Toxochelys - Wikipedia

Toxochelys Temporal range: Late Cretaceous

Fossil specimen, Smithsonian National Museum of Natural History
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Family:	†Toxochelyidae
Genus:	†Toxochelys Cope, 1873
Species
T. bauri T. browni T. latiremus T. weeksi

Toxochelys (TOKS uh KEE leez^[1]) is an extinct genus of marine turtle from the Cretaceous period. It is the most commonly found fossilized turtle species in the Smoky Hill Chalk, in western Kansas.^[2] Toxochelys was about 2 m (6 ft) in length. There are five known species in the genus: Toxocheys bauri, Toxochelys browni, Toxochelys latiremus, Toxochelys weeks, and Toxochelys moorevillensis.^[3] Phylogenetic analysis shows that Toxochelys belong to an extinct lineage of turtles transitional between modern sea turtles and other turtles.^[4]

References

^ www.nhm.org Retrieved on May 12, 2008.
^ www.oceansofkansas.com Retrieved on May 12, 2008
^ zipcodezoo.com Retrieved on May 12, 2008.
^ Kear BP, Lee MS (March 2006). "A primitive protostegid from Australia and early sea turtle evolution". Biol. Lett. 2 (1): 116–9. doi:10.1098/rsbl.2005.0406. PMC 1617175. PMID 17148342.

Cope, E. D. 1873. [On Toxochelys latiremis]. Proceedings of the Academy of Natural Sciences of Philadelphia 25:10.
Hay, O. P. 1896. On the skeleton of Toxochelys latiremis. Publ. Field Columbian Museum, Zoological Ser. (later Fieldiana: Zoology), 1(5):101–106, pls. 14 &15.
Case, E.C. 1898. Toxochelys. The University Geological Survey of Kansas, Part IV. 4:370–385. pls. 79–84.
Hay, O.P. 1903. A revision of the species of the family of fossil turtles called Toxochelyidae, with descriptions of two new species of Toxochelys and a new species of Porthochelys. Bulletin of the American Museum of Natural History 21(10):177–185.
R. Zangerl. 1953. The vertebrate fauna of the Selma Formation of Alabama. Part IV. The turtles of the family Toxochelyidae. Fieldiana: Geology Memoirs 3(4):145–277
Nicholls, E.L. 1988. New material of Toxochelys latiremis Cope, and a revision of the genus Toxochelys (Testudines, Chelonoidea). Journal of Vertebrate Paleontology, 8(2):181–187.
Druckenmiller, P. S., A. J. Daun, J. L. Skulan and J. C. Pladziewicz. 1993. Stomach contents in the upper Cretaceous shark Squalicorax falcatus. Journal of Vertebrate Paleontology. 13(supplement. to no. 3):33A.
Carrino, M.H. 2007. Taxonomic comparison and stratigraphic distribution of Toxochelys (Testudines: Cheloniidae) of South Dakota. pp. 111–132 in Martin, J.E. and Parris D.C. (eds.), The Geology and Paleontology of the Late Cretaceous Marine Deposits of the Dakotas. Geological Society of America, Special Paper 427.
Konuki, R. 2008. Biostratigraphy of sea turtles and possible bite marks on a Toxochelys (Testudine, Chelonioidea) from the Niobrara Formation (Late Santonian), Logan County, Kansas and paleoecological implications for predator–prey relationships among large marine vertebrates. Unpublished Masters thesis, Fort Hays State University, Hays, Kansas, 141 pp, Appen. I-VI.
Matzke, A.T. 2008. A juvenile Toxochelys latiremis (Testudines, Cheloniidae) from the Upper Cretaceous Niobrara Formation of Kansas, USA. Neues Jahrbuch für Geologie und Paläontologie. Abhandlungen 249(3):371–380.
Matzke, A.T. 2009. Osteology of the skull of Toxochelys (Testudines, Chelonioidea) (with 25 text-figures). Palaeontographica Abteilung A 288(4):93–150.

External links

v t e Extinct turtles
Jurassic	Achelonia Americhelydia Bashuchelys Basilochelys Caribemys Chelonides Dinochelys Dorsetochelys Eurysternum Eurysternidae Hispaniachelys Hydropelta Idiochelys Neusticemys Parachelys Palaeomedusa Plesiochelys Pleurosternidae Selenemys Sinaspideretes Tacuarembemys Thalassemys Uluops
Cretaceous	Allopleuron Atlantochelys Angolachelys Anomalochelys Araripemys Araripemydidae Archelon Arvinachelys Aurorachelys Baenoidea Basilemys Bouliachelys Caririemys Cearachelys Chelosphargis Chubutemys Chupacabrachelys Compsemys victa Corsochelys Cratochelone Ctenochelys Desmatochelys Desmatochelys padillai Emarginachelys Foxemys Gigantatypus Gilmoremys Glyptops Hoplochelys Hoyasemys Iserosaurus Jeholochelys Jiangxichelys Kinkonychelys Kirgizemys Khunnuchelys Kurmademys Leyvachelys Liaochelys Lindholmemydidae Lomalatachelys Nanhsiungchelyidae Neurankylus Notochelone Ocepechelon Ordosemys Paracryptodira Patagoniaemys Peritresius Polysternon Pleurosternon Prochelidella Protostega Protostegidae Pneumatoarthrus Rhinochelys Sankuchemys Santanachelys Sinemys Terlinguachelys Toxochelys Trionyx kansaiensis Yaminuechelys
Paleogene and Neogene	Acherontemys Allaeochelys Anosteira Arabemys Argillochelys Axestemys Baenidae Banhxeochelys Berruchelus Bothremydidae Carbonemys Carolinochelys Cedrobaena Centrochelys atlantica Centrochelys vulcanica Cerrejonemys Chelodina murrayi Chelydrops Chelydropsis Chisternon Cosmochelys Elseya uberrima Elseya nadibajagu Eochelone Eosphargis Euclastes Foxemys Galianemys Gamerabaena Glarichelys Hadrianus Hesperotestudo Hutchemys Itilochelys Kinosternon pojoaque Lapparentemys Macrocephalochelys Manchurochelys Megalochelys (Megalochelys atlas) Mesodermochelys Mexichelys Miocaretta Niolamia Pacifichelys Palaeoamyda Palatobaena Peckemys Phrynops paranaensis Planiplastron Plesiobaena Procolpochelys Protochelydra Protosphargis Psephophorus Puppigerus Rheodytes devisi Rhinoclemmys panamaensis Stupendemys Stylemys Syllomus Tasbacka Trionyx ikoviensis Warkalania
Pleistocene	Centrochelys burchardi Centrochelys robusta Hesperotestudo mlynarskii Hesperotestudo percrassa Ninjemys Meiolaniidae Meiolania
Recent	Cylindraspis species: Domed Mauritius giant tortoise Domed Rodrigues giant tortoise Réunion giant tortoise Saddle-backed Mauritius giant tortoise Saddle-backed Rodrigues giant tortoise Aldabrachelys abrupta Aldabrachelys grandidieri Chelonoidis abingdonii Lonesome George Kinosternon arizonense

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra