Bowhead whale: Difference between revisions

Bowhead whale
	Size compared to an average human
Conservation status
	; Least Concern (IUCN 3.1)
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Cetacea
Suborder:	Mysticeti
Family:	Balaenidae
Genus:	Balaena; Linnaeus, 1758
Species:	B. mysticetus
Binomial name
	Balaena mysticetus; Linnaeus, 1758
	Bowhead whale range

Browse history interactively

← Previous edit Next edit →

Content deleted Content added

VisualWikitext

Inline

Revision as of 13:18, 29 August 2016

The bowhead whale (Balaena mysticetus) is a species of the family Balaenidae, in suborder Mysticeti, and genus Balaena, once thought to include the right whale.

A stocky dark-colored whale without a dorsal fin, it can grow 14 to 18 m (46 to 59 ft) in length. This thick-bodied species can weigh from 75 to 100 tonnes (74 to 98 long tons; 83 to 110 short tons).^[3] They live entirely in fertile Arctic and sub-Arctic waters, unlike other whales that migrate to low latitude waters to feed or reproduce. The bowhead was also known as the Greenland right whale or Arctic whale. American whalemen called them the steeple-top, polar whale,^[4] or Russia or Russian whale. The bowhead has the largest mouth of any animal.^[5]

The bowhead was an early whaling target. The population was severely reduced before a 1966 moratorium was passed to protect the species. Through conservation efforts, the bowhead population has since recovered and is now rated "Least Concern" on the IUCN Red List of Threatened Species.^[2]

Taxonomy

Carl Linnaeus first described this whale in the 10th edition of his Systema Naturae (1758).^[6] Seemingly identical to its cousins in the North Atlantic, North Pacific and Southern Oceans, they were all thought to be a single species, collectively known as the "right whale", and given the binomial name Balaena mysticetus.

Today, the bowhead whale occupies a monotypic genus, separate from the right whales, as was proposed by the work of John Edward Gray in 1821.^[7] For the next 180 years, the family Balaenidae was the subject of great taxonometric debate. Authorities have repeatedly recategorized the three populations of right whale plus the bowhead whale, as one, two, three or four species, either in a single genus or in two separate genera. Eventually, it was recognized that bowheads and right whales were in fact different, but there was still no strong consensus as to whether they shared a single genus or two. As recently as 1998, Dale Rice, in his comprehensive and otherwise authoritative classification, Marine mammals of the world: systematics and distribution, listed just two species: B. glacialis (the right whales) and B. mysticetus (the bowheads).^[8]

Studies in the 2000s finally provided clear evidence that the three living right whale species do comprise a phylogenetic lineage, distinct from the bowhead, and that the bowhead and the right whales are rightly classified into two separate genera.^[9] The right whales were thus confirmed to be in a separate genus, Eubalaena. The relationship is shown in the cladogram below:

Family Balaenidae

Eubalaena (right whales)

E. glacialis North Atlantic right whale

	E. japonica North Pacific right whale

	E. australis Southern right whale

Balaena (bowhead whales)

B. mysticetus bowhead whale

The bowhead whale, genus Balaena, in the family Balaenidae (extant taxa only)^[10]

Balaena prisca, one of the five Balaena fossils from the late Miocene (~10 Mya) to early Pleistocene (~1.5 Mya), may be the same as the modern bowhead whale. The earlier fossil record shows no related cetacean after Morenocetus, found in a South American deposit dating back 23 million years.

An unknown species of right whale, the so-called "Swedenborg whale" which was proposed by Emanuel Swedenborg in the 18th century, was once thought to be a North Atlantic right whale by scientific consensus. However based on later DNA analysis those fossil bones claimed to be from "Swedenborg whales" were confirmed to be from bowhead whales.^[11]

Description

Drawing of long backbone, 13 ribs (two vestigial) large, curved upper and lower jawbones that occupy 1/3 of the body, 4 multijointed "fingers" inside pectoral fin and connecting bone, enclosed in body outline — Skeleton of a bowhead whale

The bowhead whale has a large, robust, dark-colored body and a white chin/lower jaw. The whale has a massive triangular skull, which the whale uses to break through the Arctic ice to breathe. Inuit hunters have reported bowheads surfacing through 60 cm (24 in) of ice.^[12] The bowhead also has a strongly bowed lower jaw and a narrow upper jaw. Its baleen is the longest of that of any whale, at 3 m (9.8 ft), and is used to strain tiny prey from the water. The bowhead whale has paired blowholes, at the highest point of the head, which can spout a blow 20 ft (6.1 m) high. The whale's blubber is the thickest of that of any animal, with a maximum of 43–50 cm (17–20 in).^[13] Unlike most cetaceans, the bowhead does not have a dorsal fin.^[14]

Bowhead whales are comparable in size to the three species of right whales. According to whaling captain William Scoresby Jr., the longest bowhead he measured was 17.7 m (58 ft) long, while the longest measurement he had ever heard of was of a 20.4 m (67 ft) whale caught at Godhavn, Greenland, in early 1813. He also spoke of one, caught near Spitsbergen around 1800, that was allegedly nearly 21.3 m (70 ft) long.^[15] In 1850, an American vessel claimed to have caught a 24.54 m (80.5 ft) individual in the Western Arctic.^[16] It is questionable whether these lengths were actually measured. The longest reliably measured lengths of the sexes were 16.2 m (53 ft) in a male and 18 m (59 ft) in a female, both landed by natives in Alaska.^[17] On average, female bowheads are larger than males.

Analysis of hundreds of DNA samples from living whales and from baleen used in vessels, toys, and housing material has shown that Arctic bowhead whales have lost a significant portion of their genetic diversity in the past 500 years. Bowheads originally crossed ice-covered inlets and straits to exchange genes between Atlantic and Pacific populations. This conclusion was derived from analyzing maternal lineage using mitochondrial DNA. Whaling and climatic cooling during the Little Ice Age, from the 16th century to the 19th, is supposed to have reduced the whales’ summer habitats, which explains the loss of genetic diversity.^[18]

A 2013 discovery has elucidated the function of the bowhead's large palatal retial organ. The bulbous ridge of highly vascularized tissue, the corpus cavernosum maxillaris, extends along the center of the hard plate, forming two large lobes at the rostral palate. The tissue is histologically similar to that of the corpus cavernosum of the mammalian penis. It is hypothesized that this organ provides a mechanism of cooling for the whale (which is normally protected from the cold Arctic waters by 40 cm (16 in) or more of fat). During physical exertion, the whale must cool itself to prevent hyperthermia (and ultimately brain damage). It is now believed that this organ becomes engorged with blood, causing the whale to open its mouth to allow cold seawater to flow over the organ, thus cooling the blood.^[19]

Behavior

Swimming

Bowhead whales are not social animals, typically traveling alone or in small pods of up to 6. They are able to dive and remain submerged underwater for up to an hour. However, the time spent underwater in a single dive is usually limited to 9–18 minutes.^[12] Bowheads are not thought to be deep divers but they can reach a depth of up to 500 ft (150 m). These whales are slow swimmers, normally traveling at about 2–5 km/h (1.2–3.1 mph).^[20] When fleeing from danger, they can travel at a speed of 10 km/h (6.2 mph). During periods of feeding, the average swim speed is reduced to 1.1–2.5 m/s (3.6–8.2 ft/s).^[21]

Feeding

The head of the bowhead whale comprises a third of its body length, creating an enormous feeding apparatus.^[21] Bowhead whales are filter feeders, feeding by swimming forward with mouth wide open.^[12] The whale has hundreds of overlapping baleen plates consisting of keratin hanging from each side of the upper jaw. The mouth has a large upturning lip on the lower jaw that helps to reinforce and hold the baleen plates within the mouth. This also prevents buckling or breakage of the plates from the pressure of the water passing through them as the whale advances. To feed, water is filtered through the fine hairs of keratin of the baleen plates, trapping the prey inside near the tongue where it is then swallowed.^[22] The diet consists of mostly zooplankton which includes copepods, amphipods, and many other crustaceans.^[21] Approximately 2 short tons (1.8 long tons; 1.8 t) of food is consumed each day.^[22] While foraging, bowheads are solitary or occur in groups of two to ten or more.^[13]

Vocalization

Bowhead whales are highly vocal^[23] and use low frequency (<1000 Hz) sounds to communicate while traveling, feeding, and socializing. Intense calls for communication and navigation are produced especially during migration season. During breeding season, bowheads make long, complex, variable songs for mating calls.^[20]

Reproduction

Sexual activity occurs between pairs and in boisterous groups of several males and one or two females. Breeding season is observed from March through August; conception is believed to occur primarily in March when song activity is at its highest.^[20] Reproduction can begin when a whale is 10 to 15 years old. The gestation period is 13–14 months with females producing a calf once every three to four years.^[17] Lactation typically lasts about a year. To survive in the cold water immediately after birth, calves are born with a thick layer of blubber. Within 30 minutes of birth, bowhead calves are able to swim on their own. A newborn calf is typically 4–4.5 m (13–15 ft) long, weighs approximately 1,000 kg (2,200 lb), and grows to 8.2 m (27 ft) within the first year.^[17]

Health

Lifespan

Bowhead whales are known to be among the longest-living mammals, living for over 200 years.^[24] In May 2007, a 15 m (49 ft) specimen caught off the Alaskan coast was discovered with the head of an explosive harpoon embedded deep under its neck blubber. The 3.5 in (89 mm) arrow-shaped projectile was manufactured in New Bedford, Massachusetts, a major whaling center, around 1890, suggesting the animal may have survived a similar hunt more than a century ago.^[25]^[26]^[27] This whale was estimated to be 211 years old.^[28] Other bowhead whales found on the whaling expedition were estimated to be between 135 and 172 years old. This discovery showed the longevity of the bowhead whale is much greater than originally thought.

Genetic causes

It was previously believed the more cells present in an organism, the greater the chances of mutations that cause age related diseases and cancer.^[29] Although the bowhead whale has thousands of times more cells than other mammals, the whale has a much higher resistance to cancer and aging. In 2015, scientists from the US and UK were able to successfully map the whale's genome.^[30] Through comparative analysis, two alleles that could be responsible for the whale's longevity were identified. These two specific gene mutations linked to the bowhead whale's ability to live longer are the ERCC1 gene and the proliferating cell nuclear antigen (PCNA) gene. ERCC1 is linked to DNA repair as well as increased cancer resistance. PCNA is also important in DNA repair. These mutations enable bowhead whales to better repair DNA damage, allowing for greater resistance to cancer.^[29] The whale's genome may also reveal physiological adaptations such as having low metabolic rates compared to other mammals.^[31] Changes in the gene UCP1, a gene involved in thermoregulation, can explain differences in the metabolic rates in cells.

Ecology

Range and habitat

The bowhead whale is the only baleen whale to spend its entire life in the Arctic and sub-Arctic waters.^[32] The Alaskan population spends the winter months in the southwestern Bering Sea. The group migrates northward in the spring, following openings in the ice, into the Chukchi and Beaufort seas.^[33] It has been confirmed the whale's range varies depending on climate changes and on the forming/melting of ice.^[34]

Population

The bowhead population around Alaska has increased since commercial whaling ceased. Alaska Natives continue to kill small numbers in subsistence hunts each year. This level of killing (25–40 animals annually) is not expected to affect the population's recovery. The population off Alaska's coast (the Bering-Chukchi-Beaufort stock) appears to be recovering and was at about 10,500 animals as of 2001. The status of other populations is less well known. There were about 1,200 off West Greenland in 2006, while the Svalbard population may only number in the tens. However, the numbers have been increasing in recent years.^[35]

In March 2008, Canada's Department of Fisheries and Oceans stated the previous estimates in the eastern Arctic had under-counted, with a new estimate of 14,400 animals (range 4,800–43,000).^[36] These larger numbers correspond to prewhaling estimates, indicating the population has fully recovered. However, if climate change substantially shrinks sea ice, these whales could be threatened by increased shipping traffic.^[37]

Hudson Bay to Foxe Basin

File:The Canadian field-naturalist (1997) (20330377360).jpg

Sighting locations by researchers and hunters

Hudson Bay – Foxe Basin population is distinct from the Baffin Bay – Davis Strait group.^[38] Original population size of this local group is unclear, but possibly around 500 to 600 whales annually summered in the northwestern part of the bay in 1860s.^[39] Likely, the number of whales actually inhabit within Hudson Bay is much smaller than the total population size of this group,^[40] and despite current population size is rather unclear, reports from local indigenous people indicate this population is at least increasing over decades.^[41] Larger portions of usages of the bay is considered to be summering while wintering is on smaller scale where some animals winter in Hudson Strait most notably north of Igloolik Island and northeastern Hudson Bay. Distribution patterns of whales in this regions are largely affected by presences of killer whales and bowheads can disappear from normal ranges due to recent changes in killer whales' occurrences within the bay possibly because of changes in movements of ice floes by changing climate.^[41] Whaling grounds in 19th century covered from Marble Island to Roes Welcome Sound and to Lyon Inlet and Fisher Strait, and whales still migrate through most of these areas.

Mostly, distributions within Hudson Bay is restricted in northwestern part^[38] along with Repulse Bay, Frozen Strait, northern Foxe Basin, and north of Igloolik in summer,^[41] and satellite tracking^[42] indicates that some portions of the group within the bay do not venture further south than areas south of Coasts and Mansel Islands.^{[citation needed]} Cow – calf pairs and juveniles up to 13.5 m (44 ft) in length consist of majority of summering aggregation in northern Foxe Basin while matured males and non-calving females may utilize northwestern part of Hudson Bay.^[41] Fewer whales also migrate to west coast of Hudson Bay, Mansel and Ottawa Islands.^[41] Bowhead ranges within Hudson Bay are usually considered not to cover southern parts,^[40]^[43] but at least some whales migrate into further south such as at Sanikiluaq^{[citation needed]} and Churchill river mouth.^[44]^[45]

Congregation within Foxe Basin occurs in a well-defined area at 3,700 km (2,300 mi) north of Igloolik Island to Fury and Hecla Strait and Jens Munk Island and Gifford Fiord, and into Gulf of Boothia and Prince Regent Inlet. Northward migrating along western Foxe Basin to eastern side of the basin also occurs in spring seasons.^[41]

Sea of Okhotsk

Not much is known about the endangered Sea of Okhotsk population. To learn more about the population, these mammals have been regularly observed near the Shantar Islands, very close to the shore, such as at Ongachan Bay.^[47]^[48] Several companies provide whale watching services which are mostly land-based. According to Russian scientists, this total population likely does not exceed 400 animals.^[46] Scientific research on this population was seldom done before 2009, when researchers studying belugas noticed concentrations of bowheads in the study area. Thus, bowheads in the Sea of Okhotsk were once called "forgotten whales" by researchers. With support from WWF, Russian scientists and nature conservationists cooperated to create a cetacean sanctuary in the Magadan region. This region covers vast areas of the northwestern Sea of Okhotsk, including the Shantar regions.^[49]

Possibly, vagrants from this population occasionally reach into Asian nations such as off Japan or Korean Peninsula (although this record might or might not be of a right whale^[50]). First documented report of the species in Japanese waters was of a strayed infant (7 m [23 ft]) caught in Osaka Bay on 23 June 1969,^[51] and the first living sighting was of a 10 m (33 ft) juvenile around Shiretoko Peninsula (the southernmost of ice floe range in the world) on 21 to 23 June 2015.^[52] Fossils have been excavated on Hokkaido,^[53] but it is unclear whether or not northern coasts of Japan once had been included in seasonal or occasional migration ranges.

