Spruce

This article is about the tree. For other uses, see Spruce (disambiguation).

Spruce
Norway Spruce (Picea abies)
Scientific classification
Kingdom:	Plantae
Division:	Pinophyta
Class:	Pinopsida
Order:	Pinales
Family:	Pinaceae
Subfamily:	Piceoideae Frankis
Genus:	Picea Mill.
Species
About 35; see text.

A spruce is a tree of the genus Picea /paɪˈsiːə/, a genus of about 35 species of coniferous evergreen trees in the Family Pinaceae, found in the northern temperate and boreal (taiga) regions of the earth. Spruces are large trees, from 20–60 metres (66–197 ft) tall when mature, and can be distinguished by their whorled branches and conical form. The needles, or leaves, of spruce trees are attached singly to the branches in a spiral fashion, each needle on a small peg-like structure called a pulvinus. The needles are shed when 4–10 years old, leaving the branches rough with the retained pulvinus (an easy means of distinguishing them from other similar genera, where the branches are fairly smooth).

Spruces are used as food plants by the larvae of some Lepidoptera species; see list of Lepidoptera that feed on spruces. They are also used by the larvae of gall adelgids (Adelges species).

In the mountains of western Sweden scientists have found a Norway Spruce tree, nicknamed Old Tjikko, which by reproducing through layering has reached an age of 9,550 years and is claimed to be the world's oldest known living tree.

Classification

DNA analyses have shown that traditional classifications based on the morphology of needle and cone are artificial. A recent study found that P. breweriana had a basal position, followed by P. sitchensis, and the other species were further divided into three clades, suggesting that Picea originated in North America.

Species

There are thirty-five named species of spruce in the world.

P. glauca sapling, Kluane National Park, Canada

Immature P. mariana cones, Ouimet Canyon, Ontario, Canada

P. pungens cone and foliage

Basal species:

• Picea breweriana Brewer's Spruce. Klamath Mountains, North America; local endemic.

Basal species

• Picea sitchensis Sitka Spruce. Pacific Coast of North America; the largest species, to 95m tall; important in forestry.

• Clade I

• Picea engelmannii Engelmann Spruce. Western North American mountains; important in forestry.
• Picea glauca White Spruce. Northern North America; important in forestry.

• Clade II

• Picea brachytyla Sargent's Spruce. Southwest China.
• Picea chihuahuana Chihuahua Spruce. Northwest Mexico (rare).
• Picea farreri Burmese Spruce. Northeast Burma, southwest China (mountains).
• Picea likiangensis Likiang Spruce. Southwest China.
• Picea martinezii Martinez Spruce. Northeast Mexico (very rare, endangered).
• Picea maximowiczii Maximowicz Spruce. Japan (rare, mountains).
• Picea morrisonicola Taiwan Spruce. Taiwan (high mountains).
• Picea neoveitchii Veitch's Spruce. Northwest China (rare, endangered).
• Picea orientalis Caucasian Spruce or Oriental Spruce . Caucasus, northeast Turkey.
• Picea purpurea Purple Spruce. Western China.
• Picea schrenkiana Schrenk's Spruce. Mountains of central Asia.
• Picea smithiana Morinda Spruce. Western Himalaya. Eastern Afghanistan, Northern and Northwest India
• Picea spinulosa Sikkim Spruce. Northeast India (Sikkim), Eastern Himalaya.
• Picea torano Tiger-tail Spruce. Japan.
• Picea wilsonii Wilson's Spruce. Western China.

