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Cyrtodactylus

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Cyrtodactylus
Marbled bow-fingered gecko (Cyrtodactylus marmoratus)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Family: Gekkonidae
Subfamily: Gekkoninae
Genus: Cyrtodactylus
Gray, 1827
Species

Over 300, see text.

Cyrtodactylus (Greek κυρτος kurtos "curved", from κυπτω kuptō "to stoop"; δακτυλος daktulos "finger, toe")[citation needed] is a diverse genus of Asian geckos, commonly known as bent-toed geckos, bow-fingered geckos, and forest geckos. The genus has at least 300 described species as of 2020,[1] which makes it the largest of all gecko genera.

Description

Instead of possessing dilated digits like other geckos, members of Cyrtodactylus have slender, curved toes to which the common names for the genus can be attributed.

Taxonomy

The phylogenetic relationships with this genus and the genus Geckoella has not been resolved to date. Based on morphology, Bauer (2002) suggested that it was a subgenus of Cyrtodactylus but a phylogenetic study by Feng et al. (2007), though with inadequate sampling of taxa, restored it to generic status once again.[2][3] All species of Geckoella have been placed here pending a more definitive study.

Phylogeny

The following phylogeny is from Pyron, et al. (2013),[4] and includes 22 Cyrtodactylus species. Cyrtodactylus is a sister group of Hemidactylus.[4]

Cyrtodactylus 

Species

The genus Cyrtodactylus contains more than 300 described species. In 2019, six new species of the Cyrtodactylus intermedius complex were identified in the Cardamom Mountains and Associated Highlands region of Southeast Asia.[5]