Genetic studies suggest Okhotsk population share common ancestry with whales in Bering-Chukchi-Beaufort Seas, and repeated mixings had occurred between whales in two seas.^[54]

Russian Arctic

The most endangered and historically the largest of all bowhead populations is the Svalbard/Spitsbergen population.^[55] Occurring normally in Fram Strait,^[56] Barents Sea and Severnaya Zemlya along Kara Sea^[35] to Laptev Sea and East Siberian Sea regions, these whales were seen in entire coastal regions in European and Russian Arctic, even reaching to Icelandic and Scandinavian coasts and Jan Mayen in Greenland Sea, and west of Cape Farewell and western Greenland coasts.^[57] Also, bowheads in this stock were possibly once abundant in areas adjacent to the White Sea region, where few or no animals currently migrate, such as the Kola and Kanin Peninsula. Today, the number of sightings in elsewhere are very small,^[58] but with increasing regularities^[59] with whales having strong regional connections.^[60] Whales have also started approaching townships and inhabited areas such as around Longyearbyen.^[61] The waters around the marine mammal sanctuary^[62] of Franz Josef Land is possibly functioning as the most important habitat for this population.^[63]^[64]

Current status of population structure of this stock is unclear; whether they are remnant of the historic Svalbard group, re-colonized individuals from other stocks, or if a mixing of these two or more stocks had taken place. In 2015, discoveries of the refuge along eastern Greenland where whaling ships could not reach due to ice floes^[65] and largest numbers of whales (80–100 individuals) ever sighted between Spitsbergen and Greenland^[66] indicate that more whales than previously considered survived whaling periods, and flows from the other populations are possible.

Possible moulting area on Baffin Island

During expeditions by a tour operator 'Arctic Kingdom', a large group of bowheads seemingly involved in courtship activities were discovered in very shallow bays in south of Qikiqtarjuaq in 2012.^[67] Floating skins and rubbing behaviors at sea bottom indicated possible moulting had taken place. Moulting behaviors had never or had seldomly been documented for this species before. This area is an important habitat for whales that were observed to be relatively active and to interact with humans positively, or to rest on sea floors. These whales belong to Davis Strait stock.

Isabella Bay in Niginganiq National Wildlife Area is the first wildlife sanctuary in the world to be designed specially for bowhead whales. However, moultings have not been recorded in this area due to environmental factors.^[68]

Predation

The principal predators of bowheads are humans.^[69] Killer whales are also known predators.^[70] Bowheads seek the safety of the ice and shallow waters when threatened by killer whales.^[20]

Whaling

The bowhead whale has been hunted for blubber, meat, oil, bones, and baleen. Like right whales, it swims slowly, and floats after death, making it ideal for whaling.^[71] Before commercial whaling, they were estimated to number 50,000.^[72]

Commercial bowhead whaling began in the 16th century, when the Basques killed them as they migrated south through the Strait of Belle Isle in the fall and early winter. In 1611, the first whaling expedition sailed to Spitsbergen. By mid-century, the population(s) there had practically been wiped out, forcing whalers to voyage into the "West Ice"—the pack ice off Greenland's east coast. By 1719, they had reached the Davis Strait, and by the first quarter of the 19th century, Baffin Bay.^[73]

In the North Pacific, the first bowheads were taken off the eastern coast of Kamchatka by the Danish whaleship Neptun, Captain Thomas Sodring, in 1845.^[16] In 1847, the first bowheads were caught in the Sea of Okhotsk, and the following year, Captain Thomas Welcome Roys, in the bark Superior, of Sag Harbor, caught the first bowheads in the Bering Strait region. By 1849, 50 ships were hunting bowheads in each area. By 1852, 220 ships were cruising around the Bering Strait region, which killed over 2,600 whales. Between 1854 and 1857, the fleet shifted to the Sea of Okhotsk, where 100–160 ships cruised annually. During 1858–1860, the ships shifted back to the Bering Strait region, where the majority of the fleet would cruise during the summer up until the early 20th century.^[74] An estimated 18,600 bowheads were killed in the Bering Strait region between 1848 and 1914, with 60% of the total being reached within the first two decades. An estimated 18,000 bowheads were killed in the Sea of Okhotsk during 1847–1867, 80% in the first decade.^[75]

Bowheads were first taken along the pack ice in the northeastern Sea of Okhotsk, then in Tausk Bay and Northeast Gulf (Shelikhov Gulf). Soon, ships expanded to the west, catching them around Iony Island and then around the Shantar Islands. In the Western Arctic, they mainly caught them in the Anadyr Gulf, the Bering Strait, and around St. Lawrence Island. They later spread to the western Beaufort Sea (1854) and the Mackenzie River delta (1889).^[74]

Commercial whaling, the principal cause of the population decline, is over. Bowhead whales are now hunted on a subsistence level by native peoples of North America.^[76]

Subsistence whaling in Alaska

Some of the Native peoples of North America, specifically Alaska, continue to hunt the bowhead whale on a subsistence level. Some of those peoples include:

Eskimo
- Iñupiat
- Yupik

For the Native peoples of Alaska subsistence Hunting occurs all year, and is normally managed under the same regulations as general season, drawing, and registration hunts, and a hunting license and harvest tag is usually required. For marine mammals, in specific, subsistence hunting is managed by the National Marine Fisheries Service.^[77]

Conservation

The bowhead is listed in Appendix I by CITES (that is, "threatened with extinction"). It is listed by the National Marine Fisheries Service as "endangered" under the auspices of the United States' Endangered Species Act. The IUCN Red List data are as follows:^[71]

Svalbard population – Critically endangered
Sea of Okhotsk subpopulation – Endangered
Baffin Bay-Davis Strait stock – Endangered
Hudson Bay-Foxe Basin stock – Vulnerable (Estimated to be 1,026 individuals in 2005 by DFO)^[78]
Bering-Chukchi-Beaufort stock – Lower risk – conservation dependent

The bowhead whale is listed in Appendix I^[79] of the Convention on the Conservation of Migratory Species of Wild Animals (CMS), as this species has been categorized as being in danger of extinction throughout all or a significant proportion of their range. CMS Parties strive towards strictly protecting these animals, conserving or restoring the places where they live, mitigating obstacles to migration, and controlling other factors that might endanger them.^[71]

Gallery

Media related to Balaena mysticetus at Wikimedia Commons

Whales swimming in Lindgolm strait (Russian: пролив Линдгольма) of Shantar Islands, northwestern Sea of Okhotsk^[80]
Cavorting whales in northwestern part of Sea of Okhotsk^[46]
Blowholes
Resting in Foxe Basin
Fluke up before diving
Whale showing one of pectoral fins
Tip of whitish chin visible
Map of the bowhead whale ranges centered over the North Pole

References

^ Mead, J. G.; Brownell, R. L. Jr. (2005). "Order Cetacea". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 723–743. ISBN 978-0-8018-8221-0. OCLC 62265494.
^ ^a ^b Template:IUCN2012.2
^ Rugh, David J.; Shelden, Kim E. W. (2008). "Bowhead Whale". In Perrin, William F.; Würsig, Bernd; Thewissen, J. G. M. (eds.). Encyclopedia of Marine Mammals (Second ed.). Academic Press. p. 131. ISBN 978-0-12-373553-9.
^ Scammon, Charles M. (1874) The Marine Mammals of the North-Western Coast of North America, New York: G.P. Putnam's Sons, ISBN 1-59714-061-9.
^ Guinness World Records (14 November 2007). "Whale of a time!". Archived from the original on 17 November 2007. Retrieved 4 June 2009. {{cite web}}: Unknown parameter |deadurl= ignored (|url-status= suggested) (help)
^ Linnaeus, C (1758). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis [System of nature through the three kingdoms of nature, according to the classes, orders, genera, species, with the characters, the differences, synonyms, places.] (in Latin). Vol. I (tenth, reformed ed.). Holmiae. (Laurentii Salvii). p. 824.
^ Template:IUCN2012.2 "The taxonomy is not in doubt.... Concerning common names, the species was once commonly known in the North Atlantic and adjacent Arctic as the Greenland Right Whale. However, the common name Bowhead Whale is now generally used for the species."
^ Rice, Dale W. (1998). Marine mammals of the world: systematics and distribution. Society of Marine Mammalogy Special Publication No. 4. ISBN 1-891276-03-4.
^ Kenney, Robert D. (2008). "Right Whales (Eubalaena glacialis, E. japonica, and E. australis)". In Perrin, William F.; Würsig, Bernd; Thewissen, J. G. M. (eds.). Encyclopedia of Marine Mammals. Academic Press. pp. 962–969. ISBN 978-0-12-373553-9.
^ "World-wide genetic differentiation of Eubalaena: Questioning the number of right whale species" (PDF). Molecular Ecology. 9 (11): 1793–802. 2000. doi:10.1046/j.1365-294x.2000.01066.x. PMID 11091315. {{cite journal}}: Cite uses deprecated parameter |authors= (help); Unknown parameter |lastauthoramp= ignored (|name-list-style= suggested) (help)
^ "Whale bones found in highway were not from mystery whale". ScienceNordic.com. 7 February 2013.
^ ^a ^b ^c Würsig, B. and C. Clark (1993). "Behavior". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.
^ ^a ^b Lowry, L. F. (1993). "Foods and Feeding Ecology". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.
^ Haldiman, J. T. and R. J. Tarpley (1993). "Anatomy & Physiology". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.
^ Scoresby, William (1820). An Account of the Arctic Regions with a History and a Description of the Northern Whale-Fishery. Edinburgh.
^ ^a ^b Bockstoce, J. R., and J. J. Burns (1993). "Commercial Whaling in the North Pacific Sector". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.
^ ^a ^b ^c Koski, William R., Rolph A. Davis, Gary W. Miller, and David E. Withrow (1993). "Reproduction". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy. p. 245.
^ Eilperin, Juliet (18 October 2012). "Bowhead whales lost genetic diversity, study shows". The Washington Post.
^ Ford Jr, T. J.; Werth, A. J.; George, J. C. (2013). "An intraoral thermoregulatory organ in the bowhead whale (Balaena mysticetus), the corpus cavernosum maxillaris". Anatomical Record. 296 (4): 701–708. doi:10.1002/ar.22681. PMID 23450839.
^ ^a ^b ^c ^d Finley, K.J. (2001). "Natural History and Conservation of the Greenland Whale, or Bowhead, in the Northeast Atlantic". Arctic Institute of North America. 54 (1): 55–76. doi:10.14430/arctic764.
^ ^a ^b ^c Simmon, Malene; Johnson, Mark; Tyack, Peter; Madsen, Peter T. (2009). "Behaviour and Kinematics of Continuous Ram Filtration in Bowhead Whales (Balaena mysticetus)". Biological Sciences. 276 (1674): 3819–3828. doi:10.1098/rspb.2009.1135.
^ ^a ^b Bowhead Whale. American Cetacean Society. Retrieved on 16 November 2015.
^ "Bowhead Whale : Baleen Whales : Voices in the Sea".
^ Schiffman, Joshua D.; Breen, Matthew (2015). "Comparative oncology: what dogs and other species can teach us about humans with cancer". Phil. Trans. R. Soc. B. 370 (1673): 1–13. doi:10.1098/rstb.2014.0231.
^ George, John C.; Bada, Jeffrey; Zeh, Judith; Scott, Laura; Brown, Stephen E.; O'Hara, Todd; Suydam, Robert (1999). "Age and growth estimates of bowhead whales (Balaena mysticetus) via aspartic acid racemization". Can. J. Zool. 77 (4): 571–580. doi:10.1139/z99-015. {{cite journal}}: Unknown parameter |lastauthoramp= ignored (|name-list-style= suggested) (help)
^ Conroy, Erin. (6 December 2007) Netted whale hit by lance a century ago. MSNBC.
^ 19th-century weapon found in whale. Associated Press via USA Today. 12 June 2007
^ "Can Marine Biology Help Us Live Forever? Bowhead Whale Can Live 200 Years, Is Cancer Resistant". Medical Daily. 6 January 2015. Retrieved 8 November 2015.
^ ^a ^b "Researchers hope this whale's genes will help reverse human aging". The Washington Post. 6 January 2015. Retrieved 8 November 2015.
^ "Scientists map bowhead whale's genome; discover genes responsible for long life". Technie News. 5 January 2015. Retrieved 5 January 2015.
^ "The bowhead whale lives over 200 years. Can its genes tell us why?". Science Daily. 5 January 2015. Retrieved 8 November 2015.
^ Moore, S. E., and R. R. Reeves (1993). "Distribution and Movement". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.
^ Smultea, M.; Fertl, D.; Rugh, D. J.; Bacon, C. E. (2012). Summary of systematic bowhead surveys conducted in the U.S. Beaufort and Chukchi Seas, 1975–2009. U.S. Dep. Commer., NOAA Tech. Memo. NMFS-AFSC-237. p. 48.
^ Foote, A. D.; Kaschner, K.; Schultze, S. E.; Garilao, C.; Ho, S. Y. W.; Post, K.; Higham, T. F. G.; Stokowska, C.; Van Der Es, H.; Embling, C. B.; Gregersen, K.; Johansson, F.; Willerslev, E.; Gilbert, M. T. P. (2013). "Ancient DNA reveals that bowhead whale lineages survived Late Pleistocene climate change and habitat shifts". Nature Communications. 4: 1677. Bibcode:2013NatCo...4E1677F. doi:10.1038/ncomms2714. PMID 23575681.
^ ^a ^b Norwegian Polar Institute. Bowhead whale (Balaena mysticetus). npolar.no
^ Eastern Arctic bowhead whales not threatened. Cbc.ca (16 April 2008). Retrieved on 2011-09-15.
^ Laidre, Kristin (22 January 2009) "Foraging Ecology of Bowhead Whales in West Greenland." Monster Jam. Northwest Fisheries Science Center, Seattle.
^ ^a ^b "Past, Present, and Future for Bowhead Whales (Balaena mysticetus) in Northwest Hudson Bay". A Little Less Arctic: 159–177. doi:10.1007/978-90-481-9121-5_8.
^ http://www.registrelep-sararegistry.gc.ca/species/speciesDetails_e.cfm?sid=131
^ ^a ^b http://www.nwmb.com/iku/2013-11-09-01-41-51/2013-11-09-01-46-43/2008/mar-06-2008-level-of-tah-for-bowhead-whales/551-tab15-df0-1999/file
^ ^a ^b ^c ^d ^e ^f http://www.sararegistry.gc.ca/virtual_sara/files/cosewic/sr_bowhead_whale_e.pdf
^ WWF. "June 2014: Bowheads and breaking ice – Thin Ice Blog".
^ "MPO – Recherche sur les mammifères marins au Canada – MPO Sciences".
^ http://pubs.aina.ucalgary.ca/arctic/Arctic36-1-5.pdf
^ "Bowhead Whale in Churchill Waters – Churchill Polar Bears".
^ ^a ^b ^c Shpak, Olga (19 February 2014). "Второе рождение гренландского кита" (in Russian). RIA Novosti. Retrieved 4 June 2014.
^ "Тур 'наблюдение за китами и плавание вдоль побережья Охотского моря и на Шантарските острова'" (in Russian). Arcticexpedition.ru. 15 August 2000. Retrieved 19 October 2012.
^ "Фотография: Киты подходят совсем близко к берегу" (in Russian). Turizmvnn.ru. 16 April 2011. Retrieved 19 October 2012.
^ "WWF приветствует создание нацпарка в Хабаровском крае" (in Russian). RIA Novosti. 31 December 2013. Retrieved 4 June 2014.
^ Smith, Tim D.; Reeves, Randall R.; Josephson, Elizabeth A.; Lund, Judith N. (27 April 2012). "Spatial and Seasonal Distribution of American Whaling and Whales in the Age of Sail". PLOS ONE. 7 (4): e34905. doi:10.1371/journal.pone.0034905. PMC 3338773. PMID 22558102 – via PLoS Journals.{{cite journal}}: CS1 maint: unflagged free DOI (link)
^ "大阪府漁業協同組合連合会".
^ http://www.asahi.com/articles/ASH6R5VQ2H6RIIPE026.html
^ "いしかり博物誌／第5回 - 北海道石狩市公式ホームページ".
^ Ahgeak, MacLean, Stephen (1 January 2002). "Occurrence, behavior and genetic diversity of bowhead whales in the Western Sea of Okhotsk, Russia".{{cite web}}: CS1 maint: multiple names: authors list (link)
^ Gross A., 2010 Background Document for Bowhead whale Balaena mysticetus. The OSPAR Convention and Musée des Matériaux du Centre de Recherche sur les Monuments Historiques. ISBN 978-1-907390-35-7. retrieved on 24 May 2014
^ Kovacs M.K., Bowhead whale (Balaena mysticetus). Environmental Monitoring of Svalbard and Jan Mayen. retrieved on 27 May 2014
^ Gilg O.; Born W.E. (2004). "Recent sightings of the bowhead whale (Balaena mysticetus) in Northeast Greenland and the Greenland Sea". Polar Biology. 28 (10): 796–801. doi:10.1007/s00300-005-0001-9.
^ Ritchie B. (June 2013) Arctic shorts – bowhead whale. Vimeo. Retrieved 2 June 2014
^ Sala E., 2013 Franz Josef Land Expedition: First Look at Post-Expedition Discoveries. Pristine Seas Expeditions. National Geographic. retrieved on 24 May 2014
^ WIIG Ø., Bachmann L., Janik M.V., Kovacs M.K., Lydersen C., 2007. Spitsbergen Bowhead Whales Revisited. Society for Marine Mammalogy. Retrieved 24 May 2014
^ Johannessen, R. (19 October 2011) Dette er en sensasjon!. The Aftenposten. retrieved on 27 May 2014
^ Nefedova T., Gavrilo M., Gorshkov S., 2013. Летом в Арктике стало меньше льда. Russian Geographical Society. retrieved on 24 May 2014
^ European Cetacean Society. Bowhead whales (Balaena mysticetus) sighting in the Franz Josef Land area.. retrieved on 24 May 2014
^ Scalini I. (19 February 2014) Всемирный день китов. Russian Arctic National Park. retrieved on 24 May 2014
^ Walker, Matt. "Secret whale refuge discovered".
^ "Exceptional sighting of 80+ Bowhead Whales".
^ Lennartz T. (29 April 2013) New Bowhead Whale Molting Location Found. Arctic Kingdom. retrieved on 9 June 2014
^ Polar Bears and Glaciers of Baffin Island Webinar on Vimeo. Arctic Kingdom. 2014. retrieved on 9 June 2014
^ Montague, J. J. (1993). "Introduction". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.
^ Philo, L. M., E. B. Shotts, and J. C. George (1993). "Morbidity and Mortality". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.
^ ^a ^b ^c "Bowhead Whale (Balaena mysticetus)". National Oceanic and Atmospheric Administration. Retrieved 23 November 2015.
^ Woodby, D. A. and D. B. Botkin (1993). "Stock Sizes Prior to Commercial Whaling". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.
^ Ross, W. G. (1993). "Commercial Whaling in the North Atlantic Sector". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.
^ ^a ^b Bockstoce, John (1986). Whales, Ice, & Men: The History of Whaling in the Western Arctic. University of Washington Press. ISBN 0-295-97447-8.
^ Vaughan, R. (1984). Historical survey of the European whaling industry. In Arctic Whaling: Proceedings of the International Symposium, pp. 121-145. University of Groningen.
^ "Bowhead Whale". WWF. Retrieved 1 December 2015.
^ "Subsistence Hunting, Alaska Department of Fish and Game". www.adfg.alaska.gov. Retrieved 24 February 2016.
^ Bowhead Whale, Walrus and Polar Bears of Foxe Basin on Vimeo. Arctic Kingdom. 2011. retrieved on 9 June 2014
^ "Appendix I" of the Convention on the Conservation of Migratory Species of Wild Animals (CMS). As amended by the Conference of the Parties in 1985, 1988, 1991, 1994, 1997, 1999, 2002, 2005, and 2008. Effective: 5 March 2009.
^ Vladislav Raevskii. Retrieved 1 June 2014

