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2010 in paleomammalogy

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This is an old revision of this page, as edited by 188.146.28.104 (talk) at 11:50, 26 November 2016 (Newly named eutherians). The present address (URL) is a permanent link to this revision, which may differ significantly from the current revision.

Newly named non-eutherian mammals

Name Status Authors Age Unit Location Notes Images

Acinacodus[1]

Valid

  • Lopatin
  • Maschenko
  • Averianov

Early Cretaceous

Western Siberia

 Russia

An amphidontid 'eutriconodont'.

Antechinus yammal[2]

Valid

  • Cramb
  • Hocknull

Pleistocene

 Australia

A marsupial belonging to the family Dasyuridae, a species of Antechinus.

Antechinus yuna[2]

Valid

  • Cramb
  • Hocknull

Pleistocene

 Australia

A marsupial belonging to the family Dasyuridae, a species of Antechinus.

Canchadelphys[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Didelphimorphia belonging to the superfamily Peradectoidea and the family Caroloameghiniidae. The type species is C. cristata.

Clenia brevis[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Microbiotheria, a species of Clenia.

Eomicrobiotherium matutinum[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Microbiotheria, a species of Eomicrobiotherium.

Epiklohnia[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Argyrolagoidea of uncertain phylogenetic placement. The type species is E. verticalis.

Galadi[4]

Valid

Oligo-Miocene

 Australia

A bandicoot.

Heishanobaatar[5]

Valid

Aptian/Albian

Shahai and Fuxin formations

 China

An eobaatarid multituberculate.

Hondonadia parca[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Bonapartheriiformes, a species of Hondonadia.

Hondonadia praecipitia[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Bonapartheriiformes, a species of Hondonadia.

Hondonadia pumila[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Bonapartheriiformes, a species of Hondonadia.

Hutegotherium[6]

Valid

Bathonian

Itat Formation

 Russia

A tegotheriid docodont. The type species is H. yaomingi.

Juchilestes[7]

Valid

Lower Aptian

Yixian Formation

 China

An amphidontid 'eutriconodont'. The type species is J. liaoningensis.

Kielanobaatar[5]

Valid

Aptian/Albian

Shahai and Fuxin formations

 China

An albionbaatarid multituberculate.

Klohnia major[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Argyrolagoidea of uncertain phylogenetic placement, a species of Klohnia.

Kukaodonta[8]

Valid

  • Mackness

Plio-Pleistocene

 Australia

A marsupial belonging to the subfamily Zygomaturinae; a new genus for "Euowenia" robusta De Vis (1891).

Kramadolops[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Late Eocene to early Oligocene

 Argentina

A member of Polydolopiformes. The type species is K. mayoi (Odreman Rivas, 1978); genus also includes K. abanicoi (Flynn and Wyss, 1999) and K. mckennai (Flynn and Wyss, 2004), as well as new species K. fissuratus and K. maximus.

Leptalestes toevsi[9]

Valid

  • Hunter
  • Heinrich
  • Weishampel

Late Cretaceous

St. Mary River Formation

 USA

A metatherian, a species of Leptalestes.

Litokoala dicktedfordi[10]

Valid

  • Pledge

Probably middle Miocene

Riversleigh World Heritage Area

 Australia

A marsupial belonging to the family Phascolarctidae (a relative of the koala).

Litokoala thurmerae[10]

Valid

  • Pledge

Late Oligocene

Etadunna Formation

 Australia

A marsupial belonging to the family Phascolarctidae (a relative of the koala).

Nanolestes mackennai[11]

Valid

  • Martin
  • Averianov
  • Pfretzschner

Oxfordian

Qigu Formation

 China

An amphitheriid, a species of Nanolestes.

Nidimys[9]

Valid

  • Hunter
  • Heinrich
  • Weishampel

Upper Cretaceous

St. Mary River Formation

 USA

A neoplagiaulacid multituberculate. The type species is N. occultus.

Pachybiotherium illuminatum[12]

Valid

  • Goin et al.

Miocene

Pinturas Formation

 Argentina

A marsupial belonging to the group Microbiotheria.

