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Klondike Mountain Formation

Coordinates: 48°42′N 118°42′W / 48.7°N 118.7°W / 48.7; -118.7
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Klondike Mountain Formation
Stratigraphic range: Ypresian
50.1–48.9 Ma
Klondike Mountain Formation outcrop at site A0307
TypeGeological formation
Unit ofOkanagan Highlands
Sub-unitsTom Thumb Tuff, Lower & Upper Members
UnderliesGlacial deposits
OverliesSanpoil Volcanics & others
Thicknessup to 3,200 ft (980 m)
Lithology
PrimaryVariable
OtherSee text
Location
Coordinates48°42′N 118°42′W / 48.7°N 118.7°W / 48.7; -118.7
Approximate paleocoordinates54°24′N 101°54′W / 54.4°N 101.9°W / 54.4; -101.9
RegionOkanagan highlands
Country United States
ExtentOkanagan highlands
Type section
Named forKlondike Mountain
Named byMuessig
Year defined1962
Klondike Mountain Formation is located in the United States
Klondike Mountain Formation
Klondike Mountain Formation (the United States)
Klondike Mountain Formation is located in Washington (state)
Klondike Mountain Formation
Klondike Mountain Formation (Washington (state))

The Klondike Mountain Formation is an Early Eocene (Ypresian) geological formation located in the northeast central area of Washington state. The formation, named for the type location designated in 1962, Klondike Mountain north of Republic, Washington, is composed of volcanic rocks in the upper unit and volcanics plus lacustrine (lakebed) sedimentation in which a lagerstätte with exceptionally well-preserved plant and insect fossils has been found, along with fossil epithermal hot springs.

The formation is the youngest in a group of formations which belong to the Challis Sequence rocks. The formation unconformably overlies rocks of the Eocene Sanpoil Volcanics and much older Triassic and Permian formations. The formation is bounded on its edges by a series of high-angle strike slip faults, which have contained the Klondike Mountain Formation in a series of graben structures, such as the Republic Graben. Public access to a fossilifrous outcrop at the north end of Republic is mediated by the nonprofit Stonerose Interpretive Center.

Extent

The formation is located in northern Ferry County, Washington, with the majority of the sedimentation in the Republic and Curlew Basins on the east and in the Toroda Creek area to the north west. The town of Republic, Washington is situated at the southern end of the formation, with outcrops within the city itself. The formation was named for Klondike Mountain which is just to the northeast of the city, and where the type locality is located.[1] The Curlew basin is situated north of Republic, with the northern edge along the Kettle River and the community of Curlew, Washington near the northeastern edge.[2][3]

The formation is the southernmost of a string of preserved Eocene highland lakebeds in Washington state and British Columbia.[4] The lake system, within the Okanagan highlands, extends from the Klondike Mountain Formation north approximately 1,000 kilometres (1,000,000 m) in to southern central British Columbia.[5]

Age

Early dating of the formation was based primarily on identification and correlation of the fossils found in the Tom Thumb Tuff, with Joseph Umpleby (1910) reporting a putative age of Early Miocene.[6] This date was based in examination of fossils by C. R. Eastman, who thought them to be similar to those found in the Florissant Formation of Colorado, which at the time was also considered Miocene. This age was retained by Edward W. Berry (1929), who included the Klondike Mountain Formation fossil lakebeds as part of the Latah Formation.[7] The age of the Formation has been revised in the following hundred years, with Roland W. Brown (1936) identifying the deposits as being older than the Latah Formation.[8] In a later written communication circa 1958, Brown again revised the age still older, stating the fossils found in the area of Mount Elizabeth indicated an Oligocene age. This age was used by R.L. Parker and J. A. Calkins in their 1964 work on the Curlew Quadrangle of Ferry County.[9] Since then the fossil-bearing strata of the Formation have been radiometrically dated, to give a current estimate of the Ypresian, the mid stage of the early Eocene,49.4 ± .5 million years ago.[5]