External links

"Bowhead whale: 200 year old whales". BBC Ocean Giants. October 2013.
"The Bowhead Whale". Voices in the Sea, University of California San Diego. October 2013.
"Bowhead Whale (Balaena mysticetus)". Office of Protected Resources, NOAA Fisheries. October 2013.
"Bowhead whale (Balaena mysticetus)". ARKive. October 2013.
"Harpoon may prove whale was at least 115 years old". World History Blog. October 2013.
"In Search of the Bowhead Whale". NFB.ca. October 2013. A documentary by Bill Mason from 1974 following an expedition that searches out and meets the bowhead and beluga.
"Balaena mysticetus". The TerraMar Project. March 2016.

[msw3-1] Mead, J. G.; Brownell, R. L. Jr. (2005). "Order Cetacea". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 723–743. ISBN 978-0-8018-8221-0. OCLC 62265494.

[iucn-2] Template:IUCN2012.2

[Rugh2008-3] Rugh, David J.; Shelden, Kim E. W. (2008). "Bowhead Whale". In Perrin, William F.; Würsig, Bernd; Thewissen, J. G. M. (eds.). Encyclopedia of Marine Mammals (Second ed.). Academic Press. p. 131. ISBN 978-0-12-373553-9.

[4] Scammon, Charles M. (1874) The Marine Mammals of the North-Western Coast of North America, New York: G.P. Putnam's Sons, ISBN 1-59714-061-9.

[guinness-record-5] Guinness World Records (14 November 2007). "Whale of a time!". Archived from the original on 17 November 2007. Retrieved 4 June 2009. {{cite web}}: Unknown parameter |deadurl= ignored (|url-status= suggested) (help)

[6] Linnaeus, C (1758). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis [System of nature through the three kingdoms of nature, according to the classes, orders, genera, species, with the characters, the differences, synonyms, places.] (in Latin). Vol. I (tenth, reformed ed.). Holmiae. (Laurentii Salvii). p. 824.

[iucn-bowhead-7] Template:IUCN2012.2 "The taxonomy is not in doubt.... Concerning common names, the species was once commonly known in the North Atlantic and adjacent Arctic as the Greenland Right Whale. However, the common name Bowhead Whale is now generally used for the species."

[8] Rice, Dale W. (1998). Marine mammals of the world: systematics and distribution. Society of Marine Mammalogy Special Publication No. 4. ISBN 1-891276-03-4.

[PerrinWursig2008-9] Kenney, Robert D. (2008). "Right Whales (Eubalaena glacialis, E. japonica, and E. australis)". In Perrin, William F.; Würsig, Bernd; Thewissen, J. G. M. (eds.). Encyclopedia of Marine Mammals. Academic Press. pp. 962–969. ISBN 978-0-12-373553-9.

[Rosenbaum-10] "World-wide genetic differentiation of Eubalaena: Questioning the number of right whale species" (PDF). Molecular Ecology. 9 (11): 1793–802. 2000. doi:10.1046/j.1365-294x.2000.01066.x. PMID 11091315. {{cite journal}}: Cite uses deprecated parameter |authors= (help); Unknown parameter |lastauthoramp= ignored (|name-list-style= suggested) (help)

[11] "Whale bones found in highway were not from mystery whale". ScienceNordic.com. 7 February 2013.

[BehWur-12] Würsig, B. and C. Clark (1993). "Behavior". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.

[FFELow-13] Lowry, L. F. (1993). "Foods and Feeding Ecology". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.

[AnaPhyHald-14] Haldiman, J. T. and R. J. Tarpley (1993). "Anatomy & Physiology". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.

[15] Scoresby, William (1820). An Account of the Arctic Regions with a History and a Description of the Northern Whale-Fishery. Edinburgh.

[CWNPBock-16] Bockstoce, J. R., and J. J. Burns (1993). "Commercial Whaling in the North Pacific Sector". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.

[RepKoski-17] Koski, William R., Rolph A. Davis, Gary W. Miller, and David E. Withrow (1993). "Reproduction". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy. p. 245.

[18] Eilperin, Juliet (18 October 2012). "Bowhead whales lost genetic diversity, study shows". The Washington Post.

[19] Ford Jr, T. J.; Werth, A. J.; George, J. C. (2013). "An intraoral thermoregulatory organ in the bowhead whale (Balaena mysticetus), the corpus cavernosum maxillaris". Anatomical Record. 296 (4): 701–708. doi:10.1002/ar.22681. PMID 23450839.

[Finely,_K.J._2001-20] Finley, K.J. (2001). "Natural History and Conservation of the Greenland Whale, or Bowhead, in the Northeast Atlantic". Arctic Institute of North America. 54 (1): 55–76. doi:10.14430/arctic764.

[Simmon,_Malene_2009-21] Simmon, Malene; Johnson, Mark; Tyack, Peter; Madsen, Peter T. (2009). "Behaviour and Kinematics of Continuous Ram Filtration in Bowhead Whales (Balaena mysticetus)". Biological Sciences. 276 (1674): 3819–3828. doi:10.1098/rspb.2009.1135.

[ACS_2015-22] Bowhead Whale. American Cetacean Society. Retrieved on 16 November 2015.

[23] "Bowhead Whale : Baleen Whales : Voices in the Sea".

[24] Schiffman, Joshua D.; Breen, Matthew (2015). "Comparative oncology: what dogs and other species can teach us about humans with cancer". Phil. Trans. R. Soc. B. 370 (1673): 1–13. doi:10.1098/rstb.2014.0231.

[25] George, John C.; Bada, Jeffrey; Zeh, Judith; Scott, Laura; Brown, Stephen E.; O'Hara, Todd; Suydam, Robert (1999). "Age and growth estimates of bowhead whales (Balaena mysticetus) via aspartic acid racemization". Can. J. Zool. 77 (4): 571–580. doi:10.1139/z99-015. {{cite journal}}: Unknown parameter |lastauthoramp= ignored (|name-list-style= suggested) (help)

[26] Conroy, Erin. (6 December 2007) Netted whale hit by lance a century ago. MSNBC.

[27] 19th-century weapon found in whale. Associated Press via USA Today. 12 June 2007

[28] "Can Marine Biology Help Us Live Forever? Bowhead Whale Can Live 200 Years, Is Cancer Resistant". Medical Daily. 6 January 2015. Retrieved 8 November 2015.

[Washington_Post_2015-29] "Researchers hope this whale's genes will help reverse human aging". The Washington Post. 6 January 2015. Retrieved 8 November 2015.

[30] "Scientists map bowhead whale's genome; discover genes responsible for long life". Technie News. 5 January 2015. Retrieved 5 January 2015.

[31] "The bowhead whale lives over 200 years. Can its genes tell us why?". Science Daily. 5 January 2015. Retrieved 8 November 2015.

[DisMovMoo-32] Moore, S. E., and R. R. Reeves (1993). "Distribution and Movement". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.

[33] Smultea, M.; Fertl, D.; Rugh, D. J.; Bacon, C. E. (2012). Summary of systematic bowhead surveys conducted in the U.S. Beaufort and Chukchi Seas, 1975–2009. U.S. Dep. Commer., NOAA Tech. Memo. NMFS-AFSC-237. p. 48.

[34] Foote, A. D.; Kaschner, K.; Schultze, S. E.; Garilao, C.; Ho, S. Y. W.; Post, K.; Higham, T. F. G.; Stokowska, C.; Van Der Es, H.; Embling, C. B.; Gregersen, K.; Johansson, F.; Willerslev, E.; Gilbert, M. T. P. (2013). "Ancient DNA reveals that bowhead whale lineages survived Late Pleistocene climate change and habitat shifts". Nature Communications. 4: 1677. Bibcode:2013NatCo...4E1677F. doi:10.1038/ncomms2714. PMID 23575681.

[Bowhead_whale_Balaena_mysticetus-35] Norwegian Polar Institute. Bowhead whale (Balaena mysticetus). npolar.no

[36] Eastern Arctic bowhead whales not threatened. Cbc.ca (16 April 2008). Retrieved on 2011-09-15.

[37] Laidre, Kristin (22 January 2009) "Foraging Ecology of Bowhead Whales in West Greenland." Monster Jam. Northwest Fisheries Science Center, Seattle.

[link.springer.com-38] "Past, Present, and Future for Bowhead Whales (Balaena mysticetus) in Northwest Hudson Bay". A Little Less Arctic: 159–177. doi:10.1007/978-90-481-9121-5_8.

[39] ttp://www.registrelep-sararegistry.gc.ca/species/speciesDetails_e.cfm?sid=131

[NWMB-40] ttp://www.nwmb.com/iku/2013-11-09-01-41-51/2013-11-09-01-46-43/2008/mar-06-2008-level-of-tah-for-bowhead-whales/551-tab15-df0-1999/file

[sararegistry-41] ^ ^a ^b ^c ^d ^e ^f http://www.sararegistry.gc.ca/virtual_sara/files/cosewic/sr_bowhead_whale_e.pdf

[42] WWF. "June 2014: Bowheads and breaking ice – Thin Ice Blog".

[43] "MPO – Recherche sur les mammifères marins au Canada – MPO Sciences".

[44] ttp://pubs.aina.ucalgary.ca/arctic/Arctic36-1-5.pdf

[45] "Bowhead Whale in Churchill Waters – Churchill Polar Bears".

[Olga_Shpak._2014-46] Shpak, Olga (19 February 2014). "Второе рождение гренландского кита" (in Russian). RIA Novosti. Retrieved 4 June 2014.

[47] "Тур 'наблюдение за китами и плавание вдоль побережья Охотского моря и на Шантарските острова'" (in Russian). Arcticexpedition.ru. 15 August 2000. Retrieved 19 October 2012.

[48] "Фотография: Киты подходят совсем близко к берегу" (in Russian). Turizmvnn.ru. 16 April 2011. Retrieved 19 October 2012.

[49] "WWF приветствует создание нацпарка в Хабаровском крае" (in Russian). RIA Novosti. 31 December 2013. Retrieved 4 June 2014.

[50] Smith, Tim D.; Reeves, Randall R.; Josephson, Elizabeth A.; Lund, Judith N. (27 April 2012). "Spatial and Seasonal Distribution of American Whaling and Whales in the Age of Sail". PLOS ONE. 7 (4): e34905. doi:10.1371/journal.pone.0034905. PMC 3338773. PMID 22558102 – via PLoS Journals.{{cite journal}}: CS1 maint: unflagged free DOI (link)

[51] "大阪府漁業協同組合連合会".

[52] ttp://www.asahi.com/articles/ASH6R5VQ2H6RIIPE026.html

[53] "いしかり博物誌／第5回 - 北海道石狩市公式ホームページ".

[54] Ahgeak, MacLean, Stephen (1 January 2002). "Occurrence, behavior and genetic diversity of bowhead whales in the Western Sea of Okhotsk, Russia".{{cite web}}: CS1 maint: multiple names: authors list (link)

[55] Gross A., 2010 Background Document for Bowhead whale Balaena mysticetus. The OSPAR Convention and Musée des Matériaux du Centre de Recherche sur les Monuments Historiques. ISBN 978-1-907390-35-7. retrieved on 24 May 2014

[56] Kovacs M.K., Bowhead whale (Balaena mysticetus). Environmental Monitoring of Svalbard and Jan Mayen. retrieved on 27 May 2014

[57] Gilg O.; Born W.E. (2004). "Recent sightings of the bowhead whale (Balaena mysticetus) in Northeast Greenland and the Greenland Sea". Polar Biology. 28 (10): 796–801. doi:10.1007/s00300-005-0001-9.