Paracimexomys propriscus[9]

Valid

  • Hunter
  • Heinrich
  • Weishampel

Late Cretaceous

St. Mary River Formation

 USA

A multituberculate, a species of Paracimexomys.

Paressonodon[13]

Valid

  • Wilson
  • Dechesne
  • Anderson

Late Cretaceous

Laramie Formation

 USA

A neoplagiaulacid multituberculate. The type species is P. nelsoni.

Parikimys[13]

Valid

  • Wilson
  • Dechesne
  • Anderson

Late Cretaceous

Laramie Formation

 USA

A cimolomyid multituberculate. The type species is P. carpenteri.

Paritatodon[14]

Valid

  • Martin
  • Averianov

Middle Jurassic (Callovian)

Balabansai Formation

 Kyrgyzstan

A basal docodont. A new genus for "Shuotherium" kermacki Sigogneau-Russell, 1988.

Periakros[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Polydolopimorphia belonging to the suborder Hatcheriformes and the family Glasbiidae. The type species is P. ambiguus.

Pilchenia antiqua[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Palaeothentidae, a species of Pilchenia.

Pilchenia intermedia[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Palaeothentidae, a species of Pilchenia.

Praedens[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Argyrolagoidea of uncertain phylogenetic placement. The type species is P. aberrans.

Rosendolops ebaios[3]

Valid

  • Goin
  • Abello
  • Chornogubsky

Early Oligocene

 Argentina

A member of Bonapartheriiformes, a species of Rosendolops.

Sineleutherus[11]

Valid

  • Martin
  • Averianov
  • Pfretzschner

Oxfordian

Qigu Formation

 China

A eleutherodontid allotherian.

Newly named eutherians

Name Status Authors Age Unit Location Notes Images

Acristatherium[15]

Valid

Aptian

Yixian Formation

 China

An early eutherian

Afrotarsius libycus[16]

Valid

  • Jaeger et al.

Eocene

 Libya

An early member of Anthropoidea.

Afrotragulus[17]

Valid

Early Miocene

Lothidok Formation

 Kenya

A tragulid ruminant.

Agathaeromys[18]

Valid

  • Zijlstra
  • Madern
  • van den Hoek Ostende

Probably Pleistocene

 Bonaire

An oryzomyine rodent. The type species is A. donovani; genus also incluses A. praeuniversitatis.

Ageitonomys[19]

Valid

  • Wang Ban-Yue

Early Oligocene

Ulantatal Formation

 China

A ctenodactyloid rodent. The species is A. neimongolensis.

Ailuravus mitchelli[20]

Valid

  • Hooker

Early Eocene

Blackheath Formation

 United Kingdom

Alloeumyarion[21]

Valid

  • Qiu Zhu-Ding

Early Miocene

Xiacaowan Formation

 China

A eumyarionine cricetid. The species is A. sihongensis.

Altanius magnus[22]

Valid

  • Maschenko & Takai

Early Eocene

Naran Bulak Formation

 Mongolia

A primate of uncertain phylogenetic placement, a species of Altanius.

Amphilagus wuttkei[23]

Junior synonym?

  • Mörs & Kalthoff

Late Oligocene

Enspel Formation

 Germany

A lagomorph, a species of Amphilagus. Considered to be a probable junior synonym of Amphilagus antiquus by Fostowicz-Frelik (2016).[24]

Apatemys prouti[20]

Valid

  • Hooker

Early Eocene

Blackheath Formation

 United Kingdom

A member of Apatemyidae.

Archaeonycteris relicta[25]

Valid

  • Harrison
  • Hooker

Eocene

Creechbarrow Limestone Formation

 United Kingdom

An early bat.

Arctocyonides jefferyi[20]

Valid

  • Hooker

Early Eocene

Blackheath Formation

 United Kingdom

A member of Arctocyonidae.

Australopithecus sediba[26]

Valid

Late Tertiary to Early Quaternary

Cradle of Humankind

 South Africa

One of the youngest known australopithecines to date.

Australopithecus sediba

Barrancatatus[27]

Valid

  • Carlini
  • Ciancio
  • Scillato-Yané

Early Oligocene

 Argentina
 Chile

An armadillo. The type species is "Meteutatus" rigidus Ameghino (1902); genus also includes "Meteutatus" tinguiririquensis Carlini et al. (2009), as well as new species Barrancatatus maddeni.

Basirepomys[28]

Valid

  • Kortha
  • De Blieux

Late Hemphillian (Late Miocene)

Sevier River Formation

 USA

Species:

  • B. pliocenicus (previously Peromyscus pliocenicus Wilson 1937) (type)
  • B. robertsi

Batodonoides walshi[29]

Valid

  • Kelly

Middle Eocene

Sespe Formation

 United States

A shrew-like mammal, a species of Batodonoides.

Blacktops[30]

Valid

  • Meehan
  • Martin

Early Oligocene

Brule Formation

 United States

A member of Leptictida. The type species is B. longinares; genus also includes B. latidens.

Bos buiaensis[31]

Valid

  • Martínez-Navarro et al.

Early Pleistocene

 Eritrea

A member of Bovidae, species of Bos.

Chasicotatus powelli[32]

Valid

  • Scillato–Yané, Krmpotic & Esteban

Late Miocene (Huayquerian)

 Argentina

An euphractine armadillo related to Eutatus, a species of Chasicotatus.

Chasicotatus spinozai[32]

Valid

  • Scillato–Yané, Krmpotic & Esteban

Late Miocene (Huayquerian)

 Argentina

An euphractine armadillo related to Eutatus, a species of Chasicotatus.

Chubutomys leucoreios[33]

Valid

  • Pérez
  • Vucetich
  • Kramarz

Miocene (Colhuehuapian)

 Argentina

A rodent belonging to the family Eocardiidae.

“Cricetodon” fandli[34]

Valid

  • Prieto, Böhme & Gross

Miocene

 Austria

A rodent.

Damalborea[35]

Valid

  • Gentry

Middle Pliocene

Hadar Formation

 Ethiopia

An alcelaphine bovid. Its type species is Damalborea elisabethae.

Darocasorex[36]

Valid

  • van Dam

Miocene

An anourosoricini soricidae.

Dawsonicyon[37]

Valid

  • Spaulding
  • Flynn
  • Stucky

Middle Eocene

Bridger Formation

 USA

A basal carnivoramorphan.

Deccanolestes narmadensis[38]

Valid

  • Prasad et al.

Late Cretaceous (Maastrichtian)

Deccan Intertrappean Beds

 India

Dendropithecus ugandensis[39]

Valid

  • Pickford et al.

Early Miocene

 Uganda

Desmanodon crocheti[40]

Valid

  • Prieto

Miocene

 Germany

A member of Talpidae.

Desmanodon fluegeli[41]

Valid

  • Prieto et al.

Miocene

 Austria

A member of Talpidae.

Diabolocornis[42]

Valid

  • Beatty

Early Miocene

 United States

An aletomerycine palaeomerycid. The type species is D. simonsi.

Diacodexis morrisi[20]

Valid

  • Hooker

Early Eocene

Blackheath Formation

 United Kingdom

Diacodexis parvus[43]

Valid

  • Kumar et al.

Eocene (Ypresian)

Cambay Formation

 India

Didolodus magnus[44]

Valid

  • Gelfo

Eocene

 Argentina

Dimaitherium[45]

Valid

  • Barrow
  • Seiffert
  • Simons

early Late Eocene (early Priabonian)

Birket Qarun Formation

 Egypt

A hyracoid. The type species is D. patnaiki.

Dilambdogale[46]

Valid

  • Seiffert

Earliest Priabonian

Birket Qarun Formation

 Egypt

The oldest known afrosoricid.

Diunatans[47]

Valid

  • Bosselaers
  • Post

Early Pliocene (Zanclean)

Kattendijk Formation

 Netherlands

A small rorqual

Dolosimus[48]

Valid

  • Dawson
  • Li
  • Qi

Eocene (Irdinmanhan)

 China

A rodent of uncertain phylogenetic placement. The type species is D. dolus.