• Clade III

• Picea abies Norway Spruce. Europe; important in forestry. The original Christmas tree.
• Picea alcoquiana ("P. bicolor") Alcock's Spruce. Central Japan (mountains).
• Picea alpestris Norway Spruce, Alpine Spruce. The Alps in Europe; rare, often treated as a variant of P. abies (and hybridises with it) distinct cones.
• Picea asperata Dragon Spruce. Western China; several varieties.
• Picea crassifolia. China.
• Picea glehnii Glehn's Spruce. Northern Japan, Sakhalin.
• Picea jezoensis Jezo Spruce. Northeast Asia, Kamchatka south to Japan.
• Picea koraiensis Korean Spruce. Korea, northeast China.
• Picea koyamae Koyama's Spruce. Japan (mountains).
• Picea mariana Black Spruce. Northern North America.
• Picea meyeri Meyer's Spruce. Northern China (from Inner Mongolia to Gansu).
• Picea obovata Siberian Spruce. North Scandinavia, Siberia. Often treated as a variant of P. abies (and hybridises with it) but distinct cones.
• Picea omorika Serbian Spruce. Serbia and Bosnia; local endemic; important in horticulture.
• Picea pungens Blue Spruce or Colorado Spruce. Rocky Mountains, North America; important in horticulture.
• Picea retroflexa. China.
• Picea rubens Red Spruce. Northeastern North America; important in forestry. Known as Adirondack in musical instrument making.

Uses

Timber

P. abies wood

Spruce is useful as a building wood, commonly referred to by several different names including North American timber, SPF (spruce, pine, fir) and whitewood. Spruce wood is used for many purposes, ranging from general construction work and crates to highly specialised uses in wooden aircraft, and as a tonewood in many musical instruments, including guitars, mandolins, cellos, violins, and the soundboard at the heart of a piano and the harp. The Wright brothers' first aircraft, the Flyer, was built of spruce.

Because this species has no insect or decay resistance qualities after logging, it is generally recommended for construction purposes as indoor use only (ex. indoor drywall framing). Spruce wood, when left outside cannot be expected to last more than 12–18 months depending on the type of climate it is exposed to.

Pulpwood

Spruce is one of the most important woods for paper uses, as it has long wood fibres which bind together to make strong paper. The fibres are thin walled and collapse to thin bands upon drying. Spruces are commonly used in mechanical pulping as they are easily bleached. Together with northern pines, northern spruces are commonly used to make NBSK. Spruces are cultivated over vast areas as pulpwood.

Food and medicine

Spruce (Picea mariana) essential oil in a clear glass vial

The fresh shoots of many spruces and pines are a natural source of vitamin C. Captain Cook made alcoholic sugar-based spruce beer during his sea voyages in order to prevent scurvy in his crew. The leaves and branches, or the essential oils, can be used to brew spruce beer.

The tips from the needles can be used to make spruce tip syrup . In survival situations spruce needles can be directly ingested or boiled into a tea. This replaces large amounts of vitamin C. Also, water is stored in a spruce's needles, providing an alternative means of hydration . Spruce can be used as a preventive measure for scurvy in an environment where meat is the only prominent food source .

Other uses

The resin was used in the manufacture of pitch in the past (before the use of petrochemicals); the scientific name Picea is generally thought to be derived from Latin pix, pitch (though other etymologies have been suggested).

Native Americans in North America use the thin, pliable roots of some species for weaving baskets and for sewing together pieces of birch bark for canoes. See also Kiidk'yaas for an unusual golden Sitka Spruce sacred to the Haida people.

Spruces are also popular ornamental trees in horticulture, admired for their evergreen, symmetrical narrow-conic growth habit. For the same reason, some (particularly Picea abies and P. omorika) are also extensively used as Christmas trees.

Spruce branches are also used at Aintree racecourse, Liverpool, to build several of the fences on the Grand National course. It is also used to make sculptures and Christmas trees.

Etymology

Picea used in coat-of-arms of Kuhmo, Finland

The word "spruce" entered the English language from Old French Pruce, the name of Prussia. Spruce was a generic term for commodities brought to England by Hanseatic merchants and the tree was believed to have come from Prussia.

External links

• Arboretum de Villardebelle: Cones of selected species of Picea: page 1, Arboretum de Villardebelle page 2
• Gymnosperm Database - Picea