[58] Ritchie B. (June 2013) Arctic shorts – bowhead whale. Vimeo. Retrieved 2 June 2014

[59] Sala E., 2013 Franz Josef Land Expedition: First Look at Post-Expedition Discoveries. Pristine Seas Expeditions. National Geographic. retrieved on 24 May 2014

[60] WIIG Ø., Bachmann L., Janik M.V., Kovacs M.K., Lydersen C., 2007. Spitsbergen Bowhead Whales Revisited. Society for Marine Mammalogy. Retrieved 24 May 2014

[61] Johannessen, R. (19 October 2011) Dette er en sensasjon!. The Aftenposten. retrieved on 27 May 2014

[62] Nefedova T., Gavrilo M., Gorshkov S., 2013. Летом в Арктике стало меньше льда. Russian Geographical Society. retrieved on 24 May 2014

[63] European Cetacean Society. Bowhead whales (Balaena mysticetus) sighting in the Franz Josef Land area.. retrieved on 24 May 2014

[64] Scalini I. (19 February 2014) Всемирный день китов. Russian Arctic National Park. retrieved on 24 May 2014

[65] Walker, Matt. "Secret whale refuge discovered".

[66] "Exceptional sighting of 80+ Bowhead Whales".

[67] Lennartz T. (29 April 2013) New Bowhead Whale Molting Location Found. Arctic Kingdom. retrieved on 9 June 2014

[68] Polar Bears and Glaciers of Baffin Island Webinar on Vimeo. Arctic Kingdom. 2014. retrieved on 9 June 2014

[MonIntro-69] Montague, J. J. (1993). "Introduction". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.

[MMPhil-70] Philo, L. M., E. B. Shotts, and J. C. George (1993). "Morbidity and Mortality". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.

[NOAA_2015-71] "Bowhead Whale (Balaena mysticetus)". National Oceanic and Atmospheric Administration. Retrieved 23 November 2015.

[SSWoo-72] Woodby, D. A. and D. B. Botkin (1993). "Stock Sizes Prior to Commercial Whaling". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.

[Ross-73] Ross, W. G. (1993). "Commercial Whaling in the North Atlantic Sector". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. The Bowhead Whale. Special Publication No. 2: The Society for Marine Mammalogy.

[Bockstoce1986-74] Bockstoce, John (1986). Whales, Ice, & Men: The History of Whaling in the Western Arctic. University of Washington Press. ISBN 0-295-97447-8.

[Vaughan1984-75] Vaughan, R. (1984). Historical survey of the European whaling industry. In Arctic Whaling: Proceedings of the International Symposium, pp. 121-145. University of Groningen.

[WWF_2015-76] "Bowhead Whale". WWF. Retrieved 1 December 2015.

[77] "Subsistence Hunting, Alaska Department of Fish and Game". www.adfg.alaska.gov. Retrieved 24 February 2016.

[78] Bowhead Whale, Walrus and Polar Bears of Foxe Basin on Vimeo. Arctic Kingdom. 2011. retrieved on 9 June 2014

[Appendices-79] "Appendix I" of the Convention on the Conservation of Migratory Species of Wild Animals (CMS). As amended by the Conference of the Parties in 1985, 1988, 1991, 1994, 1997, 1999, 2002, 2005, and 2008. Effective: 5 March 2009.

[80] Vladislav Raevskii. Retrieved 1 June 2014

[1]

[2]

[3]

[4]

[5]

[6]

[7]

[8]

[9]

[10]

[11]

[12]

[13]

[14]

[15]

[16]

[17]

[18]

[19]

[20]

[21]

[22]

[23]

[24]

[25]

[26]

[27]

[28]

[29]

[30]

[31]

[32]

[33]

[34]

[35]

[36]

[37]

[38]

[39]

[40]

[41]

[42]

[43]

[44]

[45]

[46]

[47]

[48]

[49]

[50]

[51]

[52]

[53]

[54]

[55]

[56]

[57]

[58]

[59]

[60]

[61]

[62]

[63]

[64]

[65]

[66]

[67]

[68]

[69]

[70]

[71]

[72]

[73]

[74]

[75]

[76]

[77]

[78]

[79]

[80]