Draconomys[49]

Valid

  • Vucetich et al.

Early Oligocene

 Argentina

An octodontoid rodent of uncertain phylogenetic placement. The type species is D. verai.

Edworthia[50]

Valid

  • Fox
  • Scott
  • Rankin

Middle Torrejonian

Paskapoo Formation

 Canada

A primitive paromomyid plesiadapiform.

Erlianomys[51]

Valid

  • Li Qian
  • Meng Jin

Early Eocene

Arshanto Formation

 China

A primitive myodont rodent. The species is E. combinatus.

Eudaemonema webbi[52]

Valid

  • Scott

Late Paleocene

 Canada

A member of Mixodectidae.

Frontanyamys[53]

Valid

  • Quer
  • Agustí

Eocene

 Spain

A rodent belonging to the family Remyidae. The type species is F. russelli.

Gaudeamus lavocati[54]

Valid

  • Coster et al.

Early Oligocene

 Libya

A rodent belonging to the group Hystricognathi.

Geringia copiosus[55]

Valid

  • Korth

Late Oligocene

 United States

A cricetid rodent, a species of Geringia.

Gracilocyon[56]

Valid

  • Smith
  • Smith

earliest Eocene

 Belgium  USA

A miacid carnivoran.

Guiomys[57]

Valid

  • Pérez

Middle Miocene

 Argentina

A primitive cavioid rodent.

Henricofilholia vucetichia[58]

Valid

  • Ribeiro
  • López
  • Bond

Oligocene

Sarmiento Formation

 Argentina

A leontiniid notoungulate.

Homo gautengensis[59]

Valid

  • Curnoe

Pleistocene

Cradle of Humankind

 South Africa

Identification of H. gautengensis was based on partial skulls, several jaws, teeth and other bones found at various times at the Caves. It emerged over 2 million years ago and died out approximately 600,000 years ago, and is believed to have arisen earlier than Homo habilis.

Hypolagus balearicus[60]

Valid

  • Quintana et al.

Early Pliocene

 Spain

A member of Leporidae belonging to the subfamily Archaeolaginae.

Iriripithecus[39]

Valid

  • Pickford et al.

Early Miocene

 Uganda

An early catarrhine of uncertain affinity. The type species is I. alekileki.

Joumocetus[61]

Valid

  • Kimura
  • Hasegawa

earliest Late Miocene

Haraichi Formation

 Japan

A cetotheriid whale. The type species is J. shimizui.

Kabirmys[62]

Valid

earliest late Eocene

Birket Qarun Formation

 Egypt

An anomaluroid rodent.

Karamojapithecus[39]

Valid

  • Pickford et al.

Early Miocene

 Uganda

An early catarrhine of uncertain affinity. The type species is K. akisimia.

Karanisia arenula[16]

Valid

  • Jaeger et al.

Eocene

 Libya

Komba walkeri[63]

Valid

  • Harrison

Early Miocene

 Kenya

A galago.

Kutchisiren[64]

Valid

Miocene

Khari Nadi Formation

 India

A new name for Kotadasiren gracilis Das & Basu, 1994 (nomen nudum). The type species is Kutchisiren cylindrica.

Leidymys juxtaparvulus[55]

Valid

  • Korth

Late Oligocene

 United States

A cricetid rodent, a species of Leidymys.

Lestobradys[65]

Valid

Late Miocene (Huayquerian)

Camacho Formation

 Uruguay

A mylodontid. The type species is L. sprechmanni.

Livyatan[66]

Valid

Miocene

Pisco Formation

 Peru

A physeteroid whale. The type species is L. melvillei.

Livyatan

Lokotunjailurus fanonei[67]

Valid

  • De Bonis et al.

Late Miocene

 Chad

A saber-toothed cat.

Luantus minor[33]

Valid

  • Pérez
  • Vucetich
  • Kramarz

Miocene (Colhuehuapian)

 Argentina

A rodent belonging to the family Eocardiidae.

Lycaon sekowei[68]

Valid

  • Hartstone-Rose et al.