Lithology

Parker and Calkins in 1964 noted the association of the Klondike Mountain Formation with the gold and silver deposits of the Republic District and suggested it as a potential host to more ore deposits in the Curlew Quadrangle.[9] The epithermal gold deposits occurring in the Sanpoil volcanics terminate directly below the unconformity where the volcanics contact the base of the Klondike Mountain Formation[10] or sometimes penetrate into the Formation's lowest unit.[11] Hydrothermal sinter deposits are known from the lowest portions of the Formation and are thought to represent hydrothermal eruption areas.[11] In general the lower portions of the Formation have a large amount of hydrothermal alteration, and areas around vents are rich in pyrite and silica. two products of natural hydrothermic sintering. The areas above that show a transition to mudstones, siltstones and sandstones grading from fine-grained material into coarser materials moving up the strata column. The finely-bedded stones show the greatest numbers of fossils and the finest preservation of details.[12]

Paleoenvironment and paleoclimate

The lake bed sediments preserve a diverse array of plants, insects, and fishes, notably the biota called the Republic flora. The Okanagan lake system, which includes the Klondike Mountain Formation, has been classified as one of the great Canadian lagerstätten.[13] The area likely had a mesic upper microthermal to lower mesothermal climate, in which winter temperatures rarely dropped low enough for snow, and which were seasonably equitable.[14]

The Okanagan highlands paleoforest surrounding the lakes have been described as precursors the to modern Temperate broadleaf and mixed forests of Eastern North America and Eastern Asia. Based on the fossil biotas the lakes were higher and cooler then the coeval coastal forests preserved in the Puget Group and Chuckanut Formation of Western Washington, which are described as lowland tropical forest ecosystems. Estimates of the paleoelevation between 0.7–1.2 km (0.43–0.75 mi) higher than the coastal forests. This is consistent with the paleoelevation estimates for the lake systems, which range between 1.1–2.9 km (1,100–2,900 m), which is similar to the modern elevation 0.8 km (0.50 mi), but higher.[14]

Estimates of the mean annual temperature have been derived from Climate leaf analysis multivariate program (CLAMP) and leaf margin analysis (LMA) of the Republic paleoflora. The CLAMP results after multiple linear regressions for Republic gave a mean annual temperature of approximately 8.0 °C (46.4 °F), while the LMA gave 9.2 ± 2.0 °C (48.6 ± 3.6 °F). This is lower then the mean annual temperature estimates given for the coastal Puget Group, which is estimated to have been between 15–18.6 °C (59.0–65.5 °F). The bioclimactic analysis for Republic suggests a mean annual precipitation amount of 115 ± 39 cm (45 ± 15 in).[14]

Paleobiota

Pteridophytes

Name Authority Year Family Notes Images

Azolla primaeva[15]

Arnold, 1955

1955

Salviniaceae

A mosquito fern

Azolla primaeva

Gingkophytes

Name Authority Year Family Notes Images

Ginkgo biloba[16]

Linnaeus

2002

Ginkgoaceae

A ginkgo

Ginkgo biloba

Ginkgo dissecta[16]

Mustoe, 2002

2002

Ginkgoaceae

A ginkgo

Ginkgo dissecta

Pinophytes

Name Authority Year Family Notes Images

Abies milleri[17]

Shorn & Wehr, 1986

1986

Pinaceae

Oldest true fir described

Abies milleri

Amentotaxus sp.[18]

Schorn & Wehr

1996

Taxaceae

A yew
Not described to species

Amentotaxus sp.

Cephalotaxus sp.[18]

Schorn & Wehr

1996

Taxaceae

A yew
Not described to species

Chamaecyparis sp.[18]

Schorn & Wehr

1996

Cupressaceae

A false cypress
Not described to species

Cryptomeria sp.[18]

Schorn & Wehr

1996

Cupressaceae

A sugi
Not described to species

Glyptostrobus sp.[18]

Schorn & Wehr

1996

Cupressaceae

A Chinese swamp cypress
Not described to species

Metasequoia occidentalis[19][20]

(Newberry) Chaney

1987

Cupressaceae

A dawn redwood

Metasequoia occidentalis

Picea sp.[18]

Schorn & Wehr

1996

Pinaceae

A spruce
Not described to species

Pinus latahensis[7]