@@ Line 5: / Line 5: @@
 | status_ref =<ref name="iucn">{{IUCN2012.2|assessors=Reilly, S.B., Bannister, J.L., Best, P.B., Brown, M., Brownell Jr., R.L., Butterworth, D.S., Clapham, P.J., Cooke, J., Donovan, G.P., Urbán, J. & Zerbini, A.N.|year=2012|id=2467|title=Balaena mysticetus|downloaded=18 October 2012}}</ref>
 | image = Bowheads42.jpg
-| image2 = Bowhead_whale_size.svg
+| image2 = Bowhead whale size.svg
 | image2_caption = Size compared to an average human
 | regnum = [[Animal]]ia
@@ Line 21: / Line 21: @@
 | range_map_caption = Bowhead whale range
 }}
+The '''bowhead whale''' (''Balaena mysticetus'') is a species of the family [[Balaenidae]], in [[Suborder (biology)|suborder]] [[Mysticeti]], and [[genus]] ''[[Balaena]]'', once thought to include the [[right whale]].
-The '''bowhead whale''' (''Balaena mysticetus'') is a species of the family [[Balaenidae]], in [[Suborder (biology)|suborder]] [[Mysticeti]], and [[genus]] ''[[Balaena]]'', once thought to include the [[right whale]]. A stocky dark-colored whale without a [[dorsal fin]], it can grow {{convert|14|to|18|m|abbr=on}} in length. This thick-bodied species can weigh from {{convert|75|to|100|t}}.<ref name=Rugh2008>{{cite book |last1=Rugh |first1=David J. |last2=Shelden |first2=Kim E. W. |editor1-last=Perrin |editor1-first=William F. |editor2-last=Würsig |editor2-first=Bernd |editor2-link=Bernd Würsig |editor3-last=Thewissen |editor3-first=J. G. M. |date=2008 |title=Encyclopedia of Marine Mammals |edition=Second |chapter=Bowhead Whale |publisher=Academic Press |isbn=978-0-12-373553-9 |page=131}}</ref> They live entirely in fertile [[Arctic Ocean|Arctic]] and sub-Arctic waters, unlike other whales that migrate to low latitude waters to feed or reproduce. The bowhead was also known as the '''Greenland right whale''' or '''Arctic whale'''. American whalemen called them the '''steeple-top''', '''polar whale''',<ref>Scammon, Charles M. (1874) [https://archive.org/details/marinemammalsofn00scam ''The Marine Mammals of the North-Western Coast of North America''], New York: G.P. Putnam's Sons, ISBN 1597140619.</ref> or '''Russia''' or '''Russian whale'''. The bowhead has the largest mouth of any animal.<ref name="guinness-record">{{cite web|author=[[Guinness World Records]] |date=14 November 2007 |title=Whale of a time! |url=http://www.guinnessworldrecords.com/news/2007/11/071114.aspx |accessdate=4 June 2009 |deadurl=yes |archiveurl=https://web.archive.org/web/20071117060430/http://www.guinnessworldrecords.com/news/2007/11/071114.aspx |archivedate=17 November 2007 }}</ref>
+A stocky dark-colored whale without a [[dorsal fin]], it can grow {{convert|14|to|18|m|abbr=on}} in length. This thick-bodied species can weigh from {{convert|75|to|100|t}}.<ref name=Rugh2008>{{cite book |last1=Rugh |first1=David J. |last2=Shelden |first2=Kim E. W. |editor1-last=Perrin |editor1-first=William F. |editor2-last=Würsig |editor2-first=Bernd |editor2-link=Bernd Würsig |editor3-last=Thewissen |editor3-first=J. G. M. |date=2008 |title=Encyclopedia of Marine Mammals |edition=Second |chapter=Bowhead Whale |publisher=Academic Press |isbn=978-0-12-373553-9 |page=131}}</ref> They live entirely in fertile [[Arctic Ocean|Arctic]] and sub-Arctic waters, unlike other whales that migrate to low latitude waters to feed or reproduce. The bowhead was also known as the '''Greenland right whale''' or '''Arctic whale'''. American whalemen called them the '''steeple-top''', '''polar whale''',<ref>Scammon, Charles M. (1874) [https://archive.org/details/marinemammalsofn00scam ''The Marine Mammals of the North-Western Coast of North America''], New York: G.P. Putnam's Sons, ISBN 1-59714-061-9.</ref> or '''Russia''' or '''Russian whale'''. The bowhead has the largest mouth of any animal.<ref name="guinness-record">{{cite web|author=[[Guinness World Records]] |date=14 November 2007 |title=Whale of a time! |url=http://www.guinnessworldrecords.com/news/2007/11/071114.aspx |accessdate=4 June 2009 |deadurl=yes |archiveurl=https://web.archive.org/web/20071117060430/http://www.guinnessworldrecords.com/news/2007/11/071114.aspx |archivedate=17 November 2007 }}</ref>
-The bowhead was an early [[whaling]] target. The population was severely reduced before a 1966 [[Moratorium (law)|moratorium]] was passed to protect the species. Through conservation efforts, the bowhead population has since recovered and is now rated "Least Concern" on the [[IUCN Red List of Threatened Species]].<ref name="iucn"/>
+The bowhead was an early [[whaling]] target. The population was severely reduced before a 1966 [[Moratorium (law)|moratorium]] was passed to protect the species. Through conservation efforts, the bowhead population has since recovered and is now rated "Least Concern" on the [[IUCN Red List of Threatened Species]].<ref name="iucn" />
-==Taxonomy==
+== Taxonomy ==
 {{See also|Evolution of cetaceans}}
 [[Carl Linnaeus]] first described this whale in the 10th edition of his ''[[Systema Naturae]]'' (1758).<ref>{{cite book |last=Linnaeus |first=C |authorlink=Carolus Linnaeus |date=1758 |title=Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. |trans-title=System of nature through the three kingdoms of nature, according to the classes, orders, genera, species, with the characters, the differences, synonyms, places. |volume=I |edition=tenth, reformed |language=Latin |publisher=Holmiae. (Laurentii Salvii) |page=824 |url=http://dz1.gdz-cms.de/index.php?id=img&no_cache=1&IDDOC=265100}}</ref> Seemingly identical to its cousins in the [[North Atlantic]], [[North Pacific]] and [[Southern Ocean]]s, they were all thought to be a single species, collectively known as the "right whale", and given the [[binomial nomenclature|binomial name]] ''Balaena mysticetus''.
-Today, the bowhead whale occupies a [[monotypic taxon|monotypic]] [[genus]], separate from the [[right whale]]s, as was proposed by the work of [[John Edward Gray]] in 1821.<ref name="iucn-bowhead">{{IUCN2012.2|assessors=Reilly, S.B., Bannister, J.L., Best, P.B., Brown, M., Brownell Jr., R.L., Butterworth, D.S., Clapham, P.J., Cooke, J., Donovan, G.P., Urbán, J. & Zerbini, A.N.|year=2012|id=2467|title=Balaena mysticetus|downloaded=18 October 2012}} "The taxonomy is not in doubt.... Concerning common names, the species was once commonly known in the North Atlantic and adjacent Arctic as the Greenland Right Whale. However, the common name Bowhead Whale is now generally used for the species."</ref> For the next 180 years, the family Balaenidae was the subject of great [[taxonometrics|taxonometric]] debate. Authorities have repeatedly recategorized the three populations of right whale plus the bowhead whale, as one, two, three or four species, either in a single genus or in two separate genera. Eventually, it was recognized that bowheads and right whales were in fact different, but there was still no strong consensus as to whether they shared a single genus or two. As recently as 1998, Dale Rice, in his comprehensive and otherwise authoritative classification, ''Marine mammals of the world: systematics and distribution'', listed just two species: ''B. glacialis'' (the right whales) and ''B. mysticetus'' (the bowheads).<ref>{{cite book |last=Rice |first=Dale W. |date=1998 |title=Marine mammals of the world: systematics and distribution |publisher=Society of Marine Mammalogy Special Publication No. 4 |isbn=1891276034}}</ref>
+Today, the bowhead whale occupies a [[monotypic taxon|monotypic]] [[genus]], separate from the [[right whale]]s, as was proposed by the work of [[John Edward Gray]] in 1821.<ref name="iucn-bowhead">{{IUCN2012.2|assessors=Reilly, S.B., Bannister, J.L., Best, P.B., Brown, M., Brownell Jr., R.L., Butterworth, D.S., Clapham, P.J., Cooke, J., Donovan, G.P., Urbán, J. & Zerbini, A.N.|year=2012|id=2467|title=Balaena mysticetus|downloaded=18 October 2012}} "The taxonomy is not in doubt.... Concerning common names, the species was once commonly known in the North Atlantic and adjacent Arctic as the Greenland Right Whale. However, the common name Bowhead Whale is now generally used for the species."</ref> For the next 180 years, the family Balaenidae was the subject of great [[taxonometrics|taxonometric]] debate. Authorities have repeatedly recategorized the three populations of right whale plus the bowhead whale, as one, two, three or four species, either in a single genus or in two separate genera. Eventually, it was recognized that bowheads and right whales were in fact different, but there was still no strong consensus as to whether they shared a single genus or two. As recently as 1998, Dale Rice, in his comprehensive and otherwise authoritative classification, ''Marine mammals of the world: systematics and distribution'', listed just two species: ''B. glacialis'' (the right whales) and ''B. mysticetus'' (the bowheads).<ref>{{cite book |last=Rice |first=Dale W. |date=1998 |title=Marine mammals of the world: systematics and distribution |publisher=Society of Marine Mammalogy Special Publication No. 4 |isbn=1-891276-03-4}}</ref>
 Studies in the 2000s finally provided clear evidence that the ''three'' living right whale species do comprise a [[phylogenetic]] lineage, ''distinct from the bowhead'', and that the bowhead and the right whales are rightly classified into two separate genera.<ref name="PerrinWursig2008">{{cite encyclopedia
@@ Line 70: / Line 72: @@
 An unknown species of right whale, the so-called "Swedenborg whale" which was proposed by [[Emanuel Swedenborg]] in the 18th century, was once thought to be a North Atlantic right whale by scientific consensus. However based on later DNA analysis those fossil bones claimed to be from "Swedenborg whales" were confirmed to be from bowhead whales.<ref>{{cite web |date=7 February 2013 |title=Whale bones found in highway were not from mystery whale |publisher=ScienceNordic.com |url=http://sciencenordic.com/whale-bones-found-highway-were-not-mystery-whale}}</ref>
-==Description==
+== Description ==
 [[File:GreenlandWhaleLyd3.jpg|thumb|Skeleton of a bowhead whale|alt=Drawing of long backbone, 13 ribs (two vestigial) large, curved upper and lower jawbones that occupy 1/3 of the body, 4 multijointed "fingers" inside pectoral fin and connecting bone, enclosed in body outline]]
 [[File:Faroe stamp 198 Baleana mysticetus.jpg|thumb|Bowhead whales (on [[Faroe Islands|Faroese]] stamp)|Stamp showing drawing of mother and calf from [[Faroe Islands]]]]
 The bowhead whale has a large, robust, dark-colored body and a white chin/lower jaw. The whale has a massive triangular [[skull]], which the whale uses to break through the [[Arctic]] ice to breathe. [[Inuit]] hunters have reported bowheads surfacing through {{convert|60|cm|abbr=on}} of ice.<ref name="BehWur">Würsig, B. and C. Clark (1993). "Behavior". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref> The bowhead also has a strongly bowed lower jaw and a narrow upper jaw. Its [[baleen]] is the longest of that of any whale, at {{convert|3|m|abbr =on}}, and is used to strain tiny prey from the water. The bowhead whale has paired blowholes, at the highest point of the head, which can spout a blow {{convert|20|ft|abbr=on}} high. The whale's [[blubber]] is the thickest of that of any animal, with a maximum of {{convert|43|-|50|cm|abbr=on}}.<ref name="FFELow">Lowry, L. F. (1993). "Foods and Feeding Ecology". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref> Unlike most cetaceans, the bowhead does not have a [[dorsal fin]].<ref name="AnaPhyHald">Haldiman, J. T. and R. J. Tarpley (1993). "Anatomy & Physiology". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref>
-Bowhead whales are comparable in size to the three species of [[right whales]]. According to whaling captain [[William Scoresby]] Jr., the longest bowhead he measured was {{convert|17.7|m|abbr=on}} long, while the longest measurement he had ever heard of was of a {{convert|20.4|m|abbr=on}} whale caught at [[Qeqertarsuaq|Godhavn]], Greenland, in early 1813. He also spoke of one, caught near [[Spitsbergen]] around 1800, that was allegedly nearly {{convert|21.3|m|abbr=on}} long.<ref>{{cite book |last=Scoresby |first=William |date=1820 |title=An Account of the Arctic Regions with a History and a Description of the Northern Whale-Fishery |publisher=Edinburgh |url=https://archive.org/details/accountofarcticr01scor}}</ref> In 1850, an American vessel claimed to have caught a {{convert|24.54|m|abbr=on}} individual in the Western Arctic.<ref name="CWNPBock"/> It is questionable whether these lengths were actually measured. The longest reliably measured lengths of the sexes were {{convert|16.2|m|abbr=on}} in a male and {{convert|18|m|abbr=on}} in a female, both landed by natives in Alaska.<ref name="RepKoski">Koski, William R., Rolph A. Davis, Gary W. Miller, and David E. Withrow (1993). "Reproduction". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy. p. 245.</ref> On average, female bowheads are larger than males.
+Bowhead whales are comparable in size to the three species of [[right whales]]. According to whaling captain [[William Scoresby]] Jr., the longest bowhead he measured was {{convert|17.7|m|abbr=on}} long, while the longest measurement he had ever heard of was of a {{convert|20.4|m|abbr=on}} whale caught at [[Qeqertarsuaq|Godhavn]], Greenland, in early 1813. He also spoke of one, caught near [[Spitsbergen]] around 1800, that was allegedly nearly {{convert|21.3|m|abbr=on}} long.<ref>{{cite book |last=Scoresby |first=William |date=1820 |title=An Account of the Arctic Regions with a History and a Description of the Northern Whale-Fishery |publisher=Edinburgh |url=https://archive.org/details/accountofarcticr01scor}}</ref> In 1850, an American vessel claimed to have caught a {{convert|24.54|m|abbr=on}} individual in the Western Arctic.<ref name="CWNPBock" /> It is questionable whether these lengths were actually measured. The longest reliably measured lengths of the sexes were {{convert|16.2|m|abbr=on}} in a male and {{convert|18|m|abbr=on}} in a female, both landed by natives in Alaska.<ref name="RepKoski">Koski, William R., Rolph A. Davis, Gary W. Miller, and David E. Withrow (1993). "Reproduction". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy. p. 245.</ref> On average, female bowheads are larger than males.
 Analysis of hundreds of DNA samples from living whales and from [[baleen]] used in vessels, toys, and housing material has shown that Arctic bowhead whales have lost a significant portion of their genetic diversity in the past 500 years. Bowheads originally crossed ice-covered inlets and straits to exchange genes between Atlantic and Pacific populations. This conclusion was derived from analyzing maternal lineage using [[mitochondrial DNA]]. Whaling and climatic cooling during the [[Little Ice Age]], from the 16th century to the 19th, is supposed to have reduced the whales’ summer habitats, which explains the loss of genetic diversity.<ref>{{cite web |last=Eilperin |first=Juliet |date=18 October 2012 |title=Bowhead whales lost genetic diversity, study shows |publisher=The Washington Post |url=http://www.washingtonpost.com/national/health-science/bowhead-whales-lost-genetic-diversity-study-shows/2012/10/18/8b0324d0-186c-11e2-a55c-39408fbe6a4b_story.html?wprss=rss_health-science&tid=pp_widget}}</ref>
@@ Line 94: / Line 96: @@
 }}</ref>
-==Behavior==
+== Behavior ==
 [[File:Grönlandwal 6-1999.jpg|thumb|upright|Resting on water surface in [[Foxe Basin]]]]
 [[File:Bowhead-1 Kate Stafford edit (16272151841).jpg|thumb|Breaching off Alaskan coast]]
-===Swimming===
+=== Swimming ===
-{{see also|Whale surfacing behaviour}}
+{{See also|Whale surfacing behaviour}}
-Bowhead whales are not social animals, typically traveling alone or in small pods of up to 6. They are able to dive and remain submerged underwater for up to an hour. However, the time spent underwater in a single dive is usually limited to 9–18 minutes.<ref name="BehWur"/> Bowheads are not thought to be deep divers but they can reach a depth of up to {{convert|500|ft|abbr=on}}. These whales are slow swimmers, normally traveling at about {{convert|2|–|5|km/h|abbr=on}}.<ref name= "Finely, K.J. 2001" /> When fleeing from danger, they can travel at a speed of {{convert|10|km/h|abbr=on}}. During periods of feeding, the average swim speed is reduced to {{convert|1.1|–|2.5|m/s|abbr=on}}.<ref name= "Simmon, Malene 2009">{{cite journal |author1=Simmon, Malene |author2=Johnson, Mark |author3=Tyack, Peter |author4=Madsen, Peter T. |date=2009 |title= Behaviour and Kinematics of Continuous Ram Filtration in Bowhead Whales (''Balaena mysticetus'') |journal=Biological Sciences |volume=276 |issue=1674 |pages=3819–3828 |doi= 10.1098/rspb.2009.1135}}</ref>
+Bowhead whales are not social animals, typically traveling alone or in small pods of up to 6. They are able to dive and remain submerged underwater for up to an hour. However, the time spent underwater in a single dive is usually limited to 9–18 minutes.<ref name="BehWur" /> Bowheads are not thought to be deep divers but they can reach a depth of up to {{convert|500|ft|abbr=on}}. These whales are slow swimmers, normally traveling at about {{convert|2|–|5|km/h|abbr=on}}.<ref name="Finely, K.J. 2001" /> When fleeing from danger, they can travel at a speed of {{convert|10|km/h|abbr=on}}. During periods of feeding, the average swim speed is reduced to {{convert|1.1|–|2.5|m/s|abbr=on}}.<ref name="Simmon, Malene 2009">{{cite journal |author1=Simmon, Malene |author2=Johnson, Mark |author3=Tyack, Peter |author4=Madsen, Peter T. |date=2009 |title= Behaviour and Kinematics of Continuous Ram Filtration in Bowhead Whales (''Balaena mysticetus'') |journal=Biological Sciences |volume=276 |issue=1674 |pages=3819–3828 |doi= 10.1098/rspb.2009.1135}}</ref>
