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Yellow-throated marten (M. flavigula)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Mustelidae
Subfamily: Guloninae
J.E.Gray, 1825[1]

Guloninae[2][3] is a subfamily of the mammal family Mustelidae distributed across Eurasia and the Americas. It includes martens and the fisher, tayra and wolverine.[2][3] These genera were formerly included within a paraphyletic definition of the mustelid subfamily Mustelinae.[4]

Most gulonine species are arboreal to a degree. Some of the fashion furs come from this subfamily, e.g. sable, marten.[5]


Extant species[edit]

Tribe Image Genus Living species
Gulonini Eira Hamilton Smith, 1842
Gulo Pallas, 1780
Martini Martes Pinel, 1792 (martens)
Pekania Gray, 1865

Extinct genera[edit]

  • Aragonictis Valenciano et al., 2022[6] - Middle Miocene Europe
    • A. araid
  • Circamustela Petter, 1967[6] - Middle to Late Miocene Europe
    • C. dechaseauxi
    • C. peignei
    • C.? laevidens
  • Dehmictis Ginsburg and Morales, 1992[6] - Early Miocene Europe
  • Eiricitis [7] - Early Pliocene Asia
    • E. pachygnatha
  • Laphictis Viret, 1933[6]
  • Ischyrictis Helbing, 1930[6]
  • Plesiogulo? Zdansky, 1924[8][7] - Middle Miocene to Pliocene
    • P. brachygnathus (Schlosser, 1903)
    • P. botori Haile-Selassie, Hlusko & Howell, 2004
    • P. crassa Teilhard de Chardin, 1945
    • P. marshalli (Martin, 1928)
    • P. lindsayi Harrison, 1981
    • P. monspessulanus Viret, 1939
    • P. praecocidens Kurtén, 1970
  • Sinictis Zdansky, 1924[6]
    • S. dolichognathus
  • Sminthosinis Bjork, 1970[7] - Middle Miocene North America
    • S. bowleri


  1. ^ Gray, J.E. (1825). "Outline of an attempt at the disposition of the Mammalia into tribes and families with a list of the genera apparently appertaining to each tribe". Annals of Philosophy. New Series. 10: 337–344.
  2. ^ a b Nascimento, F. O. do (2014). "On the correct name for some subfamilies of Mustelidae (Mammalia, Carnivora)". Papéis Avulsos de Zoologia. 54 (21): 307–313. doi:10.1590/0031-1049.2014.54.21.
  3. ^ a b Law, C. J.; Slater, G. J.; Mehta, R. S. (2018-01-01). "Lineage Diversity and Size Disparity in Musteloidea: Testing Patterns of Adaptive Radiation Using Molecular and Fossil-Based Methods". Systematic Biology. 67 (1): 127–144. doi:10.1093/sysbio/syx047. PMID 28472434.
  4. ^ Koepfli KP, Deere KA, Slater GJ, et al. (2008). "Multigene phylogeny of the Mustelidae: Resolving relationships, tempo and biogeographic history of a mammalian adaptive radiation". BMC Biol. 6: 4–5. doi:10.1186/1741-7007-6-10. PMC 2276185. PMID 18275614.
  5. ^ "History of Fur in Fashion: Introduction". 4 December 2011.
  6. ^ a b c d e f Valenciano, A.; Morales, J.; et al. (January 2022). "Aragonictis araid, gen. et sp. nov., a small-sized hypercarnivore (Carnivora, Mustelidae) from the upper middle Miocene of the Iberian Peninsula (Spain)". Journal of Vertebrate Paleontology. 41 (5): e2005615. doi:10.1080/02724634.2021.2005615.
  7. ^ a b c Samuels, Joshua X.; Bredehoeft, Keila E.; Wallace, Steven C. (2018-04-18). "A new species of Gulo from the Early Pliocene Gray Fossil Site (Eastern United States); rethinking the evolution of wolverines". PeerJ. 6: e4648. doi:10.7717/peerj.4648. ISSN 2167-8359. PMC 5910791. PMID 29682423.
  8. ^ Valenciano, Alberto; Govender, Romala (2020-06-01). "New insights into the giant mustelids (Mammalia, Carnivora, Mustelidae) from Langebaanweg fossil site (West Coast Fossil Park, South Africa, early Pliocene)". PeerJ. 8: e9221. doi:10.7717/peerj.9221. ISSN 2167-8359. PMC 7271888. PMID 32547866.

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