Berry, 1929

1929

Pinaceae

A 5 needle pine

Pinus latahensis

"Pinus macrophylla"[7]

Berry, 1929

1929

Pinaceae

A 3 needle pine, jr homoym to Pinus macrophylla Lindley 1839

"Pinus macrophylla"

Pinus monticolensis[7]

Berry, 1929

1929

Pinaceae

A pine seed morphogenus

Pinus monticolensis

Pinus tetrafolia[7]

Berry, 1929

1929

Pinaceae

A possible 4 needled pine,
Noted by Berry as
"highly improbable that this should represent a distinct botanic species"

Pseudolarix wehrii[21]

Gooch, 1992

1940

Pinaceae

A golden larch,
originally described as Pseudolarix americana.[21]

Pseudolarix wehrii

Sequoia affinis[19]

Lesquereux

1935

Cupressaceae

A coast redwood,
Reported as "Sequioa langsdorfii" by Brown, 1935

Sequoia affinis

Taxodium dubium[7][19]

(Sternberg) Heer, 1855

1929

Cupressaceae

A bald cypress

Taxus sp.[18]

Schorn & Wehr

1996

Taxaceae

A yew
Not described to species

Thuja sp.[18]

Schorn & Wehr

1996

Cupressaceae

An arborvitae
Not described to species

Torreya bonseri[7][22]

(Knowlton) LaMotte, 1926

1929

Taxaceae

A plum-yew relative

Tsuga sp.[18]

Schorn & Wehr

1996

Cupressaceae

A hemlock
Not described to species

Flowering plants

Rhus hybrid leaf with lobed terminal leaflet

Over a dozen different Rosaceae genera, both extant and extinct, have been identified in the formation. These fossils are some of the oldest reliable macrofossils (excluding ancient fossil pollen) for the family.[23] Fossils of Sorbus genus and Rhus genus leaves that show evidence of being interspecies hybrids have been noted from the formation. Flynn, DeVore and Pigg in 2019 described four species of sumac which formed multiple hybrids.[24] Between three and four Trochodendraceae species that have been described from the Klondike Mountain Formation. Broadly circumscribed four species in three genera have been identified at Republic, Paraconcavistylon wehrii, Pentacentron sternhartae, Tetracentron hopkinsii, and Trochodendron nastae. Additionally the species Trochodendron drachukii is known from related Kamloops group shales at the McAbee Fossil Beds near Cache Creek, British Columbia. Manchester et al. 2018 noted that Tr. drachukii is likely the fruits of Tr. nastae, while Pe. sternhartae are likely the fruits of Te. hopkinsii.[25] If fossils of the fruits and foliage in attachment are found, that would bring the species count down to three whole plant taxa.[25] Additionally, the extinct genus Nordenskioldia is also known from the formation. The placement of Nordenskioldia is debated, with some treatments placing it into Trochodendraceae, while a 2020 analysis placed it outside of the crown-group Trochodendaceae.[26] Wesley Wehr in 1994 reported Bignoniaceae seeds along with a single Rubiaceae fruit and an isolated Fabaceae leaf.[27] An update of the floral list by Wehr and Manchester published in 1996 added an additional fifteen taxa identified from reproductive structures such as flowers fruits or seeds.[28] One of the fossils illustrated was tentatively identified as from the banana genus Ensete while another was placed in the extinct myrtle genus Paleomyrtinaea. However further fossil finds resulted in the re-identification of the first as a rhizome section likely from the water-lily Nuphar carlquistii, while the second is a seed mass from the same water-lilly.[29]

Name Authors Year Family Notes Images

Acer arcticum[30]

Heer, 1876

1987

Sapindaceae

A maple

Acer hillsi[30]

Wolfe & Tanai, 1987

1987

Sapindaceae

A maple

Acer republicense[30]

Wolfe & Tanai, 1987

1987

Sapindaceae

A maple

Acer spitzi[30]

Wolfe & Tanai, 1987

1987

Sapindaceae

A maple

Acer stonebergae[30]

Wolfe & Tanai, 1987

1987

Sapindaceae

A maple

Acer toradense[30]