-===Feeding===
+=== Feeding ===
-The head of the bowhead whale comprises a third of its body length, creating an enormous feeding apparatus.<ref name= "Simmon, Malene 2009"/> Bowhead whales are [[filter feeders]], feeding by swimming forward with mouth wide open.<ref name="BehWur"/> The whale has hundreds of overlapping [[baleen]] plates consisting of [[keratin]] hanging from each side of the upper jaw. The mouth has a large upturning lip on the lower jaw that helps to reinforce and hold the baleen plates within the mouth. This also prevents buckling or breakage of the plates from the pressure of the water passing through them as the whale advances. To feed, water is filtered through the fine hairs of keratin of the baleen plates, trapping the prey inside near the tongue where it is then swallowed.<ref name= "ACS 2015">[http://acsonline.org/fact-sheets/bowhead-whale/ Bowhead Whale]. American Cetacean Society. Retrieved on 16 November 2015.</ref> The diet consists of mostly [[zooplankton]] which includes [[copepods]], [[amphipods]], and many other [[crustaceans]].<ref name= "Simmon, Malene 2009"/> Approximately {{convert|2|ST|LT MT}} of food is consumed each day.<ref name= "ACS 2015"/> While foraging, bowheads are solitary or occur in groups of two to ten or more.<ref name="FFELow"/>
+The head of the bowhead whale comprises a third of its body length, creating an enormous feeding apparatus.<ref name="Simmon, Malene 2009" /> Bowhead whales are [[filter feeders]], feeding by swimming forward with mouth wide open.<ref name="BehWur" /> The whale has hundreds of overlapping [[baleen]] plates consisting of [[keratin]] hanging from each side of the upper jaw. The mouth has a large upturning lip on the lower jaw that helps to reinforce and hold the baleen plates within the mouth. This also prevents buckling or breakage of the plates from the pressure of the water passing through them as the whale advances. To feed, water is filtered through the fine hairs of keratin of the baleen plates, trapping the prey inside near the tongue where it is then swallowed.<ref name="ACS 2015">[http://acsonline.org/fact-sheets/bowhead-whale/ Bowhead Whale]. American Cetacean Society. Retrieved on 16 November 2015.</ref> The diet consists of mostly [[zooplankton]] which includes [[copepods]], [[amphipods]], and many other [[crustaceans]].<ref name="Simmon, Malene 2009" /> Approximately {{convert|2|ST|LT MT}} of food is consumed each day.<ref name="ACS 2015" /> While foraging, bowheads are solitary or occur in groups of two to ten or more.<ref name="FFELow" />
-===Vocalization===
+=== Vocalization ===
-Bowhead whales are highly vocal<ref>[http://cetus.ucsd.edu/voicesinthesea_org/species/baleenWhales/bowhead.html Voices of the Sea]</ref> and use low frequency (<1000&nbsp;Hz) sounds to communicate while traveling, feeding, and socializing. Intense calls for communication and navigation are produced especially during migration season. During breeding season, bowheads make long, complex, variable songs for mating calls.<ref name= "Finely, K.J. 2001">{{cite journal |author=Finley, K.J. |date=2001 |title= Natural History and Conservation of the Greenland Whale, or Bowhead, in the Northeast Atlantic |journal=Arctic Institute of North America |volume=54 |issue=1 |pages=55–76 |doi=10.14430/arctic764}}</ref>
+Bowhead whales are highly vocal<ref>{{cite web|url=http://cetus.ucsd.edu/voicesinthesea_org/species/baleenWhales/bowhead.html|title=Bowhead Whale : Baleen Whales : Voices in the Sea|publisher=}}</ref> and use low frequency (<1000&nbsp;Hz) sounds to communicate while traveling, feeding, and socializing. Intense calls for communication and navigation are produced especially during migration season. During breeding season, bowheads make long, complex, variable songs for mating calls.<ref name="Finely, K.J. 2001">{{cite journal |author=Finley, K.J. |date=2001 |title= Natural History and Conservation of the Greenland Whale, or Bowhead, in the Northeast Atlantic |journal=Arctic Institute of North America |volume=54 |issue=1 |pages=55–76 |doi=10.14430/arctic764}}</ref>
-===Reproduction===
+=== Reproduction ===
 {{See also|Whale reproduction}}
-Sexual activity occurs between pairs and in boisterous groups of several males and one or two females. Breeding season is observed from March through August; conception is believed to occur primarily in March when song activity is at its highest.<ref name= "Finely, K.J. 2001" /> Reproduction can begin when a whale is 10 to 15 years old. The [[gestation]] period is 13–14 months with females producing a calf once every three to four years.<ref name="RepKoski"/> [[Lactation]] typically lasts about a year. To survive in the cold water immediately after birth, calves are born with a thick layer of [[blubber]]. Within 30 minutes of birth, bowhead calves are able to swim on their own. A newborn calf is typically {{convert|4|-|4.5|m|abbr=on}} long, weighs approximately {{convert|1000|kg|abbr=on}}, and grows to {{convert|8.2|m|abbr=on}} within the first year.<ref name="RepKoski"/>
+Sexual activity occurs between pairs and in boisterous groups of several males and one or two females. Breeding season is observed from March through August; conception is believed to occur primarily in March when song activity is at its highest.<ref name="Finely, K.J. 2001" /> Reproduction can begin when a whale is 10 to 15 years old. The [[gestation]] period is 13–14 months with females producing a calf once every three to four years.<ref name="RepKoski" /> [[Lactation]] typically lasts about a year. To survive in the cold water immediately after birth, calves are born with a thick layer of [[blubber]]. Within 30 minutes of birth, bowhead calves are able to swim on their own. A newborn calf is typically {{convert|4|-|4.5|m|abbr=on}} long, weighs approximately {{convert|1000|kg|abbr=on}}, and grows to {{convert|8.2|m|abbr=on}} within the first year.<ref name="RepKoski" />
-==Health==
+== Health ==
-===Lifespan===
+=== Lifespan ===
 Bowhead whales are known to be among the [[longest-living animals|longest-living mammals]], living for over 200 years.<ref>{{cite journal |author1=Schiffman, Joshua D. |author2=Breen, Matthew |date=2015 |title=Comparative oncology: what dogs and other species can teach us about humans with cancer |journal=Phil. Trans. R. Soc. B |volume=370 |issue=1673 |pages=1–13 |doi=10.1098/rstb.2014.0231}}</ref> In May 2007, a {{convert|15|m|abbr=on}} specimen caught off the [[Alaska]]n coast was discovered with the head of an explosive [[harpoon]] embedded deep under its neck blubber. The {{convert|3.5|in|abbr=on}} arrow-shaped projectile was manufactured in [[New Bedford, Massachusetts|New Bedford]], [[Massachusetts]], a major whaling center, around 1890, suggesting the animal may have survived a similar hunt more than a century ago.<ref>{{cite journal |author1=George, John C. |author2=Bada, Jeffrey |author3=Zeh, Judith |author4=Scott, Laura |author5=Brown, Stephen E. |author6=O'Hara, Todd |author7=Suydam, Robert |lastauthoramp=yes |date=1999 |title=Age and growth estimates of bowhead whales (''Balaena mysticetus'') via aspartic acid racemization |journal=Can. J. Zool. |volume=77 |issue=4 |pages=571–580 |doi=10.1139/z99-015}}</ref><ref>Conroy, Erin. (6 December 2007) [http://www.msnbc.msn.com/id/19195624/ Netted whale hit by lance a century ago]. MSNBC.</ref><ref>[http://usatoday30.usatoday.com/news/nation/2007-06-12-century-old-whale_N.htm 19th-century weapon found in whale]. Associated Press via USA Today. 12 June 2007</ref> This whale was estimated to be 211 years old.<ref>{{cite web |title=Can Marine Biology Help Us Live Forever? Bowhead Whale Can Live 200 Years, Is Cancer Resistant |url=http://www.medicaldaily.com/can-marine-biology-help-us-live-forever-bowhead-whale-can-live-200-years-cancer-316424 |date=6 January 2015 |work=Medical Daily |accessdate=8 November 2015}}</ref> Other bowhead whales found on the whaling expedition were estimated to be between 135 and 172 years old. This discovery showed the [[longevity]] of the bowhead whale is much greater than originally thought.
-===Genetic causes===
+=== Genetic causes ===
-It was previously believed the more cells present in an organism, the greater the chances of [[mutations]] that cause age related diseases and [[cancer]].<ref name= "Washington Post 2015">{{cite web |title=Researchers hope this whale’s genes will help reverse human aging |url=https://www.washingtonpost.com/news/speaking-of-science/wp/2015/01/06/researchers-hope-this-whales-genes-will-help-reverse-human-aging/ |date=6 January 2015 |work=The Washington Post |accessdate=8 November 2015}}</ref> Although the bowhead whale has thousands of times more cells than other mammals, the whale has a much higher resistance to cancer and aging. In 2015, scientists from the US and UK were able to successfully map the whale's [[genome]].<ref>{{cite web |title=Scientists map bowhead whale’s genome; discover genes responsible for long life |url=http://www.techienews.co.uk/9722127/scientists-map-bowhead-whales-genome-discover-genes-responsible-long-life/ |date=5 January 2015 |work=Technie News |accessdate=5 January 2015}}</ref> Through comparative analysis, two [[alleles]] that could be responsible for the whale's longevity were identified. These two specific gene mutations linked to the bowhead whale's ability to live longer are the [[ERCC1]] gene and the proliferating cell nuclear antigen ([[PCNA]]) gene. [[ERCC1]] is linked to [[DNA repair]] as well as increased cancer resistance. [[PCNA]] is also important in DNA repair. These mutations enable bowhead whales to better repair DNA damage, allowing for greater resistance to cancer.<ref name= "Washington Post 2015"/> The whale's genome may also reveal physiological adaptations such as having low metabolic rates compared to other mammals.<ref>{{cite web |title=The bowhead whale lives over 200 years. Can its genes tell us why? |url=http://www.sciencedaily.com/releases/2015/01/150105101421.htm |date=5 January 2015 |work=Science Daily |accessdate=8 November 2015}}</ref> Changes in the gene [[UCP1]], a gene involved in [[thermoregulation]], can explain differences in the metabolic rates in cells.
+It was previously believed the more cells present in an organism, the greater the chances of [[mutations]] that cause age related diseases and [[cancer]].<ref name="Washington Post 2015">{{cite web |title=Researchers hope this whale’s genes will help reverse human aging |url=https://www.washingtonpost.com/news/speaking-of-science/wp/2015/01/06/researchers-hope-this-whales-genes-will-help-reverse-human-aging/ |date=6 January 2015 |work=The Washington Post |accessdate=8 November 2015}}</ref> Although the bowhead whale has thousands of times more cells than other mammals, the whale has a much higher resistance to cancer and aging. In 2015, scientists from the US and UK were able to successfully map the whale's [[genome]].<ref>{{cite web |title=Scientists map bowhead whale’s genome; discover genes responsible for long life |url=http://www.techienews.co.uk/9722127/scientists-map-bowhead-whales-genome-discover-genes-responsible-long-life/ |date=5 January 2015 |work=Technie News |accessdate=5 January 2015}}</ref> Through comparative analysis, two [[alleles]] that could be responsible for the whale's longevity were identified. These two specific gene mutations linked to the bowhead whale's ability to live longer are the [[ERCC1]] gene and the proliferating cell nuclear antigen ([[PCNA]]) gene. [[ERCC1]] is linked to [[DNA repair]] as well as increased cancer resistance. [[PCNA]] is also important in DNA repair. These mutations enable bowhead whales to better repair DNA damage, allowing for greater resistance to cancer.<ref name="Washington Post 2015" /> The whale's genome may also reveal physiological adaptations such as having low metabolic rates compared to other mammals.<ref>{{cite web |title=The bowhead whale lives over 200 years. Can its genes tell us why? |url=http://www.sciencedaily.com/releases/2015/01/150105101421.htm |date=5 January 2015 |work=Science Daily |accessdate=8 November 2015}}</ref> Changes in the gene [[UCP1]], a gene involved in [[thermoregulation]], can explain differences in the metabolic rates in cells.
-==Ecology==
+== Ecology ==
 [[File:Balaena mysticetus.jpg|thumb|Drawing of an adult in 1884]]
-===Range and habitat===
+=== Range and habitat ===
 The bowhead whale is the only [[baleen whale]] to spend its entire life in the [[Arctic]] and sub-Arctic waters.<ref name="DisMovMoo">Moore, S. E., and R. R. Reeves (1993). "Distribution and Movement". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref> The [[Alaska]]n population spends the winter months in the southwestern [[Bering Sea]]. The group migrates northward in the spring, following openings in the ice, into the [[Chukchi Sea|Chukchi]] and [[Beaufort Sea|Beaufort]] seas.<ref>{{cite book |last1=Smultea |first1=M. |last2=Fertl |first2=D. |last3=Rugh |first3=D. J. |last4=Bacon |first4=C. E. |date=2012 |title=Summary of systematic bowhead surveys conducted in the U.S. Beaufort and Chukchi Seas, 1975–2009 |publisher=U.S. Dep. Commer., NOAA Tech. Memo. NMFS-AFSC-237 |page=48 |url=http://purl.fdlp.gov/GPO/gpo29638}}</ref> It has been confirmed the whale's range varies depending on climate changes and on the forming/melting of ice.<ref>{{Cite journal | doi = 10.1038/ncomms2714| pmid = 23575681| title = Ancient DNA reveals that bowhead whale lineages survived Late Pleistocene climate change and habitat shifts| journal = Nature Communications| volume = 4| pages = 1677| year = 2013| last1 = Foote | first1 = A. D. | last2 = Kaschner | first2 = K. | last3 = Schultze | first3 = S. E. | last4 = Garilao | first4 = C. | last5 = Ho | first5 = S. Y. W. | last6 = Post | first6 = K. | last7 = Higham | first7 = T. F. G. | last8 = Stokowska | first8 = C. | last9 = Van Der Es | first9 = H. | last10 = Embling | first10 = C. B. | last11 = Gregersen | first11 = K. | last12 = Johansson | first12 = F. | last13 = Willerslev | first13 = E. | last14 = Gilbert | first14 = M. T. P. | bibcode = 2013NatCo...4E1677F}}</ref>
-===Population===
+=== Population ===
-The bowhead population around [[Alaska]] has increased since commercial whaling ceased. [[Alaska Natives]] continue to kill small numbers in [[Subsistence economy|subsistence hunt]]s each year. This level of killing (25–40 animals annually) is not expected to affect the population's recovery. The population off Alaska's coast (the Bering-Chukchi-Beaufort stock) appears to be recovering and was at about 10,500 animals as of 2001. The status of other populations is less well known. There were about 1,200 off West Greenland in 2006, while the [[Svalbard]] population may only number in the tens. However, the numbers have been increasing in recent years.<ref name="Bowhead whale Balaena mysticetus">[[Norwegian Polar Institute]]. [http://www.npolar.no/en/species/bowhead-whale.html Bowhead whale (''Balaena mysticetus'')]. npolar.no</ref>
+The bowhead population around [[Alaska]] has increased since commercial whaling ceased. [[Alaska Natives]] continue to kill small numbers in [[Subsistence economy|subsistence hunts]] each year. This level of killing (25–40 animals annually) is not expected to affect the population's recovery. The population off Alaska's coast (the Bering-Chukchi-Beaufort stock) appears to be recovering and was at about 10,500 animals as of 2001. The status of other populations is less well known. There were about 1,200 off West Greenland in 2006, while the [[Svalbard]] population may only number in the tens. However, the numbers have been increasing in recent years.<ref name="Bowhead whale Balaena mysticetus">[[Norwegian Polar Institute]]. [http://www.npolar.no/en/species/bowhead-whale.html Bowhead whale (''Balaena mysticetus'')]. npolar.no</ref>
 In March 2008, Canada's Department of Fisheries and Oceans stated the previous estimates in the eastern Arctic had under-counted, with a new estimate of 14,400 animals (range 4,800–43,000).<ref>[http://www.cbc.ca/news/technology/eastern-arctic-bowhead-whales-not-threatened-after-all-government-says-1.717576 Eastern Arctic bowhead whales not threatened]. Cbc.ca (16 April 2008). Retrieved on 2011-09-15.</ref> These larger numbers correspond to prewhaling estimates, indicating the population has fully recovered. However, if climate change substantially shrinks sea ice, these whales could be threatened by increased shipping traffic.<ref>Laidre, Kristin (22 January 2009) [https://web.archive.org/web/20090510234455/http://www.nwfsc.noaa.gov/events/monsterdocs/win2009msjall01.22.09.pdf "Foraging Ecology of Bowhead Whales in West Greenland."] Monster Jam. Northwest Fisheries Science Center, Seattle.</ref>
-====Hudson Bay to Foxe Basin====
+==== Hudson Bay to Foxe Basin ====
 [[File:The Canadian field-naturalist (1997) (20330377360).jpg|thumb|Sighting locations by researchers and hunters]]