Species Authority Common name Geographic range
C. aaroni Günther & Rösler, 2003[6] New Guinea
C. adleri Das, 1997 Nicobar Islands, Andaman Sea, India
C. adorus Shea, Couper, Wilmer & Amey, 2011 [7] Queensland, Australia
C. aequalis Bauer, 2003[8] Mon State, Myanmar
C. agamensis (Bleeker, 1860) Agam bow-fingered gecko Sumatra, Indonesia
C. agusanensis Taylor, 1915 Mindanao bow-fingered gecko Mindanao Island, The Philippines
C. albofasciatus (Boulenger, 1885) Boulenger's Indian gecko western Ghats, India
C. amphipetraeus Chomdej, Suwannapoom, Pawangkhanant, Pradit, Nazarov, Grismer, & Poyarkov, 2020 Tak bent-toed gecko Tak, Thailand
C. angularis (M.A. Smith, 1921) angulated bow-fingered gecko Eastern Thailand
C. annandalei Bauer, 2003[8] Annandale's leaf-toed gecko Sagaing Division, Myanmar
C. annulatus Taylor, 1915[9] Annulated bow-fingered gecko Mindanao Island, The Philippines
C. arcanus Oliver, Richards, & Sistrom 2012 [10][11] Papua New Guinea
C. astrum Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012 [12][13] Starry bent-toed gecko Southern Thailand, Peninsular Malaysia
C. atremus Kraus & Weijola, 2019 Papua New Guinea
C. aunglini Murdoch, Grismer, Wood, Thura, Win, Grismer, Trueblood & Quah, 2018[5] Kyauk Nagar Cave bent-toed gecko Myanmar
C. auralensis Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5] Phnom Aural bent-toed gecko Phnom Aural, Cambodia
C. aurensis Grismer, 2005[14] Aur Island bent-toed gecko Aur Island, Malaysia
C. auribalteatus Sumontha, Panitvong & Deein, 2010[15] Golden-belted bent-toed gecko Phitsanulok, Thailand
C. australotitiwangsaensis Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012 [12] Southern Titiwangsa bent-toed gecko Peninsular Malaysia
C. ayeyarwadyensis Bauer, 2003[8] Rakhine State, Myanmar
C. badenensis Nguyen, Orlov & Darevsky, 2006[16] Southern Vietnam
C. baluensis Mocquard, 1890[17] Balu bow-fingered gecko Mount Kinabalu, Borneo
C. bansocensis Luu, Nguyen, Le, Bonkowski & Ziegler, 2016[18] Ban Soc bent-toed gecko Laos
C. batik Iskandar, Rachmansah & Umilaela, 2011[19] Batik bent-toed gecko Indonesia
C. battalensis Khan, 1993 Reticulate plump-bodied gecko Pakistan
C. batucolus L.L. Grismer, Onn, J.L. Grismer, Wood & Belabut, 2008[20] Besar Island bent-toed gecko Pulau Besar of the Water Islands Archipelago off the west coast of Melaka, Peninsular Malaysia
C. bayinnyiensis Grismer, Wood Jr., Thura, Quah, Murdoch, Grismer, Herr, Lin & Kyaw, 2018 Bayin Nyi Cave bent-toed gecko Myanmar
C. bhupathyi Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018[21] Northeast India
C. bichnganae Tri & Grismer, 2010 Bich Ngan's bent-toed gecko Vietnam
C. bidoupimontis Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler, 2012[22] Vietnam
C. bintangrendah Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012[12] Bintang lowland bent-toed gecko Peninsular Malaysia
C. bintangtinggi Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels, 2012[12] Peninsular Malaysia
C. biordinis Brown & Mccoy, 1980[23] Guadalcanal bow-fingered gecko Mount Austen, Guadalcanal, Solomon Islands
C. bobrovi Nguyen et al., 2015[24] Bobrov's bent-toed gecko Vietnam
C. bokorensis Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5] Bokor Plateau, Cambodia
C. boreoclivus Oliver, Krey, Mumpuni & Richards, 2011[25] Papua New Guinea
C. brevidactylus Bauer, 2002[26] Myanmar
C. brevipalmatus (M.A. Smith, 1923)[27] Kampuchea bow-fingered gecko, short-hand forest gecko Khao Luang, Nakhon Si Thammarat, Peninsular Thailand
C. buchardi David, Teynie & Ohler, 2004[28] Laos
C. bugiamapensis Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler, 2012[22] Vietnam
C. calamei Luu, Bonkowki, Nguyen, Le, Schneider, Ngo & Ziegler, 2016 Calame's bent-toed gecko central Laos
C. camortensis Chandramouli, 2020 Kamorta, Nicobar Islands
C. caovansungi Orlov & Quang Truong, 2007 [29] Vietnam
C. capreoloides Rösler, Richards & Günther, 2007 Papua New Guinea
C. cardamomensis Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5] Cardamom Mountains bent-toed gecko Cardamom Mountains, Cambodia
C. cattienensis Geissler, Nazarov, Orlov, Böhme, Phung, Nguyen & Ziegler, 2009[30] Cattien bent-toed gecko Vietnam
C. cavernicolus Inger & King, 1962 Sarawak bow-fingered gecko, Niah Cave gecko Sarawak
C. cayuensis Li, 2007 Cayu bent-toed gecko China (Cayu County, Xizang)
C. celatus Kathriner et al., 2014[31] Indonesia
C. chamba AgarwaL, Khandekar & Bauer, 2018 Chamba bent-toed gecko Himachal Pradesh, India
C. chanhomeae Bauer, Sumontha & Pauwels, 2003[32] Thailand
C. chaunghanakwaensis Grismer, Wood Jr., Thura, Quah, Murdoch, Grismer, Herr, Lin & Kyaw, 2018 Chaunghanakwa Hill bent-toed gecko Myanmar
C. chauquangensis Quang, Orlov, et al. 2007[33] Vietnam
C. chrysopylos Bauer, 2003[8] Panluang-Pyadakin Wildlife Sanctuary, Shan State, Myanmar
C. chungi Ostrowski, Le, Ngo, Pham, Phung, Nguyen & Ziegler, 2021[34] Chung's bent-toed gecko Binh Thuan province, Vietnam
C. collegalensis (Beddome, 1870) forest spotted gecko, Kollegal ground gecko India
C. condorensis (M.A. Smith, 1921) Pulo Condore bow-fingered gecko, Con Dao bow-fingered gecko Pulo Condore Island, Vietnam
C. consobrinoides Annandale, 1905 southern Myanmar
C. consobrinus W. Peters, 1871 Peters's bow-fingered gecko, thin-banded forest gecko Malaysian peninsula, Borneo
C. cracens Batuwita & Bahir, 2005 Sri Lanka
C. crustulus Oliver, Hartman, Turner, Wilde, Austin & Richards, 2020 Manus bent-toed gecko Papua New Guinea (Manus Island)
C. cryptus Heidrich, Rösler, Thanh, Böhme & Ziegler, 2007[35] Vietnam
C. cucdongensis Schneider & Phung TM, 2014[36] Cucdong bent-toed gecko Vietnam
C. cucphuongensis Ngo & Onn, 2011[37] Vietnam
C. culaochamensis Tri, Grismer, Thai & Wood, 2020 Cù Lao Chàm bent–toed gecko Vietnam
C. dammathetensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Dammathet cave bent-toed gecko Myanmar
C. darevskii Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov, 2014 Darevski's bent-toed gecko Laos
C. darmandvillei (Weber, 1890) Darmandville bow-fingered gecko Flores, Komodo, Pulau Kalao, Sumbawa, Sikka, Lombok in Indonesia
C. dati Tri, 2013[39] Dat bent-toed gecko Vietnam
C. dattanensis (Khan, 1980) plump banded gecko, Khan's bow-fingered gecko nw Pakistan
C. dattkyaikensis Grismer, Wood, Quah, Grismer, Thura, Oaks & Lin, 2020 Datt Kyaik Hill bent-toed gecko Myanmar
C. dayangbuntingensis Quah, Grismer, Wood Jr. & Sah, 2019 peninsular Malaysia
C. deccanensis (Günther, 1864) Günther's Indian gecko Northern Western Ghats of India
C. derongo Brown & Parker, 1973 Derongo bow-fingered gecko Papua, New Guinea
C. deveti (Brongersma, 1948) Moluccan bow-fingered gecko Moluccas, Indonesia
C. doisuthep Kunya et al. 2014[40] Doi Suthep bent-toed gecko Thailand
C. dumnuii Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome & Kunya, 2010[41] Dumnui's bent-toed gecko Thailand
C. durio Grismer, Anuar, Quah, Muin, Onn, Grismer & Ahmad, 2010[42] Durian bent-toed gecko Malaysia
C. edwardtaylori Batuwita & Bahir, 2005 Sri Lanka
C. eisenmanae Tri, 2008[43] Eisenman's bent-toed gecko Vietnam
C. elok Dring, 1979 Malaysia bow-fingered gecko western Malaysia
C. epiroticus Kraus, 2008 Papua New Guinea
C. equestris Oliver, Richards, Mumpuni & Rosler, 2016 Papua New Guinea
C. erythrops Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome & Kunya, 2009[44] Red-eyed bent-toed gecko Thailand
C. evanquahi Wood, Grismer, Muin, Anuar & Oaks, 2020 Evan Quah's banded bent-toed gecko Peninsular Malaysia
C. fasciolatus (Blyth, 1861)[45] banded bent-toed gecko Northwest India
C. feae (Boulenger, 1893) Myanmar
C. fraenatus (Günther, 1864) Sri Lanka bow-fingered gecko Sri Lanka
C. fumosus (F. Müller, 1895)[46] Sulawesi, Indonesia
C. gansi Bauer, 2003[8] Min Dat District, Chin State, Myanmar
C. gialaiensis Luu, Dung, Nguyen, Le & Ziegler, 2017 Vietnam
C. gordongekkoi Das, 1994 [47] Lombok, Indonesia
C. grismeri Tri, 2008 Grismer's bent-toed gecko Vietnam
C. guakanthanensis Grismer, Belabut, Quah, Onn & Wood, 2014[48] Gua Kanthan bent-toed gecko Malaysia
C. gubaot Welton, Siler, Linkem, Diesmos & Brown, 2010[49] the Philippines
C. gubernatoris (Annandale, 1913) Sikkimese bent-toed gecko Darjeeling, India
C. gunungsenyumensis Grismer et al., 2016[50] Gunung Senyum bent-toed gecko Malaysia
C. guwahatiensis Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 India
C. halmahericus Mertens, 1929 [51] Indonesia
C. hidupselamanya Grismer, Wood, Anuar, Grismer, Quah, Murdoch, Muin, Davis, Auilar, Klabacka, Cobos, Aowphol & Sites, 2016 Chiku bent-toed gecko peninsular Malaysia
C. hikidai Riyanto A, 2012[52] Bunguran island, Indonesia
C. himalayanus Duda & Sani, 1977 Himalaya bow-fingered gecko Jammu, India
C. himalayicus (Annandale, 1906) Himalaya bent-toed gecko India (West Bengal, Darjeeling)
C. hinnamnoensis Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler, 2016 Hinnamno bent-toed gecko Laos
C. hitchi Riyanto, Kurniati & Engilis, 2016 southeastern Sulawesi, Indonesia
C. hontreensis Tri, Grismer & Grismer, 2008[53] Hon Tre bent-toed gecko Vietnam
C. hoskini Shea, Couper, Wilmer & Amey, 2011[54] Queensland, Australia
C. houaphanensis Schneider, Luu, Sitthivong, Teynié, Le, Nguyen & Ziegler, 2020 Houaphan bent–toed gecko Laos
C. huongsonensis Luu, Do & Ziegler, 2011[55] Vietnam
C. huynhi Ngo & Bauer, 2008[56] Huynh's bent-toed gecko Vietnam
C. ingeri Hikida, 1990 Sabah bow-fingered gecko Sabah, Borneo
C. interdigitalis (Ulber, 1992) skin-toe forest gecko Thailand, Laos
C. intermedius Ulber, 1993 intermediate bow-fingered gecko Cambodia, Vietnam, and Thailand
C. inthanon Kunya et al., 2015[57] Doi Inthanon bent-toed gecko Thailand
C. irianjayaensis Rösler, 2001 Papua, New Guinea
C. irregularis (M.A. Smith, 1921) irregular bow-fingered gecko Vietnam
C. jaegeri Luu, Calame, Bonkowski, T. Nguyen & Ziegler, 2014[58] Laos
C. jaintiaensis Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 India
C. jambangan Welton, Siler, Diesmos & Brown, 2010[59] the Philippines
C. jarakensis L.L. Grismer, Onn, J.L. Grismer, Wood & Belabut, 2008[20] Jarak Island bent-toed gecko Pulau Jarak off the west coast of Perak, Peninsular Malaysia
C. jarujini (Ulber, 1992) Jarujin's forest gecko Laos and northern Thailand
C. jatnai Amarasinghe, Riyanto, Mumpuni & Grismer, 2020 Jatna's bent–toed gecko west Bali, Indonesia
C. jelawangensis Grismer, Wood, Anuar, Quah et al., 2014[60] Jelawang bent-toed gecko Thai-Malay Peninsula
C. jellesmae Boulenger, 1897 Kabaena bow-fingered gecko Masarang in Indonesia
C. jeyporensis (Beddome, 1877) Jeypore rock gecko eastern Ghats, India
C. kazirangaensis Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 India
C. khammouanensis Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov, 2014 Khammouane bent-toed gecko Laos
C. khasiensis (Jerdon, 1870) Khasi Hills bent-toed gecko India and northern Burma
C. khelangensis Pauwels, Sumontha, Panitvong & Varaguttanonda, 2014[61] Lampang bent-toed gecko Thailand
C. kimberleyensis Aaron & Doughty, 2012[62] Kimberley bent-toed gecko, East Montalivet Island gecko Australia
C. kingsadai Ziegler, Phung, Le & Nguyen, 2013[63] Vietnam
C. klugei Kraus, 2008 Papua New Guinea
C. kohrongensis Grismer, Onn, Oaks, Neang, Sokun, Murdoch, Stuart & Grismer, 2020 Koh Rong Island bent–toed gecko Cambodia
C. kunyai Pauwels, Sumontha, Keeratikiat & Phanamphon, 2014[64] Kunya's bent-toed gecko Thailand
C. laangensis Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5] Phnom Laang bent-toed gecko Phnom Laang, Cambodia
C. laevigatus Darevsky, 1964 Komodo Island
C. langkawiensis Grismer, Wood, et al., 2012[12] Langkawi Island bent-toed gecko Langkawi, Malaysia
C. lateralis Werner, 1896 Sumatra bow-fingered gecko Sumatra
C. lawderanus (Stoliczka, 1871) Lawder's bent-toed gecko northwestern India
C. leegrismeri Chan & Norhayati, 2010[65] Tenggol Island bent-toed gecko Malaysia
C. lekaguli Grismer et al., 2012[12] Southern Thailand
C. lenggongensis Grismer, Wood, Anuar, Grismer, Quah, Murdoch, Muin, Davis, Auilar, Klabacka, Cobos, Aowphol & Sites, 2016 Lenggong bent-toed gecko peninsular Malaysia
C. lenya Mulcahy, Thura & Zug, 2017 Lenya banded bent-toed gecko Myanmar
C. limajalur Davis, Bauer, Jackman, Nashriq & Das, 2019 five-banded bent-toed gecko East Malaysia
C. linnoensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Linno Cave bent-toed gecko Myanmar
C. linnwayensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Linn-Way bent-toed gecko Myanmar
C. lomyenensis Ngo & Pauwels, 2010[66] Lomyen bent-toed gecko Laos
C. loriae (Boulenger, 1897) Boulenger's bow-fingered gecko Papua New Guinea
C. louisiadensis (DeVis, 1892) ring-tailed gecko Louisiade Archipelago, Papua New Guinea
C. macrotuberculatus Grismer & Ahmad, 2008[67] tuberculate bent-toed gecko, large tubercled bent-toed gecko Malaysia
C. maelanoi Grismer, Rujirawan, Termprayoon, Ampai, Yodthong, Wood, Oaks, & Aowphol, 2020 Mae La Noi bent-toed gecko Mae Hong Son Province, Thailand
C. majulah Grismer, Wood & Lim 2012 Singapore; Indonesia
C. malayanus De Rooij, 1915 Borneo bow-fingered gecko Borneo
C. mamanwa Welton, Siler, Linkem, Diesmos & Brown, 2010[49][68] The Philippines
C. mandalayensis Mahony, 2009[69] Myanmar
C. manos Oliver, Karkkainen, Rösler & Richards, 2019 yellow-snouted bent-toed gecko Papua New Guinea
C. markuscombaii Darevsky, Helfenberger, Orlov, & Shah, 1998[70] striped gecko Nepal
C. marmoratus Gray, 1831 marbled bow-fingered gecko Thailand, Malay Peninsula, Indonesia, Papua New Guinea.
C. martini Tri, 2011[71] Martin's bent-toed gecko Vietnam
C. martinstolli Darevsky, Helfenberger, Orlov, & Shah, 1998[70] barred gecko Nepal
C. matsuii Hikida, 1990 Hikida's bow-fingered gecko,
Matsui's bent-toed gecko		 Sabah, Borneo
C. mcdonaldi Shea, Couper, Wilwer & Amey 2011 [72] Australia
C. medioclivus Oliver, Richards & Sistrom, 2012 [73] Papua New Guinea
C. meersi L. Grismer, Wood, Quah, Murdoch, M. Grismer, Herr, Espinoza, Brown & Lin, 2018 Bago Yoma bent-toed gecko Myanmar (Yangon)
C. metropolis Grismer, Wood, Onn, Anuar & Muin, 2014[74] Malaysia
C. mimikanus (Boulenger, 1914) false bow-fingered gecko, Mimika bow-fingered gecko,
Mimika bent-toed gecko		 Papua New Guinea
C. minor Oliver & Richards, 2012[75] Papua New Guinea
C. mombergi Grismer, Wood, Quah, Thura, Herr & Lin, 2019 Indawgyi bent-toed gecko Myanmar
C. montanus Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 India
C. muangfuangensis Sitthivong, Luu, Ha, Nguyen, Le & Ziegler, 2019 Muangfuang bent-toed gecko Laos
C. multiporus Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov, 2014 Multipored bent-toed gecko Laos
C. muluensis Davis, Bauer, Jackman, Nashriq & Das, 2019 Mulu bent-toed gecko East Malaysia
C. murua Kraus & Allison, 2006[76] Papua New Guinea
C. myaleiktaung Grismer, Wood, Thura, Win, Grismer, Trueblood & Quah, 2018 Mya Leik Taung bent-toed gecko Myanmar
C. myintkyawthurai L. Grismer, Wood, Quah, Murdoch, M. Grismer, Herr, Espinoza, Brown & Lin, 2018 Mt. Popa bent-toed gecko Myanmar (Mandalay region, Bago Yoma Range, Bago Region)
C. nagalandensis Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 India
C. naungkayaingensis Grismer, Wood, Thura, Quah, Murdoch, Grismer, Herr, Lin & Kyaw, 2018 Myanmar
C. nebulosus (Beddome, 1870) Clouded Indian gecko India
C. nepalensis Schleich & Kästle, 1998[77] Nepalese rock gecko Nepal
C. ngoiensis Schneider, Luu, Sittivong, Teynie, Le, Nguyen & Ziegler, 2020 Ngoi bent–toed gecko Laos
C. nicobaricus Chandramouli, 2020 Nicobar Islands
C. nigriocularis Nguyễn, Orlov & Darevsky, 2007 southern Vietnam
C. novaeguineae (Schlegel, 1844) New Guinea bow-fingered gecko New Guinea
C. nuaulu Oliver, Edgar, Mumpuni, Iskandar & Lilley, 2009[78] Indonesia
C. nyinyikyawi Grismer, Wood, Thura, Win & Quah, 2019 Shwe Settaw bent-toed gecko Myanmar
C. oldhami (Theobald, 1876) Oldham's bow-fingered gecko Myanmar and Thailand
C. otai Nguyen et al., 2015[24] Ota's bent-toed gecko Vietnam
C. pageli Schneider et al., 2011[79] Laos
C. pantiensis L.L. Grismer, Onn, J.L. Grismer, Wood & Belabut, 2008[20] Panti Mountain bent-toed gecko Gunung Panti Forest Reserve, Johor, Peninsular Malaysia
C. papilionoides Ulber & Grossmann, 1991 butterfly forest gecko Thailand
C. papuensis (Brongersma, 1934) Papua bow-fingered gecko Papua New Guinea
C. paradoxus Darevsky & Szczerbak, 1997 Kieng Giang, Phu Quoc, and Hon Thom Islands in Vietnam
C. payacola Johnson, Quah, Anuar, Muin, Wood, Grismer, Greer, Onn, Ahmad, Bauer & Grismer, 2012 [80] Malaysia
C. payarhtanensis Mulcahy, Thura & Zug, 2017 Tenasserim Mountain bent-toed gecko southern Myanmar
C. peguensis (Boulenger, 1893) Thai bow-fingered gecko, Pegu forest gecko Thailand, western Malaysia, and Myanmar
C. petani Riyanto, Grismer & Wood, 2015[81] Farmer's bent-toed gecko Indonesia
C. pharbaungensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Pharbaung Cave bent-toed gecko Myanmar
C. phetchaburiensis Pawels, Sumontha & Bauer, 2016 Thailand
C. philippinicus Steindachner, 1867 Philippine bow-fingered gecko northern Philippines
C. phnomchiensis Nyang, Henson & Stuart, 2020[82] Prey Lang bent-toed gecko[83] Prey Lang, Cambodia
C. phongnhakebangensis Ziegler, Rösler, Herrmann & Vũ, 2002 Phong Nha-Ke Bang bow-fingered gecko Vietnam
C. phuketensis Sumontha, Pawels, Kunya, Nitikul, Samphanthamit & Grismer, 2012 [84] Phuket bent-toed gecko Phuket, Thailand
C. phumyensis Ostrowski, Do, Le, Ngo, Pham, Nguyen, Nguyen & Ziegler, 2020 Phu My bent-toed gecko Vietnam
C. phuocbinhensis S. Nguyen, Le, Tran, Orlov, Lathrop, MacCulloch, Le, Jin, L. Nguyen, T. Nguyen, Hoang, Che, Murphy & Zang, 2013 [85] Vietnam
C. phuquocensis Tri, Grismer & Grismer, 2010[86][87] Phu Quoc bent–toed gecko Vietnam
C. pinlaungensis Grismer, Wood, Quah, Thura, Oaks & Lin, 2019 Pinlaung bent-toed gecko Myanmar
C. pronarus Shea, Couper, Wilmer & Amey, 2011[54] Australia
C. psarops Harvey et al., 2015[88] Speckle-faced bent-toed gecko Indonesia
C. pseudoquadrivirgatus Rösler, Nguyễn, Vũ, Ngô & Ziegler, 2008 central Vietnam
C. pubisulcus Inger, 1958 Inger's bow-fingered gecko Borneo
C. puhuensis Nguyen, Yang, Thi Le, Nguyen, Orlov, Hoang, Nguyen, Jin, Rao, Hoang, Che, Murphy & Zhang, 2014[89] Pù Hu bent-toed gecko Vietnam
C. pulchellus Gray, 1827 Malayan forest gecko Penang, Malaysia
C. pyadalinensis Grismer, Wood, Thura, Win & Quah, 2019 Myanmar
C. pyinyaungensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Pyinyaung bent-toed gecko Myanmar
C. quadrivirgatus Taylor, 1962 Taylor's bow-fingered gecko, four-striped forest gecko,
marbled bent-toed gecko		 Thailand, Malaysia, Singapore, and Indonesia
C. ramboda Batuwita & Bahir, 2005 Sri Lanka
C. ranongensis Sumontha, Pauwels, Panitvong, Kunya & Grismer, 2015[90] Ranong bent-toed gecko Thailand
C. redimiculus King, 1962 Palawan bow-fingered gecko Palawan, Philippines
C. rex Oliver, Richards, Mumpuni & Rösler, 2016 Papua New Guinea
C. rishivalleyensis Agarwal, 2016 Rishi Valley geckoella India
C. robustus Kraus, 2008 Papua New Guinea
C. roesleri Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh & Schmitz, 2010 [91] Roesler's bent-toed gecko Vietnam & Laos
C. rosichonarieforum Riyanto, Grismer & Wood, 2015[92] Rosichonariefi's bent-toed gecko Indonesia