Wolfe & Tanai, 1987

1987

Sapindaceae

A maple

Acer washingtonense[30]

Wolfe & Tanai, 1987

1987

Sapindaceae

A maple

Acer wehri[30]

Wolfe & Tanai, 1987

1987

Sapindaceae

A maple

Alnus parvifolia[20]

(Berry) Wolfe & Wehr, 1929

1929

Betulaceae

An Alder

Alnus parvifolia

Barghoornia oblongifolia[20]

Wolfe & Wehr, 1987

1987

Burseraceae

An extinct Bursera relative

Barghoornia oblongifolia

Betula leopoldae[20]

Wolfe & Wehr, 1987

1987

Betulaceae

A birch

Betula leopoldae

Bohlenia americana[20]

(Brown) Wolf & Wehr, 1935

1935

Sapindaceae

An extinct sapindale

Bohlenia americana

Calycites ardtunensis[27]

Crane, 1988

1995

incertae sedis

A winged fruit of unidentified affinities

Calycocarpum sp.[28]

(Nuttall) Spach, 1839

1996

Menispermaceae

A moonseed
Not described

Carpinus perryae[31]

Pigg, Manchester, & Wehr, 2003

2003

Betulaceae

A hornbeam

Cedrelospermum sp.[28]

Saporta, 1889

1996

Ulmaceae

An elm relative
Not described

Cercidiphyllum obtritum[20]

(Dawson) Wolfe & Wehr, 1890

1935

Cercidiphyllaceae

A katsura

Cercidiphyllum obtritum

Comptonia columbiana[20]

Dawson, 1890

1890

Myricaceae

A Comptonia

Comptonia columbiana

Corylopsis reedae[32]

Radtke, Pigg, & Wehr, 2001

2001

Hamamelidaceae

A winter-hazel

Corylus johnsonii[31]

Pigg, Manchester, & Wehr, 2003

2003

Betulaceae

A hazel nut

Corylus johnsonii

Craigia sp.[27]

Wehr, 1995

1995

Tiliaceae

A Cragia species
Not described

Cruciptera simsonii[33]

(Brown) Manchester

1991

Juglandaceae

A walnut family relative.

Deviacer sp.[27]

Wehr, 1995

Polygalaceae

A milkwort relative
Not described

Dillhoffia cachensis[34]

Manchester & Pigg, 2008

2008

incertae sedis

A flower of unknown relationships

Dillhoffia cachensis

Dipteronia brownii[35]

McClain & Machester, 2001

2001

Sapindaceae

A Dipteronia

Dipteronia brownii

Eucommia montana[36]

Brown, 1940

1997

Eucommiaceae

A "hard rubber tree"

Eucommia montana

Fagopsis undulata[20]

(Knowlton) Hollick, 1899

1987

Fagaceae

A beech

Fagopsis undulata

Fagus langevinii[37]

Manchester & Dillhoff, 2004

2004

Fagaceae

A beech

Florissantia quilchenensis

(Mathewes & Brooke) Manchester

Malvaceae

A chocolate relative

Florissantia quilchenensis

Fothergilla malloryi[32]

Radtke, Pigg, & Wehr, 2005

2005

Hamamelidaceae

A witch alder

Fothergilla malloryi

Koelreuteria dilcheri[38]

Wang et al., 2012

2012

Sapindaceae

A Koelreuteria species

Koelreuteria dilcheri

Langeria magnifica[20]

Wolfe & Wehr, 1987

1987

Hamamelidaceae

A witch hazel relative

Langeria magnifica

Macginicarpa sp.[27]

Manchester, 1986

1995

Platanaceae

A plane tree
Not described

Macginitiea gracilis[20]

(Lesquereux) Wolfe & Wehr

1929

Platanaceae

A plane tree

Macginitiea gracilis

Meliosma sp.[28]

Colebrook, 1819

1996

Sabiaceae

A Meliosma species
Not described

Nordenskioldia sp.[27]

Wehr, 1995

1995

Trochodendraceae?[26]

A trochodendroid of uncertain placement.
Not described.

Nuphar carlquistii[29]

DeVore, Taylor, & Pigg, 2015

2015

Nymphaeaceae

A waterlily,
Rhizome sections were first identified as Ensete.

Nuphar carlquistii seeds

Oemleria janhartfordae[23]

Benedict, DeVore, & Pigg, 2011

2011

Rosaceae

An Osoberry

Orontium wolfei[39]

Bogner, Johnson, Kvaček & Upchurch, 2007

2007

Araceae

A golden club

Orontium wolfei

Palaeocarpinus barksdaleae[31]

Pigg, Manchester, & Wehr, 2003

2003

Betulaceae

A birch relative

Palaeocarpinus barksdaleae

Paleoallium billgenseli[40]

Pigg, Bryan, & DeVore, 2018

2018

Amaryllidaceae

An onion relative

Paleoallium billgenseli

Palaeophytocrene sp.[27]

Wehr, 1995

1995

Icacinaceae

A Phytocrene relative
Not described

Paraconcavistylon wehrii[25][26]

(Manchester et al., 2018)

2018

Trochodendraceae

A Trochodendrale first described as "Concavistylon" wehrii
moved to a new genus in 2020.

Pentacentron sternhartae[25]

Manchester et al., 2018

2018

Trochodendraceae

A Trochodendrale

Pentacentron sternhartae

Photinia pageae[20]

Wolfe & Wehr, 1987

1987

Rosacaea

A Christmas-berry relative

Photinia pagae

Plafkeria sp.[27]

Wehr, 1995

1995

Tiliaceae

A linden relative
Not described

Platananthus[41]

Pigg & Wehr, 2002

2002

Platanaceae

A sycamore stamen head
Not described

Populus lindgreni[7]

Knowlton, 1898

1929

Salicaceae

A cottonwood

Prunus cathybrownae[23]

Benedict, DeVore, & Pigg, 2011

2011

Rosaceae

A cherry relative

Prunus cathybrownae

Pterocarya sp.[27]

Wehr, 1995

1995

Juglandaceae

A wing nut
Not described

Pteroceltis[41]

Pigg & Wehr, 2002

2002

Celtidaceae

A cannabaceous fruit
Not described

Pteronepelys wehrii[42]

Manchester

1994

incertae sedis

A samara of uncertain affinities.

Pteronepelys wehrii

Quercus sp.[27]

Wehr, 1995

1995

Fagaceae

An oak
Not described

Republica hickeyi[20]

Wolfe & Wehr, 1987

1987

incertae seids

An incertae sedis angiosperm
possibly of Hamamelididae affiliations

Republica hickeyi

Rhus boothillensis[24]

Flynn, DeVore, & Pigg, 2019

2019

Anacardiaceae

A sumac,
formed hybrids with the other Rhus species in the Klondike Mountain Formation

Rhus boothillensis

Rhus garwellii[24]

Flynn, DeVore, & Pigg, 2019

2019

Anacardiaceae

A sumac,
formed hybrids with the other Rhus species in the Klondike Mountain Formation

Rhus garwellii

Rhus malloryi[20][24]

(Wolfe and Wehr) Flynn, DeVore and Pigg

1987

Anacardiaceae

A sumac,
formed hybrids with the other Rhus species in the Klondike Mountain Formation

Rhus malloryi

Rhus republicensis[24]

Flynn, DeVore, & Pigg, 2019

2019

Anacardiaceae

A sumac,
formed hybrids with the other Rhus species in the Klondike Mountain Formation

Sabia sp.[28]

Colebrook, 1819

1996

Sabiaceae

A Sabia species
Not described

Sassafras hesperia[20]

Berry, 1929

1929

Lauraceae

A sassafras

Sassafras hesperia

Schoepfia republicensis[20]

(LaMotte) Wolfe & Wehr, 1944

1929

Schoepfiaceae

A Schoepfia relative

Schoepfia republicensis

Sloanea[41]

Pigg & Wehr, 2002

2002

Elaeocarpaceae

An elaeocarpaceous fruit
Not described

Tetracentron hopkinsii[25]