Pliocene to Pleistocene

 South Africa

A relative of the African wild dog.

Mazateronodon[69]

Valid

  • Marigó, Minwer-Barakat & Moyà-Solà

Middle Eocene

 Spain

A notharctid adapiform. The type species is Mazateronodon endemicus.

Mazzonicebus[70]

Valid

  • Kay

Early Miocene

Sarmiento Formation

 Argentina

A New World monkey related to Homunculus patagonicus and members of the genus Soriacebus. The type species is M. almendrae.

Mazzoniphractus[27]

Valid

  • Carlini
  • Ciancio
  • Scillato-Yané

Late Eocene

Sarmiento Formation

 Argentina

An armadillo. The type species is M. ingens.

Mendozahippus[71]

Valid

  • Cerdeño
  • Vera

Late Oligocene

Agua de la Piedra Formation

 Argentina

A notohippid notoungulate.

Metaliomys[28]

Valid

  • Kortha
  • De Blieux

Late Hemphillian (Late Miocene)

Sevier River Formation

 USA

Species: Metaliomys sevierensis

Miacis rundlei[20]

Valid

  • Hooker

Early Eocene

Blackheath Formation

 United Kingdom

Originally described as a species of Miacis,[20] subsequently transferred to the genus Gracilocyon.[72]

Miocyon vallisrubrae[73]

Valid

  • Friscia
  • Rassmussen

Middle Eocene

Uinta Formation

 USA

Mormopterus barrancae[74]

Valid

  • Czaplewski

Colhuehuapian

 Argentina

A free-tailed bat, a species of Mormopterus.

Myotragus palomboi[75]

Valid

  • Bover
  • Quintana
  • Alcover

Early Pliocene

 Spain

A member of Caprinae.

Neomatronella gassoni[20]

Valid

  • Hooker

Early Eocene

Blackheath Formation

 United Kingdom

A member of Erinaceomorpha belonging to the family Amphilemuridae.

Nosmips[76]

Valid

  • Seiffert et al.

Eocene (Priabonian)

Birket Qarun Formation

 Egypt

A primate of uncertain phylogenetic placement. The type species is N. aenigmaticus.

Nyctereutes lockwoodi[77]

Valid

  • Geraads et al.

Middle Pliocene

 Ethiopia

A species of Nyctereutes (a relative of the raccoon dog).

Ochotona kormosi[78]

Valid

  • Fostowicz-Frelik
  • Frelik
  • Gasparik

Pleistocene

 Hungary

A pika.

Omoiosicista[79]

Valid

  • Kimura

Early Miocene

 China

A dipodid rodent.

Orelladjidaumo amplus[80]

Valid

  • Korth

Late Oligocene

 United States

An eomyid rodent, a species of Orelladjidaumo.

Paciculus dakotensis[55]

Valid

  • Korth

Late Oligocene

 United States

A cricetid rodent, a species of Paciculus.

Patagonhippus[81]

Valid

  • López
  • Ribeiro
  • Bond

Oligocene

 Argentina

A notohippid notoungulate. The type species is P. canterensis; genus also includes P. dukei.

Phiomys hammudai[82]

Valid

  • Jaeger et al.

Eocene

 Libya

A rodent belonging to the family Phiomyidae.

Plesiobalaenoptera[83]

Valid

  • Bisconti

Late Miocene (Tortonian)

Sediments of the Stirone River

 Italy

A parabalaenopterine balaenopterid. The type species is P. quarantellii.

Platalearostrum[84]

Valid

  • Post
  • Kompanje

early or middle Pleistocene

 the Netherlands

A prehistoric pilot whale (Family Delphinidae). The type species is P. hoekmani

Pliolophus barnesi[20]

Valid

  • Hooker

Early Eocene

Blackheath Formation

 United Kingdom

Presbytherium[85]

Valid

  • Scott

Paleocene

Paskapoo Formation

 Canada

A cyriacotheriid pantodont. Species:

  • P. rhodorugatus
  • P. taurus

Prolagus osmolskae[86]

Valid

  • Fostowicz-Frelik

Pliocene

 Poland

A member of Lagomorpha, a species of Prolagus.

