hautier, L.; Southon, J.; Rouillard, J.-M.; Fernicola, J. C.; Vizcaíno, S. F.; MacPhee, R. D.E.; Poinar, H. N. (2016-02-22). "The phylogenetic affinities of the extinct glyptodonts". Current Biology. 26 (4): R155–R156. doi:10.1016/j.cub.2016.01.039.

[6] Gillian C. Gibb, Fabien L. Condamine, Melanie Kuch, Jacob Enk, Nadia Moraes-Barros, Mariella Superina, Hendrik N. Poinar, and Frédéric Delsuc (2016). "Shotgun Mitogenomics Provides a Reference Phylogenetic Framework and Timescale for Living Xenarthrans". Molecular Biology and Evolution. doi:10.1093/molbev/msv250.{{cite journal}}: CS1 maint: multiple names: authors list (link)

[1]

[2]

[3]

[4]

[5]

[6]