Pigg et al., 2007

2018

Trochodendraceae

A Trochodendrale,
possibly the leaves of Pentacentron sternhartae

Tetracentron hopkinsii

Tilia johnsoni[20]

Wolfe & Wehr, 1987

1987

Malvaceae

A Linden

Tilia johnsoni

Trochodendron nastae[43]

Pigg, Wehr, & Ickert-Bond, 2001

2001

Trochodendraceae

A Trochodendron

Trochodendron nastae

Tsukada davidiifolia[20]

Wolfe & Wehr, 1987

1987

Nyssaceae

A dove-tree relative

Tsukada davidiifolia

Ulmus chuchuanus[44]

(Berry) LaMotte, 1926

2005

Ulmaceae

An elm species

Ulmus okanaganensis

Denk & Dillhoff, 2005

2005

Ulmaceae

An elm species

Ulmus okanaganensis

Vitis sp.[27]

Wehr, 1995

1995

Vitaceae

A grape
Not described

Arthropods

The insect fauna of the Klondike Mountain Formation includes representatives from over 13 orders, based on a 1992 estimate, including immature though adult specimens and both terrestrial and aquatic taxa.[45] The most prevalent orders are Diptera and Hemiptera, each making up approximately 30% of the fossil insects known in 1992. A solitary lepidopteran fossil has been recovered, but no full descriptive work has been made on the specimen, aside from a single PhD dissertation. Early examination placed the moth in the family Geometridae, but later work has identified it as the oldest member of the tiger moth subfamily Arctiinae.[46]

The neuropteran insects (lacewings and their allies) identified as of 2014 include species from the families Berothidae, Chrysopidae, Hemerobiidae, Ithonidae (including Polystoechotidae), Nymphidae, Osmylidae, and Psychopsidae.[5] A number of mecopteran species belonging to the families Cimbrophlebiidae, Dinopanorpidae, Eorpidae, and Panorpidae are also known.[47] Fossil wings first described in 2015 were identified as being from Susumanioidea stick-insects, a group that had been known from the Jurassic to the Paleocene only.[48] The order Dermaptera was first reported in 1992[45] and is known from a series of isolated partial specimens, mostly abdominal sections with the distinct anal forceps attached. Based on the forceps structuring the specimens were tentatively assigned to the modern family Forficulidae, as the oldest North American representatives of the family known at that time.[49] A list of Coleopteran families identified by 1992 included Carabidae, Cerambycidae, Chrysomelidae, Curculionidae, Dytiscidae, Elateridae and Lucanidae,[45] but the beetle fauna has not been described in depth yet, with only two weevil species having been fully described.[50][51] A review of the Okanagan highlands hymenoptera published in 2018 identified four "Symphyta" families in the formation Cimbicidae, Pamphiliidae, Siricidae, and Tenthredinidae. Of the "Apocrita" families thirteen are represented, the "Parasitica" families are Braconidae, Diapriidae Ichneumonidae, Proctotrupidae, and Roproniidae while the Vespodea families are Formicidae, Pompilidae, Scoliidae and Vespidae. Within Apoidea the "Spheciformes" families include Angarosphecidae and Sphecidae while Halictidae is the sole "Apiformes" family known from body fossils. Prunus and Ulmus leaves have been found having damage that is consistent with the damage pattern left by Megachilidae species bees when they remove sections of tissue for nest lining. There are several additional Apoidea fossils that were left as incertae sedis in the group based on the similarity between them and Paleorhopalosoma menatensis, a Paleocene species described from the Menat Formation Auvergne, France. The placement of P. menatensis is uncertain, having been initially described as a member of Rhopalosomatidae, but is possibly an Angarosphecidae or closely related taxon,based on the wing and body morphology.[52]

Name Authors Year Family Notes Images

Adamsochrysa wilsoni[53]

Makarkin & Archibald

2013

Chrysopidae

A green lacewing

Ainigmapsychops inexspectatus[5]

Makarkin & Archibald

2014

Psychopsidae?