Protophiomys durattalahensis[82]

Valid

  • Jaeger et al.

Eocene

 Libya

A rodent belonging to the family Phiomyidae.

Pseudoloris pyrenaicus[87]

Valid

  • Minwer-Barakat
  • Marigó
  • Moyà-Solà

Eocene

 Spain

A primate belonging to the family Omomyidae.

Qatranilestes[46]

Valid

  • Seiffert

Rupelian

Jebel Qatrani Formation

 Egypt

The youngest known afrosoricid.

Rusingaceros[88]

Valid

  • Geraads

Miocene

 Kenya

A rhinoceros, a new genus for "Dicerorhinus" leakeyi (Hooijer, 1966).

Saadanius[89]

Valid

Oligocene

Shumaysi Formation

 Saudi Arabia

A catarrhine primate. The species was named S. hijazensis.

Saadanius

Sadypus minutus[27]

Valid

  • Carlini
  • Ciancio
  • Scillato-Yané

Early Oligocene

Sarmiento Formation

 Argentina

An armadillo, a species of Sadypus.

Scarrittia barranquensis[58]

Valid

  • Ribeiro
  • López
  • Bond

Oligocene

Sarmiento Formation

 Argentina

A leontiniid notoungulate.

Shazurus[90]

Valid

  • Sallam
  • Seiffert
  • Simons

Late Eocene

 Egypt

An anomalure. The type species is S. minutus.

Simiolus andrewsi[91]

Valid

  • Harrison

Middle Miocene

 Kenya

A dendropithecid.

Sinodonomys[79]

Valid

  • Kimura

Early Miocene

 China

A dipodid rodent.

Sparnacomys georgei[20]

Valid

  • Hooker

Early Eocene

Blackheath Formation

 United Kingdom

A rodent belonging to the family Paramyidae and the subfamily Microparamyinae.

Talahphiomys[82]

Valid

  • Jaeger et al.

Eocene to early Oligocene

 Egypt
 Libya

A rodent belonging to the family Phiomyidae. The type species is "Phiomys" lavocati Wood (1968); genus also includes new species T. libycus.

Talahpithecus[16]

Valid

  • Jaeger et al.

Eocene

 Libya

A member of Oligopithecidae. The type species is T. parvus.

Tapirus lundeliusi[92]

Valid

  • Hulbert

Early Pleistocene

 United States

A tapir.

Turkanapithecus rusingensis[39]

Valid

  • Pickford et al.

Early Miocene

 Uganda

Vallehermosomys[49]

Valid

  • Vucetich et al.

Early Oligocene

 Argentina

An octodontoid rodent of uncertain phylogenetic placement. The type species is V. mazzonii; genus might also include Vallehermosomys? merlinae.

Viverravus lawsoni[20]

Valid

  • Hooker

Early Eocene

Blackheath Formation

 United Kingdom

A member of Viverravidae.

Notes

  1. ^ Travouillon, Gurovich, Beck, Muirhead.
  2. ^ a b Kusuhashi, Hu, Wang, Setoguchi, Marsuoka.
  3. ^ Averianov, Lopatin, Krasnolutskii, Ivantsov.
  4. ^ Gao, Wilson, Luo, Maga, Meng, Wang.
  5. ^ Hu, Meng, Li, Wang.
  6. ^ Sánchez, Quiralte, Morales, Pickford.
  7. ^ Berger, de Ruiter, Churchill, Schmid, Carlson, Dirks, Kibii.
  8. ^ Sallam, Seiffert, Simons, Brindley.
  9. ^ Bajpai, Domning, Das, Velez-Juarbe, Mishra.
  10. ^ Rinderknechta, Bostelmann, Pereab, Lecuona.
  11. ^ Lambert, Bianucci, Post, de Muizon, Salas-Gismondi, Urbina, Reumer.
  12. ^ Zalmout, Sanders, MacLatchy, Gunnell, Al-Mufarreh, Ali, Nasser, Al-Masari, Al-Sobhi, Nadhra, Matari, Wilson, Gingerich.

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