-[[Hudson Bay]] – [[Foxe Basin]] population is distinct from the [[Baffin Bay]]{{snds}}[[Davis Strait]] group.<ref name="link.springer.com">{{cite journal | url = http://link.springer.com/chapter/10.1007%2F978-90-481-9121-5_8#page-1 | doi=10.1007/978-90-481-9121-5_8 | title=Past, Present, and Future for Bowhead Whales (Balaena mysticetus) in Northwest Hudson Bay | journal=A Little Less Arctic | pages=159–177}}</ref> Original population size of this local group is unclear, but possibly around 500 to 600 whales annually summered in the northwestern part of the bay in 1860s.<ref>http://www.registrelep-sararegistry.gc.ca/species/speciesDetails_e.cfm?sid=131</ref> Likely, the number of whales actually inhabit within Hudson Bay is much smaller than the total population size of this group,<ref name=NWMB>http://www.nwmb.com/iku/2013-11-09-01-41-51/2013-11-09-01-46-43/2008/mar-06-2008-level-of-tah-for-bowhead-whales/551-tab15-df0-1999/file</ref> and despite current population size is rather unclear, reports from local indigenous people indicate this population is at least increasing over decades.<ref name=sararegistry/> Larger portions of usages of the bay is considered to be summering while wintering is on smaller scale where some animals winter in [[Hudson Strait]] most notably north of [[Igloolik Island]] and northeastern Hudson Bay. Distribution patterns of whales in this regions are largely affected by presences of [[killer whale]]s and bowheads can disappear from normal ranges due to recent changes in killer whales' occurrences within the bay possibly because of changes in movements of ice floes by [[climate change|changing climate]].<ref name=sararegistry>http://www.sararegistry.gc.ca/virtual_sara/files/cosewic/sr_bowhead_whale_e.pdf</ref> Whaling grounds in 19th century covered from [[Marble Island]] to [[Roes Welcome Sound]] and to
+[[Hudson Bay]] – [[Foxe Basin]] population is distinct from the [[Baffin Bay]]{{snds}}[[Davis Strait]] group.<ref name="link.springer.com">{{cite journal | url = http://link.springer.com/chapter/10.1007%2F978-90-481-9121-5_8#page-1 | doi=10.1007/978-90-481-9121-5_8 | title=Past, Present, and Future for Bowhead Whales (Balaena mysticetus) in Northwest Hudson Bay | journal=A Little Less Arctic | pages=159–177}}</ref> Original population size of this local group is unclear, but possibly around 500 to 600 whales annually summered in the northwestern part of the bay in 1860s.<ref>http://www.registrelep-sararegistry.gc.ca/species/speciesDetails_e.cfm?sid=131</ref> Likely, the number of whales actually inhabit within Hudson Bay is much smaller than the total population size of this group,<ref name=NWMB>http://www.nwmb.com/iku/2013-11-09-01-41-51/2013-11-09-01-46-43/2008/mar-06-2008-level-of-tah-for-bowhead-whales/551-tab15-df0-1999/file</ref> and despite current population size is rather unclear, reports from local indigenous people indicate this population is at least increasing over decades.<ref name=sararegistry /> Larger portions of usages of the bay is considered to be summering while wintering is on smaller scale where some animals winter in [[Hudson Strait]] most notably north of [[Igloolik Island]] and northeastern Hudson Bay. Distribution patterns of whales in this regions are largely affected by presences of [[killer whale]]s and bowheads can disappear from normal ranges due to recent changes in killer whales' occurrences within the bay possibly because of changes in movements of ice floes by [[climate change|changing climate]].<ref name=sararegistry>http://www.sararegistry.gc.ca/virtual_sara/files/cosewic/sr_bowhead_whale_e.pdf</ref> Whaling grounds in 19th century covered from [[Marble Island]] to [[Roes Welcome Sound]] and to
 [[Lyon Inlet]] and [[Fisher Strait]], and whales still migrate through most of these areas.
-Mostly, distributions within Hudson Bay is restricted in northwestern part<ref name="link.springer.com"/> along with [[Repulse Bay]], [[Frozen Strait]], northern Foxe
+Mostly, distributions within Hudson Bay is restricted in northwestern part<ref name="link.springer.com" /> along with [[Repulse Bay]], [[Frozen Strait]], northern Foxe
-Basin, and north of [[Igloolik]] in summer,<ref name=sararegistry/> and satellite tracking<ref>http://arctic.blogs.panda.org/field/june-2014-bowheads-and-breaking-ice/</ref> indicates that some portions of the group within the bay do not venture further south than areas south of [[Coats Island|Coasts]] and [[Mansel Island]]s.{{citation needed|date=April 2016}} Cow - calf pairs and juveniles up to {{convert|13.5|m|abbr=on}} in length consist of majority of summering aggregation in northern Foxe Basin while matured males and non-calving females may utilize northwestern part of Hudson Bay.<ref name=sararegistry/> Fewer whales also migrate to west coast of Hudson Bay, [[Mansel Island|Mansel]] and [[Ottawa Islands]].<ref name=sararegistry/> Bowhead ranges within Hudson Bay are usually considered not to cover southern parts,<ref name=NWMB /><ref>http://www.dfo-mpo.gc.ca/science/coe-cde/cemam/themes/dynami-fra.htm</ref> but at least some whales migrate into further south such as at [[Sanikiluaq]]{{citation needed|date=April 2016}} and [[Churchill River (Hudson Bay)|Churchill river mouth]].<ref>http://pubs.aina.ucalgary.ca/arctic/Arctic36-1-5.pdf</ref><ref>http://churchillpolarbears.org/2015/07/bowhead-whale-in-churchill-waters/</ref>
+Basin, and north of [[Igloolik]] in summer,<ref name=sararegistry /> and satellite tracking<ref>{{cite web|url=http://arctic.blogs.panda.org/field/june-2014-bowheads-and-breaking-ice/|title=June 2014: Bowheads and breaking ice – Thin Ice Blog|first=|last=WWF|publisher=}}</ref> indicates that some portions of the group within the bay do not venture further south than areas south of [[Coats Island|Coasts]] and [[Mansel Island]]s.{{citation needed|date=April 2016}} Cow – calf pairs and juveniles up to {{convert|13.5|m|abbr=on}} in length consist of majority of summering aggregation in northern Foxe Basin while matured males and non-calving females may utilize northwestern part of Hudson Bay.<ref name=sararegistry /> Fewer whales also migrate to west coast of Hudson Bay, [[Mansel Island|Mansel]] and [[Ottawa Islands]].<ref name=sararegistry /> Bowhead ranges within Hudson Bay are usually considered not to cover southern parts,<ref name=NWMB /><ref>{{cite web|url=http://www.dfo-mpo.gc.ca/science/coe-cde/cemam/themes/dynami-fra.htm|title=MPO – Recherche sur les mammifères marins au Canada – MPO Sciences|publisher=}}</ref> but at least some whales migrate into further south such as at [[Sanikiluaq]]{{citation needed|date=April 2016}} and [[Churchill River (Hudson Bay)|Churchill river mouth]].<ref>http://pubs.aina.ucalgary.ca/arctic/Arctic36-1-5.pdf</ref><ref>{{cite web|url=http://churchillpolarbears.org/2015/07/bowhead-whale-in-churchill-waters/|title=Bowhead Whale in Churchill Waters – Churchill Polar Bears|publisher=}}</ref>
-Congregation within Foxe Basin occurs in a well-defined area at {{convert|3,700|km|abbr=on}} north of Igloolik Island to [[Fury and Hecla Strait]] and [[Jens Munk Island]] and [[wikt:sv:Gifford Fiord|Gifford Fiord]], and into [[Gulf of Boothia]] and [[Prince Regent Inlet]]. Northward migrating along western Foxe Basin to eastern side of the basin also occurs in spring seasons.<ref name=sararegistry/>
+Congregation within Foxe Basin occurs in a well-defined area at {{convert|3,700|km|abbr=on}} north of Igloolik Island to [[Fury and Hecla Strait]] and [[Jens Munk Island]] and [[wikt:sv:Gifford Fiord|Gifford Fiord]], and into [[Gulf of Boothia]] and [[Prince Regent Inlet]]. Northward migrating along western Foxe Basin to eastern side of the basin also occurs in spring seasons.<ref name=sararegistry />
-[[File:A bowhead whale spyhops off the coast north western Sea of Okhotsk by Olga Shpak, Marine Mammal Council, IEE RAS.jpg|thumb|Whale spyhops in Ulbansky Bay, northwestern [[Okhotsk Sea]].<ref name="Olga Shpak. 2014"/>]]
+[[File:A bowhead whale spyhops off the coast north western Sea of Okhotsk by Olga Shpak, Marine Mammal Council, IEE RAS.jpg|thumb|Whale spyhops in Ulbansky Bay, northwestern [[Okhotsk Sea]].<ref name="Olga Shpak. 2014" />]]
-====Sea of Okhotsk====
+==== Sea of Okhotsk ====
 Not much is known about the endangered [[Sea of Okhotsk]] population. To learn more about the population, these mammals have been regularly observed near the [[Shantar Islands]], very close to the shore, such as at [[Ongachan Bay]].<ref>{{cite web|url=http://www.arcticexpedition.ru/?tourname=place-power9250&photogallery=1&direction=FarEast-NearSea&powerplace=place-power92 |title=Тур 'наблюдение за китами и плавание вдоль побережья Охотского моря и на Шантарските острова' |publisher=Arcticexpedition.ru |date=15 August 2000 |accessdate=19 October 2012 |language=ru}}</ref><ref>{{cite web|url=http://www.turizmvnn.ru/cont/show/14672/ |title=Фотография: Киты подходят совсем близко к берегу |language=ru |publisher=Turizmvnn.ru |date=16 April 2011 |accessdate=19 October 2012}}</ref> Several companies provide [[whale watching]] services which are mostly land-based. According to Russian scientists, this total population likely does not exceed 400 animals.<ref name="Olga Shpak. 2014">{{cite web |language=ru |last=Shpak |first=Olga |date=19 February 2014 |url=http://ria.ru/earth/20140219/995484804.html |title=Второе рождение гренландского кита |publisher=[[RIA Novosti]] |accessdate=4 June 2014}}</ref> Scientific research on this population was seldom done before 2009, when researchers studying [[Beluga whale|belugas]] noticed concentrations of bowheads in the study area. Thus, bowheads in the Sea of Okhotsk were once called "forgotten whales" by researchers. With support from [[World Wide Fund for Nature|WWF]], Russian scientists and nature conservationists cooperated to create a cetacean sanctuary in the [[Magadan]] region. This region covers vast areas of the northwestern Sea of Okhotsk, including the Shantar regions.<ref>{{cite web |date=31 December 2013 |url=http://ria.ru/hab/20131231/987503119.html |title=WWF приветствует создание нацпарка в Хабаровском крае |language=ru |publisher=[[RIA Novosti]] |accessdate=4 June 2014}}</ref>
-Possibly, vagrants from this population occasionally reach into Asian nations such as off Japan or [[Korean Peninsula]] (although this record might or might not be of a [[right whale]]<ref>http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0034905#pone-0034905-g002</ref>). First documented report of the species in Japanese waters was of a strayed infant ({{convert|7|m|abbr=on|disp=sqbr}}) caught in [[Osaka Bay]] on 23 June 1969,<ref>http://osakagyoren.or.jp/naniwa/</ref> and the first living sighting was of a {{convert|10|m|abbr=on}} juvenile around [[Shiretoko Peninsula]] (the southernmost of [[ice stream|ice floe]] range in the world) on 21 to 23 June 2015.<ref>http://www.asahi.com/articles/ASH6R5VQ2H6RIIPE026.html</ref> Fossils have been excavated on [[Hokkaido]],<ref>http://www.city.ishikari.hokkaido.jp/soshiki/bunkazaih/2785.html</ref> but it is unclear whether or not northern coasts of Japan once had been included in seasonal or occasional migration ranges.
+Possibly, vagrants from this population occasionally reach into Asian nations such as off Japan or [[Korean Peninsula]] (although this record might or might not be of a [[right whale]]<ref>{{cite journal|url=http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0034905|title=Spatial and Seasonal Distribution of American Whaling and Whales in the Age of Sail|first1=Tim D.|last1=Smith|first2=Randall R.|last2=Reeves|first3=Elizabeth A.|last3=Josephson|first4=Judith N.|last4=Lund|date=27 April 2012|publisher=|journal=PLOS ONE|volume=7|issue=4|pages=e34905|via=PLoS Journals|doi=10.1371/journal.pone.0034905|pmid=22558102|pmc=3338773}}</ref>). First documented report of the species in Japanese waters was of a strayed infant ({{convert|7|m|abbr=on|disp=sqbr}}) caught in [[Osaka Bay]] on 23 June 1969,<ref>{{cite web|url=http://osakagyoren.or.jp/naniwa/|title=大阪府漁業協同組合連合会|publisher=}}</ref> and the first living sighting was of a {{convert|10|m|abbr=on}} juvenile around [[Shiretoko Peninsula]] (the southernmost of [[ice stream|ice floe]] range in the world) on 21 to 23 June 2015.<ref>http://www.asahi.com/articles/ASH6R5VQ2H6RIIPE026.html</ref> Fossils have been excavated on [[Hokkaido]],<ref>{{cite web|url=http://www.city.ishikari.hokkaido.jp/soshiki/bunkazaih/2785.html|title=いしかり博物誌／第5回 - 北海道石狩市公式ホームページ|publisher=}}</ref> but it is unclear whether or not northern coasts of Japan once had been included in seasonal or occasional migration ranges.
-Genetic studies suggest Okhotsk population share common ancestry with whales in Bering-Chukchi-Beaufort Seas, and repeated mixings had occurred between whales in two seas.<ref>[http://oaktrust.library.tamu.edu/handle/1969.1/ETD-TAMU-2002-THESIS-M33 Occurrence, behavior and genetic diversity of bowhead whales in the Western Sea of Okhotsk, Russia]</ref>
+Genetic studies suggest Okhotsk population share common ancestry with whales in Bering-Chukchi-Beaufort Seas, and repeated mixings had occurred between whales in two seas.<ref>{{cite web|url=http://oaktrust.library.tamu.edu/handle/1969.1/ETD-TAMU-2002-THESIS-M33|title=Occurrence, behavior and genetic diversity of bowhead whales in the Western Sea of Okhotsk, Russia|first=MacLean, Stephen|last=Ahgeak|date=1 January 2002|publisher=}}</ref>
-====Russian Arctic====
+==== Russian Arctic ====
-{{see also|Russian Arctic National Park}}
+{{See also|Russian Arctic National Park}}
-The most endangered and historically the largest of all bowhead populations is the Svalbard/Spitsbergen population.<ref>Gross A., 2010 [http://qsr2010.ospar.org/media/assessments/Species/P00494_Bowhead_whale.pdf Background Document for Bowhead whale Balaena mysticetus]. The [[OSPAR Convention]] and [[Musée des Matériaux du Centre de Recherche sur les Monuments Historiques]]. ISBN 978-1-907390-35-7. retrieved on 24 May 2014</ref> Occurring normally in [[Fram Strait]],<ref>Kovacs M.K., [http://mosj.npolar.no/en/fauna/marine/indicators/bowhead_whale.html? Bowhead whale (''Balaena mysticetus'')]. Environmental Monitoring of Svalbard and Jan Mayen. retrieved on 27 May 2014</ref> [[Barents Sea]] and [[Severnaya Zemlya]] along [[Kara Sea]]<ref name="Bowhead whale Balaena mysticetus"/> to [[Laptev Sea]] and [[East Siberian Sea]] regions, these whales were seen in entire coastal regions in European and Russian Arctic, even reaching to [[Iceland]]ic and [[Norway|Scandinavian]] coasts and [[Jan Mayen]] in [[Greenland Sea]], and west of [[Cape Farewell, Greenland|Cape Farewell]] and western [[Greenland]] coasts.<ref>{{cite journal|author1=Gilg O.|author2= Born W.E.|year= 2004 |url=http://www.researchgate.net/publication/226243275_Recent_sightings_of_the_bowhead_whale_(Balaena_mysticetus)_in_Northeast_Greenland_and_the_Greenland_Sea |title=Recent sightings of the bowhead whale (''Balaena mysticetus'') in Northeast Greenland and the Greenland Sea|journal= Polar Biology|volume= 28|issue= 10|pages= 796–801|doi=10.1007/s00300-005-0001-9}}</ref> Also, bowheads in this stock were possibly once abundant in areas adjacent to the [[White Sea]] region, where few or no animals currently migrate, such as the [[Kola Peninsula|Kola]] and [[Kanin Peninsula]]. Today, the number of sightings in elsewhere are very small,<ref>Ritchie B. (June 2013) [https://vimeo.com/83027638 Arctic shorts – bowhead whale]. [[Vimeo]]. Retrieved 2 June 2014</ref> but with increasing regularities<ref>Sala E., 2013 [http://newswatch.nationalgeographic.com/2013/09/11/franz-josef-land-expedition-first-look-at-post-expedition-discoveries/ Franz Josef Land Expedition: First Look at Post-Expedition Discoveries]. Pristine Seas Expeditions. [[National Geographic Channel|National Geographic]]. retrieved on 24 May 2014</ref> with whales having strong regional connections.<ref>WIIG Ø., Bachmann L., Janik M.V., Kovacs M.K., Lydersen C., 2007. [http://www.researchgate.net/publication/249474046_SPITSBERGEN_BOWHEAD_WHALES_REVISITED/file/504635298afa4eb130.pdf Spitsbergen Bowhead Whales Revisited]. [[Society for Marine Mammalogy]]. Retrieved 24 May 2014</ref> Whales have also started approaching townships and inhabited areas such as around [[Longyearbyen]].<ref>Johannessen, R. (19 October 2011) [http://www.aftenposten.no/nyheter/iriks/--Dette-er-en-sensasjon-6407203.html#.U4QNo-lZrIU Dette er en sensasjon!]. The [[Aftenposten]]. retrieved on 27 May 2014</ref> The waters around the marine mammal sanctuary<ref>Nefedova T., Gavrilo M., Gorshkov S., 2013. [http://www.rgo.ru/ru/article/letom-v-arktike-stalo-menshe-lda Летом в Арктике стало меньше льда]. Russian Geographical Society. retrieved on 24 May 2014</ref> of [[Franz Josef Land]] is possibly functioning as the most important habitat for this population.<ref>European Cetacean Society. [http://liege.europeancetaceansociety.eu/content/bowhead-whales-balaena-mysticetus-sighting-franz-josef-land-area Bowhead whales (''Balaena mysticetus'') sighting in the Franz Josef Land area.]. retrieved on 24 May 2014</ref><ref>Scalini I. (19 February 2014) [http://web.archive.org/web/20141129204614/http://www.rus-arc.ru/ru/News/Details/e155b1cd-4f34-436f-979c-7e0a36cc5821 Всемирный день китов]. Russian Arctic National Park. retrieved on 24 May 2014</ref>