A possible psychopsid lacewing

Ainigmapsychops inexspectatus

Allorapisma chuorum[54]

Makarkin & Archibald

2009

Ithonidae

A moth lacewing

Allorapisma chuorum

Antiquiala snyderae[55]

Archibald & Cannings

2019

Aeshnidae

A darner dragonfly

Aphrophora sp.[45]

Lewis, 1992

1992

Aphrophoridae

An aphrophorid spittlebug
Not described.

Cimbrophlebia brooksi[47]

Archibald

2009

Cimbrophlebiidae

A Cimbrophlebiid scorpionfly

Cimbrophlebia brooksi

Cimbrophlebia westae[47]

Archibald

2009

Cimbrophlebiidae

A Cimbrophlebiid scorpionfly

Cimbrophlebia westae

Dinokanaga andersoni[56]

Archibald

2005

Dinopanorpidae

A scorpion fly species

Dinokanaga andersoni

Dinokanaga dowsonae[56]

Archibald

2005

Dinopanorpidae

A scorpion fly species

Dinokanaga sternbergi[56]

Archibald

2005

Dinopanorpidae

A scorpion fly species

Eoceneithycerus carpenteri[50]

Legalov

2013

Brentidae

An Ithycerinae weevil

Eoprephasma hichensi[48]

Archibald & Bradler

2015

Susumanioidea

A stick insect species

Eorpa elverumi[57]

Archibald, Mathewes, & Greenwood

2013

Eorpidae

A mecopteran scorpionfly

Eorpa elverumi

?Eorpa ypsipeda[57]

Archibald, Mathewes, & Greenwood

2013

Eorpidae

A mecopteran scorpionfly, tentatively identified

Possible E. ypsipeda[57]

Ithyceroides klondikensis[51]

Legalov

2015

Brentidae

An Ithycerinae weevil

Idemlinea versatilis[55]

Archibald & Cannings

2019

Aeshnidae

A darner dragonfly

Klondikia whiteae[58]

Dlussky & Rasnitsyn

2003

Formicidae

An ant

Metanephrocerus belgardeae[59]

Archibald, Kehlmaier, & Mathewes

2014

Pipunculidae

A pipunculid

Metanephrocerus belgardeae

Myrmeciites[60]

Archibald, Cover, & Moreau

2006

Formicidae

A bulldog ant form genus

Myrmeciities sp.

Neoephemera antiqua[61]

Sinitchenkova

1999

Neoephemeridae

A mayfly

Nymphes georgei[62]

Archibald, Makarkin, & Ansorge

2009

Nymphidae

A nymphid lacewing

Nymphes georgei

Oecophylla kraussei[63]

(Dlussky & Rasnitsyn)

1999

Formicidae

An ant, described as Camponotites kraussei,
Moved to Oecophylla kraussei in 2017[64]

Palaeopsychops marringerae[65]

Archibald & Makarkin, 2006

2006

Ithonidae[66]

A giant lacewing

Palaeopsychops marringerae

Palaeopsychops timmi[65]

Archibald & Makarkin, 2006

2006

Ithonidae[66]

A giant lacewing

Palaeopsychops timmi

Palecphora sp.[45]

Lewis, 1992

1992

Cercopidae

A cercopid froghopper
Not described.

Petrolystra sp.[45]

Lewis, 1992

1992

Aphrophoridae

An aphrophorid spittlebug
Not described.

Polystoechotites barksdalae[60]

Archibald & Makarkin, 2006

2006

Ithonidae[66]

A giant lacewing

Polystoechotites barksdalae

Polystoechotites falcatus[60]

Archibald & Makarkin, 2006

2006

Ithonidae[66]

A giant lacewing

Polystoechotites falcatus

Polystoechotites lewisi[60]

Archibald & Makarkin, 2006

2006

Ithonidae[66]

A giant lacewing

Polystoechotites lewisi

Proneuronema wehri[67][68]

Makarkin, Archibald, & Oswald

2003

Hemerobiidae

A hemerobiid lacewing, originally placed in Cretomerobius,
moved to Proneuronema in 2016

Propalosoma gutierrezae[63]

Dlussky & Rasnitsyn

1999

Formicidae

An ant, described as a Rhopalosomatidae wasp,
moved to myrmeciinae in 2018[69]

Propalosoma gutierrezae

Ulteramus republicensis[70]

Archibald & Rasnitsyn

2015

Pamphiliidae

A parasitic wasp

Ypshna brownleei[55]

Archibald & Cannings

2019

Aeshnidae

A darner dragonfly

"Forficulid species 1"[49]

Lewis, 1994

1994

Forficulidae?