C. rubidus (Blyth, 1861) red bow-fingered gecko Andaman Islands; Myanmar
C. rufford Luu et al., 2016[93] Rufford bent-toed gecko Laos
C. russelli Bauer, 2003[8] Htamanthi Wildlife Sanctuary, Sagaing Division and Indawgyi Wildlife Sanctuary, Rakhine State, Myanmar
C. sadanensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Sadan Cave bent-toed gecko Myanmar
C. sadansinensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Sadan Sin Cave bent-toed gecko Myanmar
C. sadleiri Wells & Wellington, 1984 Sadleir's bow-fingered gecko Christmas Island
C. saiyok Panitvong, Sumontha, Tunprasert & Pauwels, 2014[94] Sai Yok bent-toed gecko Thailand
C. salomonensis Rösler, Richards & Günther, 2007[95] Solomon Islands
C. samroiyot Pauwels & Sumontha, 2014[96] Sam Roi Yot bent-toed gecko Thailand
C. sangi Pauwels, Nazarov, Bobrov & Poyarkov, 2018 southern Vietnam
C. sanook Pauwels, Sumontha, Latinne & Grismer, 2013[97] Sanook bent-toed gecko Thailand
C. sanpelensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Sanpel Cave bent-toed gecko Myanmar
C. semenanjungensis Grismer & Leong, 2005 Peninsular bent-toed gecko western Malaysia
C. semiadii Riyanto, Bauer & Yudha, 2014[98] Indonesia
C. semicinctus Harvey et al., 2015[88] Kerinci bent-toed gecko Indonesia
C. septentrionalis Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 India
C. septimontium Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5] Bảy Núi bent-toed gecko Bảy Núi Hills, Vietnam
C. seribuatensis Youmans & Grismer, 2006 Seribuat bent-toed gecko western Malaysia
C. sermowaiensis (de Rooij, 1915) DeRooij's bow-fingered gecko New Guinea
C. serratus Kraus, 2007[99] Papua New Guinea
C. sharkari Grismer et al., 2014[60] Sharkari's bent-toed gecko Thai-Malay Peninsula
C. shwetaungorum Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Shwetaung bent-toed gecko Myanmar
C. sinyineensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Sin Yine Cave bent-toed gecko Myanmar
C. slowinskii Bauer, 2002 Myanmar
C. soba Batuwita & Bahir, 2005 Dumbara bent-toed gecko, Knuckles bent-toed gecko Sri Lanka
C. sommerladi Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler, 2016 Sommerlad's bent-toed gecko central Laos
C. soni Le, Nguyen, Le & Ziegler, 2016 Son's bent-toed gecko Vietnam
C. sonlaensis Nguyen, Pham, Ziegler, Ngo & Le, 2017 northwestern Vietnam
C. soudthichaki Luu, Calame, Nguyen, Bonkowski & Ziegler, 2015[100] Soudthichak's bent-toed gecko Laos
C. speciosus (Beddome, 1870) forest spotted gecko India
C. spelaeus Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov, 2014 Laos
C. spinosus Linkem, McGuire, Hayden, Setiadi, Bickford & Brown, 2008 Sulawesi, Indonesia
C. srilekhae Agarwal, 2016 Bangalore geckoella India
C. stresemanni Rösler & Glaw, 2008 Stresemann's bent-toed gecko Malaysia
C. subsolanus Batuwita & Bahir, 2005 Sinharaja bent-toed gecko Sri Lanka
C. sumonthai Bauer, 2002 Sumontha's gecko Thailand
C. sumuroi Welton, Siler, Linkem, Diesmos & Brown, 2010[49] the Philippines
C. surin Chan-ard & Makchai (2011)[101] Thailand
C. sworderi (M.A. Smith, 1925) Johore bow-fingered gecko, Kota-tinggi forest gecko,
Sworder's bent-toed gecko		 Malaysia
C. tahuna Riyanto, Arida & Koch, 2018[102] Sangir Island, Indonesia
C. takouensis Van Tri * Gamble, 2010[103] Takou bent-toed gecko Vietnam
C. tamaiensis (M.A. Smith, 1940)[104] Tamai bent-toed gecko Myanmar
C. tambora Riyanto, Mulyadi, McGuire, Kusrini, Febylasmia, Basyir & Kaiser, 2017 Tambora bent-toed gecko Sumbawa, Indonesia
C. tanahjampea Riyanto, Hamidy & McGuire, 2018 Tanahjampea bent-toed gecko Tanahjampea Island, Indonesia
C. tanim Nielsen & Oliver, 2017 Papua New Guinea
C. taungwineensis Grismer, Wood, Quah, Grismer, Thura, Oaks & Lin, 2020 Taung Wine Hill bent-toed gecko Myanmar
C. tautbatorum Welton, Siler, Diesmos & Brown, 2009[105] the Philippines