+The most endangered and historically the largest of all bowhead populations is the Svalbard/Spitsbergen population.<ref>Gross A., 2010 [http://qsr2010.ospar.org/media/assessments/Species/P00494_Bowhead_whale.pdf Background Document for Bowhead whale Balaena mysticetus]. The [[OSPAR Convention]] and [[Musée des Matériaux du Centre de Recherche sur les Monuments Historiques]]. ISBN 978-1-907390-35-7. retrieved on 24 May 2014</ref> Occurring normally in [[Fram Strait]],<ref>Kovacs M.K., [http://mosj.npolar.no/en/fauna/marine/indicators/bowhead_whale.html? Bowhead whale (''Balaena mysticetus'')]. Environmental Monitoring of Svalbard and Jan Mayen. retrieved on 27 May 2014</ref> [[Barents Sea]] and [[Severnaya Zemlya]] along [[Kara Sea]]<ref name="Bowhead whale Balaena mysticetus" /> to [[Laptev Sea]] and [[East Siberian Sea]] regions, these whales were seen in entire coastal regions in European and Russian Arctic, even reaching to [[Iceland]]ic and [[Norway|Scandinavian]] coasts and [[Jan Mayen]] in [[Greenland Sea]], and west of [[Cape Farewell, Greenland|Cape Farewell]] and western [[Greenland]] coasts.<ref>{{cite journal|author1=Gilg O.|author2= Born W.E.|year= 2004 |url=http://www.researchgate.net/publication/226243275_Recent_sightings_of_the_bowhead_whale_(Balaena_mysticetus)_in_Northeast_Greenland_and_the_Greenland_Sea |title=Recent sightings of the bowhead whale (''Balaena mysticetus'') in Northeast Greenland and the Greenland Sea|journal= Polar Biology|volume= 28|issue= 10|pages= 796–801|doi=10.1007/s00300-005-0001-9}}</ref> Also, bowheads in this stock were possibly once abundant in areas adjacent to the [[White Sea]] region, where few or no animals currently migrate, such as the [[Kola Peninsula|Kola]] and [[Kanin Peninsula]]. Today, the number of sightings in elsewhere are very small,<ref>Ritchie B. (June 2013) [https://vimeo.com/83027638 Arctic shorts – bowhead whale]. [[Vimeo]]. Retrieved 2 June 2014</ref> but with increasing regularities<ref>Sala E., 2013 [http://newswatch.nationalgeographic.com/2013/09/11/franz-josef-land-expedition-first-look-at-post-expedition-discoveries/ Franz Josef Land Expedition: First Look at Post-Expedition Discoveries]. Pristine Seas Expeditions. [[National Geographic Channel|National Geographic]]. retrieved on 24 May 2014</ref> with whales having strong regional connections.<ref>WIIG Ø., Bachmann L., Janik M.V., Kovacs M.K., Lydersen C., 2007. [http://www.researchgate.net/publication/249474046_SPITSBERGEN_BOWHEAD_WHALES_REVISITED/file/504635298afa4eb130.pdf Spitsbergen Bowhead Whales Revisited]. [[Society for Marine Mammalogy]]. Retrieved 24 May 2014</ref> Whales have also started approaching townships and inhabited areas such as around [[Longyearbyen]].<ref>Johannessen, R. (19 October 2011) [http://www.aftenposten.no/nyheter/iriks/--Dette-er-en-sensasjon-6407203.html#.U4QNo-lZrIU Dette er en sensasjon!]. The [[Aftenposten]]. retrieved on 27 May 2014</ref> The waters around the marine mammal sanctuary<ref>Nefedova T., Gavrilo M., Gorshkov S., 2013. [http://www.rgo.ru/ru/article/letom-v-arktike-stalo-menshe-lda Летом в Арктике стало меньше льда]. Russian Geographical Society. retrieved on 24 May 2014</ref> of [[Franz Josef Land]] is possibly functioning as the most important habitat for this population.<ref>European Cetacean Society. [http://liege.europeancetaceansociety.eu/content/bowhead-whales-balaena-mysticetus-sighting-franz-josef-land-area Bowhead whales (''Balaena mysticetus'') sighting in the Franz Josef Land area.]. retrieved on 24 May 2014</ref><ref>Scalini I. (19 February 2014) [http://web.archive.org/web/20141129204614/http://www.rus-arc.ru/ru/News/Details/e155b1cd-4f34-436f-979c-7e0a36cc5821 Всемирный день китов]. Russian Arctic National Park. retrieved on 24 May 2014</ref>
-Current status of population structure of this stock is unclear; whether they are remnant of the historic Svalbard group, re-colonized individuals from other stocks, or if a mixing of these two or more stocks had taken place. In 2015, discoveries of the refuge along eastern Greenland where whaling ships could not reach due to ice floes<ref>http://www.bbc.com/earth/story/20150721-secret-whale-refuge-discovered</ref> and largest numbers of whales (80–100 individuals) ever sighted between Spitsbergen and Greenland<ref>https://oceanwide-expeditions.com/blog/exceptional-sighting-of-80-bowhead-whales</ref> indicate that more whales than previously considered survived whaling periods, and flows from the other populations are possible.
+Current status of population structure of this stock is unclear; whether they are remnant of the historic Svalbard group, re-colonized individuals from other stocks, or if a mixing of these two or more stocks had taken place. In 2015, discoveries of the refuge along eastern Greenland where whaling ships could not reach due to ice floes<ref>{{cite web|url=http://www.bbc.com/earth/story/20150721-secret-whale-refuge-discovered|title=Secret whale refuge discovered|first=Matt|last=Walker|publisher=}}</ref> and largest numbers of whales (80–100 individuals) ever sighted between Spitsbergen and Greenland<ref>{{cite web|url=https://oceanwide-expeditions.com/blog/exceptional-sighting-of-80-bowhead-whales|title=Exceptional sighting of 80+ Bowhead Whales|publisher=}}</ref> indicate that more whales than previously considered survived whaling periods, and flows from the other populations are possible.
-====Possible moulting area on Baffin Island====
+==== Possible moulting area on Baffin Island ====
 During expeditions by a tour operator 'Arctic Kingdom', a large group of bowheads seemingly involved in courtship activities were discovered in very shallow bays in south of [[Qikiqtarjuaq]] in 2012.<ref>Lennartz T. (29 April 2013) [http://arctickingdom.com/new-bowhead-whale-molting-location-found/ New Bowhead Whale Molting Location Found]. Arctic Kingdom. retrieved on 9 June 2014</ref> Floating skins and rubbing behaviors at sea bottom indicated possible [[moulting]] had taken place. Moulting behaviors had never or had seldomly been documented for this species before. This area is an important habitat for whales that were observed to be relatively active and to interact with humans positively, or to rest on sea floors. These whales belong to [[Davis Strait]] stock.
 Isabella Bay in [[List of protected areas of Nunavut|Niginganiq National Wildlife Area]] is the first wildlife sanctuary in the world to be designed specially for bowhead whales. However, moultings have not been recorded in this area due to environmental factors.<ref>[https://vimeo.com/90248199 Polar Bears and Glaciers of Baffin Island Webinar] on [[Vimeo]]. Arctic Kingdom. 2014. retrieved on 9 June 2014</ref>
-===Predation===
+=== Predation ===
-The principal predators of bowheads are [[human]]s.<ref name="MonIntro">Montague, J. J. (1993). "Introduction". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref> [[Killer whales]] are also known predators.<ref name="MMPhil">Philo, L. M., E. B. Shotts, and J. C. George (1993). "Morbidity and Mortality". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref> Bowheads seek the safety of the ice and shallow waters when threatened by killer whales.<ref name= "Finely, K.J. 2001" />
+The principal predators of bowheads are [[human]]s.<ref name="MonIntro">Montague, J. J. (1993). "Introduction". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref> [[Killer whales]] are also known predators.<ref name="MMPhil">Philo, L. M., E. B. Shotts, and J. C. George (1993). "Morbidity and Mortality". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref> Bowheads seek the safety of the ice and shallow waters when threatened by killer whales.<ref name="Finely, K.J. 2001" />
-==Whaling==
+== Whaling ==
 [[File:18th century arctic whaling.jpg|thumb|left|Eighteenth century engraving showing Dutch whalers hunting bowhead whales in the [[Arctic]]|alt=Two whaleboats beached in foreground, 5 rowed and 4 sailing whaleboats chasing/attacking 5 whales, two larger whaling ships nearby, and sun peeking around snow-covered mountain in background]]
-The bowhead whale has been [[whaling|hunted]] for [[blubber]], meat, [[whale oil|oil]], bones, and [[baleen]]. Like [[right whales]], it swims slowly, and floats after death, making it ideal for whaling.<ref name= "NOAA 2015">{{cite web |title=Bowhead Whale (''Balaena mysticetus'') |url=http://www.nmfs.noaa.gov/pr/species/mammals/cetaceans/bowheadwhale.htm |work=National Oceanic and Atmospheric Administration |accessdate=23 November 2015}}</ref> Before commercial whaling, they were estimated to number 50,000.<ref name="SSWoo">Woodby, D. A. and D. B. Botkin (1993). "Stock Sizes Prior to Commercial Whaling". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref>
+The bowhead whale has been [[whaling|hunted]] for [[blubber]], meat, [[whale oil|oil]], bones, and [[baleen]]. Like [[right whales]], it swims slowly, and floats after death, making it ideal for whaling.<ref name="NOAA 2015">{{cite web |title=Bowhead Whale (''Balaena mysticetus'') |url=http://www.nmfs.noaa.gov/pr/species/mammals/cetaceans/bowheadwhale.htm |work=National Oceanic and Atmospheric Administration |accessdate=23 November 2015}}</ref> Before commercial whaling, they were estimated to number 50,000.<ref name="SSWoo">Woodby, D. A. and D. B. Botkin (1993). "Stock Sizes Prior to Commercial Whaling". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref>
 Commercial bowhead whaling began in the 16th century, when the Basques killed them as they migrated south through the [[Strait of Belle Isle]] in the fall and early winter. In 1611, the first whaling expedition sailed to [[Spitsbergen]]. By mid-century, the population(s) there had practically been wiped out, forcing whalers to voyage into the "West Ice"—the pack ice off [[Greenland]]'s east coast. By 1719, they had reached the [[Davis Strait]], and by the first quarter of the 19th century, [[Baffin Bay]].<ref name="Ross">Ross, W. G. (1993). "Commercial Whaling in the North Atlantic Sector". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref>
@@ Line 178: / Line 180: @@
 In the North Pacific, the first bowheads were taken off the eastern coast of Kamchatka by the Danish whaleship ''Neptun'', Captain Thomas Sodring, in 1845.<ref name="CWNPBock">Bockstoce, J. R., and J. J. Burns (1993). "Commercial Whaling in the North Pacific Sector". In Burns, J. J.; Montague, J. J.; and Cowles, C. J. ''The Bowhead Whale''. Special Publication No. 2: The Society for Marine Mammalogy.</ref> In 1847, the first bowheads were caught in the Sea of Okhotsk, and the following year, Captain [[Thomas Welcome Roys]], in the bark ''Superior'', of [[Sag Harbor, New York|Sag Harbor]], caught the first bowheads in the [[Bering Strait]] region. By 1849, 50 ships were hunting bowheads in each area. By 1852, 220 ships were cruising around the Bering Strait region, which killed over 2,600 whales. Between 1854 and 1857, the fleet shifted to the Sea of Okhotsk, where 100–160 ships cruised annually. During 1858–1860, the ships shifted back to the Bering Strait region, where the majority of the fleet would cruise during the summer up until the early 20th century.<ref name="Bockstoce1986">{{cite book |last=Bockstoce |first=John |title=Whales, Ice, & Men: The History of Whaling in the Western Arctic |year=1986 |publisher=University of Washington Press |isbn=0-295-97447-8 }}</ref> An estimated 18,600 bowheads were killed in the Bering Strait region between 1848 and 1914, with 60% of the total being reached within the first two decades. An estimated 18,000 bowheads were killed in the Sea of Okhotsk during 1847–1867, 80% in the first decade.<ref name=Vaughan1984>Vaughan, R. (1984). ''Historical survey of the European whaling industry''. In ''Arctic Whaling: Proceedings of the International Symposium'', pp. 121-145. University of Groningen.</ref>
-Bowheads were first taken along the pack ice in the northeastern Sea of Okhotsk, then in [[Taui Bay|Tausk Bay]] and Northeast Gulf ([[Shelikhov Gulf]]). Soon, ships expanded to the west, catching them around [[Iony Island]] and then around the [[Shantar Islands]]. In the Western Arctic, they mainly caught them in the [[Gulf of Anadyr|Anadyr Gulf]], the Bering Strait, and around [[St. Lawrence Island]]. They later spread to the western [[Beaufort Sea]] (1854) and the [[Mackenzie River]] delta (1889).<ref name="Bockstoce1986"/>
+Bowheads were first taken along the pack ice in the northeastern Sea of Okhotsk, then in [[Taui Bay|Tausk Bay]] and Northeast Gulf ([[Shelikhov Gulf]]). Soon, ships expanded to the west, catching them around [[Iony Island]] and then around the [[Shantar Islands]]. In the Western Arctic, they mainly caught them in the [[Gulf of Anadyr|Anadyr Gulf]], the Bering Strait, and around [[St. Lawrence Island]]. They later spread to the western [[Beaufort Sea]] (1854) and the [[Mackenzie River]] delta (1889).<ref name="Bockstoce1986" />
 [[File:Bowhead Whale 2002-08-10.jpg|thumb|Inuit woman and child standing on bowhead whale after a 2002 subsistence hunt]]
-Commercial whaling, the principal cause of the population decline, is over. Bowhead whales are now hunted on a [[subsistence]] level by [[Indigenous peoples of the Americas|native peoples of North America]].<ref name= "WWF 2015">{{cite web |title=Bowhead Whale |url=http://wwf.panda.org/what_we_do/endangered_species/cetaceans/about/right_whales/bowhead_whale/ |work=WWF |accessdate=1 December 2015}}</ref>
+Commercial whaling, the principal cause of the population decline, is over. Bowhead whales are now hunted on a [[subsistence]] level by [[Indigenous peoples of the Americas|native peoples of North America]].<ref name="WWF 2015">{{cite web |title=Bowhead Whale |url=http://wwf.panda.org/what_we_do/endangered_species/cetaceans/about/right_whales/bowhead_whale/ |work=WWF |accessdate=1 December 2015}}</ref>
 === Subsistence whaling in Alaska ===
@@ Line 190: / Line 192: @@
 For the Native peoples of Alaska subsistence Hunting occurs all year, and is normally managed under the same regulations as general season, drawing, and registration hunts, and a hunting license and harvest tag is usually required. For [[marine mammal]]s, in specific, subsistence hunting is managed by the [[National Marine Fisheries Service]].<ref>{{Cite web|title = Subsistence Hunting, Alaska Department of Fish and Game|url = http://www.adfg.alaska.gov/index.cfm?adfg=subsistence.hunting|website = www.adfg.alaska.gov|access-date = 2016-02-24}}</ref>
-==Conservation==
+== Conservation ==
-The bowhead is listed in Appendix I by [[CITES]] (that is, "threatened with extinction"). It is listed by the [[National Marine Fisheries Service]] as "endangered" under the auspices of the United States' [[Endangered Species Act]]. The [[IUCN Red List]] data are as follows:<ref name= "NOAA 2015"/>
+The bowhead is listed in Appendix I by [[CITES]] (that is, "threatened with extinction"). It is listed by the [[National Marine Fisheries Service]] as "endangered" under the auspices of the United States' [[Endangered Species Act]]. The [[IUCN Red List]] data are as follows:<ref name="NOAA 2015" />
 *[[Svalbard]] population – Critically endangered
 *[[Sea of Okhotsk]] subpopulation – Endangered
@@ Line 198: / Line 200: @@
 *[[Bering Sea|Bering]]-[[Chukchi Sea|Chukchi]]-[[Beaufort Sea|Beaufort]] stock – Lower risk – conservation dependent
-The bowhead whale is listed in Appendix I<ref name="Appendices">"[https://web.archive.org/web/20110611112003/http://www.cms.int/documents/appendix/Appendices_COP9_E.pdf Appendix I]" of the Convention on the Conservation of Migratory Species of Wild Animals (CMS). As amended by the Conference of the Parties in 1985, 1988, 1991, 1994, 1997, 1999, 2002, 2005, and 2008. Effective: 5 March 2009.</ref> of the Convention on the Conservation of Migratory Species of Wild Animals ([[Bonn Convention|CMS]]), as this species has been categorized as being in danger of extinction throughout all or a significant proportion of their range. CMS Parties strive towards strictly protecting these animals, conserving or restoring the places where they live, mitigating obstacles to migration, and controlling other factors that might endanger them.<ref name= "NOAA 2015"/>
+The bowhead whale is listed in Appendix I<ref name="Appendices">"[https://web.archive.org/web/20110611112003/http://www.cms.int/documents/appendix/Appendices_COP9_E.pdf Appendix I]" of the Convention on the Conservation of Migratory Species of Wild Animals (CMS). As amended by the Conference of the Parties in 1985, 1988, 1991, 1994, 1997, 1999, 2002, 2005, and 2008. Effective: 5 March 2009.</ref> of the Convention on the Conservation of Migratory Species of Wild Animals ([[Bonn Convention|CMS]]), as this species has been categorized as being in danger of extinction throughout all or a significant proportion of their range. CMS Parties strive towards strictly protecting these animals, conserving or restoring the places where they live, mitigating obstacles to migration, and controlling other factors that might endanger them.<ref name="NOAA 2015" />
-==Gallery==
+== Gallery ==
 {{Commons-inline|Balaena mysticetus}}
 <gallery>
 File:Bowhead whales swimming in Lingolm strait by Vladislav Raevskii.JPG|Whales swimming in Lindgolm strait ''({{lang-ru|пролив Линдгольма}})'' of [[Shantar Islands]], northwestern Sea of Okhotsk<ref>[https://plus.google.com/photos/103061088309322077718/albums/5506708198561849937?banner=pwa Vladislav Raevskii]. Retrieved 1 June 2014</ref>
-File:Bowhead whales cavorting in the coastal water of north western Sea of Okhotsk by Olga Shpak, Marine Mammal Council, IEE RAS.jpg|Cavorting whales in northwestern part of Sea of Okhotsk<ref name="Olga Shpak. 2014"/>
+File:Bowhead whales cavorting in the coastal water of north western Sea of Okhotsk by Olga Shpak, Marine Mammal Council, IEE RAS.jpg|Cavorting whales in northwestern part of Sea of Okhotsk<ref name="Olga Shpak. 2014" />
 File:Grönlandwal 1-1999.jpg|Blowholes
 File:Grönlandwal 2-1999.jpg|Resting in [[Foxe Basin]]
@@ Line 213: / Line 215: @@
 </gallery>
-==See also==
+== See also ==
 {{Portal|Cetaceans|Marine life}}
-*[[List of cetaceans|List of whale and dolphin species]]
+*[[List of cetaceans]] - whale and dolphin species
 {{-}}
-==References==
+== References ==
 {{Reflist|30em}}
-==External links==
+== External links ==
 {{Commons category|Balaena mysticetus}}
 {{Wikispecies|Balaena mysticetus}}