A possible forficulid earwig with long cerci
Not described

"Forficulid species 2"[49]

Lewis, 1994

1994

Forficulidae?

A possible forficulid earwig with short cerci
Not described

Vertebrates

Five species of fish have been identified from the formation, four of which are known from skeletal elements, while the fifth is only known from isolated scales.[71] Of the five species, two are unique to the formation, Hiodon woodruffi and Libotonius pearsoni were both described by paleoichthyologist Mark V. H. Wilson in 1978 and 1979 respectively. The other three species, "Amia" hesperia, Amyzon aggregatum, and Eosalmo driftwoodensis, were first described from Okanagan Highlands formations in British Columbia and subsequently also identified from Ferry County fossils. The first notation of fish fossils in the Republic area was by Joseph Umpleby in his 1910 visit to the area, who collected fish near the Tom Thumb Mine, and sent them to the National Museum of Natural History. After examining the fossils, Charles R. Eastman listed the specimens as belonging to the extinct species Amyzon brevipinne in his Fossil fishes in the collection of the United States National Museum.[72] Research tapered off until a series of fish were collected in the Toroda Creek Graben northwest of Republic by R. C. Pearson during his compilation of the Geologic map of the Bodie Mountain quadrangle, Ferry and Okanogan Counties, Washington. The fossils were tentatively identified by paleoichthyologist David Dunkle as members of the genera Amyzon, Tricophanes, Erismatopterus and an undefined salmonid.[73] Pearson sent almost all of the specimens collected to the Smithsonian, but the fossils were never accessioned into the collections there and are now considered lost. He did retain one fossil from the initial collection which was later donated to the USGS collections. The largest single work on the fish of the Okanagan Highlands was published by Mark Wilson in 1977 and covered fossils collected from the known British Columbian Okanagan Highlands fossil sites of the time.[74] While not covering the Washington State fossils, Wilson named two of the species that are currently recognized from the Klondike Mountain Formation Amyzon aggregatum and Eosalmo driftwoodensis. Additionally scales attributed to the genus Amia were discussed and the genus Libotonius was named from fossils in the Allenby Formation.[75][76] In the late 1960s a collection of fish from near the Tom Thumb Mine in Republic was compiled by resident R. Woodward. During the summers of 1976 and 1977 the University of Alberta conducted field collecting in both the Republic and Toroda Creek areas, along with the donation of the Woodward collection, yielded a number of fossil catostomids, along with a single percopsid, a salmonid, a hiodontid, and an Amia scale. The hiodontids were subsequently described as the species Eohiodon woodruffi in 1978 based on differences between the Tom thumb Tuff fossils and those found in British Columbian sites.[73] A year later the percopsid fossils were also described as Libotonius pearsoni, extending the range of the genus south from the Allenby Formation.[75]

Bird fossils are limited to mostly isolated feathers that are preserved in the finer grained strata of the lake bed, though one partial bird skeleton has also been recovered.

Name Authors Year Family Notes Images

"Amia" hesperia

Wilson

1977

Amiidae

A bowfin

Amyzon aggregatum

Wilson

1977

Catostomidae

A sucker

Amyzon aggregatum

Eosalmo driftwoodensis[77]

Wilson

1977

Salmonidae

A Salmon

Eosalmo driftwoodensis

Hiodon woodruffi[73]

Wilson

1978

Hiodontidae

A mooneye, first described as "Eohiodon" woodruffi.[73]

Hiodon woodruffi

Libotonius pearsoni[75]

Wilson

1979

Libotoniidae

A sand roller relative.

"Unnamed"[78]

Mayr et al.

2019

Aves incertae sedis

indeterminate feathers and a skeleton

Unidentified feather

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