C. taybacensis Pham, Le, Ngo, Ziegler & Nguyen, 2019 Taybac bent-toed gecko Vietnam
C. taynguyenensis S. Nguyen, Le, Tran, Orlov, Lathrop, Macculloch, Le, Jin, L. Nguyen, T. Nguyen, Hoang, Che, Murphy & Zhang 2013[106] Vietnam
C. tebuensis Grismer, Anuar, Muin, Quah & Wood, 2013[107] Tebu Mountain bent-toed gecko Malaysia
C. teyniei David, Nguyen, Schneider & Ziegler, 2011[108] Laos
C. thathomensis Nazarov, Pauwels, Konstantinov, Chulisov, Orlov & Poyarkov, 2018 Thathom bent-toed gecko Laos
C. thirakhupti Pauwels, Bauer, Sumontha & Chanhome, 2004[109] Thirakhupt's bent-toed gecko Thailand
C. thochuensis Ngo & Grismer, 2012[110] Thochu bent–toed gecko Tho Chu Island, Vietnam
C. thuongae Phung, Schingen, Ziegler & T. Nguyen, 2014[111] Vietnam
C. thylacodactylus Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer, 2019[5] Phnom Dalai, Cambodia
C. tibetanus Boulenger, 1905 Tibet
C. tigroides Bauer, Sumontha & Pauwels, 2003[32] Tiger bent-toed gecko Thailand
C. timur Muin, Mohamed, Onn, Sumarli, Loredo & Heinz, 2014[60] Banjaran Timur bent-toed gecko Thai-Malay Peninsula
C. tiomanensis Das & Lim, 2000[112] Tioman Island bent-toed gecko Malaysia
C. triedrus (Günther, 1864) spotted bent-toed gecko, Sri Lanka gecko,
spotted bow-fingered gecko, spotted ground gecko		 Sri Lanka
C. trilatofasciatus Grismer & Wood, 2012[113] Cameron Highlands bent-toed gecko Malaysia
C. tripartitus Kraus, 2008 Papua New Guinea
C. tripuraensis Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 India
C. tuberculatus Lucas & Frost, 1900 Queensland, Australia
C. urbanus Purkayastha, Das, Bohra, Bauer & Agarwal, 2020 urban bent-toed gecko India (Assam)
C. varadgirii Agarwal et al., 2016[114] Giri's Geckoella Mumbai, India
C. variegatus Blyth, 1859 variegated bow-fingered gecko, Moulmein forest gecko Myanmar and Thailand
C. vilaphongi Schneider, Nguyen, Le, Nophaseud, Bonkowski & Ziegler, 2014 Laos
C. wakeorum Bauer, 2003[8] Southern Rakhine State, Myanmar
C. wallacei Hayden, Brown, Gillespie, Setiadi, Linkem, Iskandar, Umilaela, Bickford, Riyanto, Mumpuni & McGuire,2008 Sulawesi, Indonesia
C. wangkulangkulae Sumontha et al., 2014[115] Wangkulangkul's bent-toed gecko Thailand
C. wayakonei Nguyen, Kingsada, Rösler, Auer & Ziegler, 2010[116] Northern Laos & Yunnan, China [117]
C. welpyanensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Wel Pyan Cave bent-toed gecko Myanmar
C. wetariensis Dunn, 1927 Wetar bow-fingered gecko Wetar, Indonesia
C. yakhuna (Deraniyagala, 1945) northern Sri Lanka gecko, spotted bow-fingered gecko,
blotch bow-fingered gecko, demon gecko		 Sri Lanka
C. yangbayensis Tri & Onn, 2010[118] Yangbay bent-toed gecko Vietnam
C. yathepyanensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Lin, Kyaw & Lwin, 2017[38] Yathe Pyan Cave bent-toed gecko Myanmar
C. yoshii Hikida, 1990 Yoshi's bow-fingered gecko Sabah, Borneo
C. ywanganensis Grismer, Wood, Thura, Quah, Murdoch, M. Grismer, Herr, Espinoza & Lin, 2018 Myanmar (Shan)
C. zebraicus (Taylor, 1962) Thailand
C. zhaoermii Shi & Zhao 2010 [119] Tibet, China
C. ziegleri Nazarov, Orlov, Nguyen & Ho, 2008 [120] Vietnam
C. zugi Oliver, Tjaturadi, Mumpuni, Krey & Richards, 2008[121] Melanesia

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Further reading

  • Bauer, Aaron M. [in French] (2002). "Two New Species of Cyrtodactylus (Squamata: Gekkonidae) from Myanmar". Proceedings of the California Academy of Sciences. 53 (7): 73–86.
  • Gray, J.E. (1827). "A Synopsis of the Genera of Saurian Reptiles, in which some new Genera are indicated, and others reviewed by actual Examination". The Philosophical Magazine, or Annals of Chemistry, Mathematics, Astronomy, Natural History, and General Science. 2: 54–58. (Cyrtodactylus, new genus, p. 56).
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