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'''''Shorea''''' is a [[genus]] of about 196 [[species]] of mainly [[rainforest]] [[trees]] in the [[Family (biology)|family]] [[Dipterocarpaceae]]. The genus is named after Sir [[John Shore, 1st Baron Teignmouth|John Shore]], the Governor-General of the [[British East India Company]], 1793–1798. |
'''''Shorea''''' is a [[genus]] of about 196 [[species]] of mainly [[rainforest]] [[trees]] in the [[Family (biology)|family]] [[Dipterocarpaceae]]. The genus is named after Sir [[John Shore, 1st Baron Teignmouth|John Shore]], the Governor-General of the [[British East India Company]], 1793–1798. The [[timber]] of trees of the genus is sold under the common names '''lauan, luan, lawaan, meranti, seraya, balau, bangkirai''' and '''Philippine mahogany'''.<ref>[https://www.wood-database.com/lauan/ Lauan - The Wood Database]</ref> |
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== Description == |
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Many economically important [[timber]] trees belong to the genus, sold under names including "meranti", "lauan", "luan", "lawaan", "seraya", "balau", "bangkirai" and "Philippine mahogany". |
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Shorea is native to southeast [[Asia]], from Northern [[India]] to [[Malaysia]], [[Indonesia]] and the [[Philippines]]. In west [[Malesia]] and the Philippines this genus dominates the skyline of the [[tropical]] [[forest]]s. The tallest documented tropical [[angiosperm]] is an 88.3-metre-tall ''[[Shorea faguetiana]]'' in the [[Tawau Hills National Park]], in [[Sabah]] on the island of [[Borneo]], and in that park at least five other species of the genus have been measured to be over 80 m tall: ''[[Shorea argentifolia|S. argentifolia]]'', ''[[Shorea gibbosa|S. gibbosa]]'', ''[[Shorea johorensis|S. johorensis]]'', ''[[Shorea smithiana|S. smithiana]]'' and ''[[Shorea superba|S. superba]]''.<ref>{{cite web | url = http://www.nativetreesociety.org/worldtrees/sea_ei/borneo_ii.htm | title = Borneo | publisher = [[Eastern Native Tree Society]] | accessdate= 2008-06-21}}</ref> Borneo is also the hotspot of ''Shorea'' diversity with 138 species, of which 91 are [[Endemism|endemic]] to the island.<ref name = Ashton>Ashton, P. S. "Dipterocarpaceae". In ''Tree Flora of Sabah and Sarawak,'' Volume 5, 2004. Soepadmo, E.; Saw, L. G. and Chung, R. C. K. eds. Government of Malaysia, Kuala Lumpur, Malaysia. {{ISBN|983-2181-59-3}}</ref> |
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== Species == |
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{{Main|List of Shorea species}} |
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==Reproductive biology== |
==Reproductive biology== |
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''Shorea'' are [[insect pollinated]] and a variety of insects have been implicated, with species within the sections of ''Shorea'' sharing the same insect pollinators. Flowering within a section is sequential within one habitat and species association to prevent competition for pollinators.<ref>{{cite journal |last= LaFrankie |first=James V. Jr. |author2=H. T. Chan |date=June 1991 |title= Confirmation of Sequential Flowering in ''Shorea'' (Dipterocarpaceae) |journal=Biotropica |volume=23 |issue=2 |pages=200–203 |doi= 10.2307/2388308 |jstor=2388308}}</ref> |
''Shorea'' are [[insect pollinated]] and a variety of insects have been implicated, with species within the sections of ''Shorea'' sharing the same insect pollinators. Flowering within a section is sequential within one habitat and species association to prevent competition for pollinators.<ref>{{cite journal |last= LaFrankie |first=James V. Jr. |author2=H. T. Chan |date=June 1991 |title= Confirmation of Sequential Flowering in ''Shorea'' (Dipterocarpaceae) |journal=Biotropica |volume=23 |issue=2 |pages=200–203 |doi= 10.2307/2388308 |jstor=2388308}}</ref> |
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==Uses== |
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Many economically important [[timber]] trees belong to ''Shorea''. They are sold under various trade names including " |
Many economically important [[timber]] trees belong to ''Shorea''. They are sold under various trade names including "Lauan", "Lawaan", "Meranti", "Seraya", "Balau", "Bangkirai" and "Philippine mahogany". (For a list of species associated with each name, see the article on [[Dipterocarp timber classification]]). The "Philippine mahogany" sold in North America is not a true [[mahogany]] at all, but a mixture of woods from the genus ''Shorea''. |
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Other products from ''Shorea'' spp. include [[dammar]] and [[Illepe]]. Dammar is a resin collected from a variety of species. It varies in colour among the different taxonomic groups. ''[[Shorea wiesneri]]'' is listed in many websites as an important source of dammar;<ref>{{cite encyclopedia |url= http://www.britannica.com/eb/article-9028654/dammar |title = Dammar |encyclopedia = Encyclopædia Britannica |accessdate= 2007-11-14}}</ref> however, this appears to be either a trade name or a [[Synonym (taxonomy)|synonym]].<ref>{{cite web | url = http://193.62.154.38/diptero/diptax.html | title = Dipterocarpaceae Data Base—Taxonomic Information | publisher = Royal Botanical Gardens, Edinburgh | accessdate= 2007-11-14}}</ref><ref>{{cite web | url = http://www.kew.org/searchepic/summaryquery.do;jsessionid=BD255C82F8006D581ED0D08C7CBC1300?scientificName=Shorea+wiesneri+&searchAll=true&categories=names&categories=bibl&categories=colln&categories=taxon&categories=flora&categories=misc | title = Electronic Plant Information Centre | publisher = Royal Botanical Gardens, Kew | accessdate= 2007-11-14}}</ref> |
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The "Philippine mahogany" sold in North America is not a [[mahogany]] at all, but a mixture of woods from the genus ''Shorea''. |
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[[Borneo tallow nut oil]] is extracted from the egg-shaped winged fruit of ''Shorea'' species.<ref>{{cite web|title=Minor oil crops - Individual monographs (Balanites-Borneo tallow nut-Brazil nut-Caryocar spp)|url=http://www.fao.org/docrep/X5043E/x5043E05.htm#Borneo%20tallow%20nut|website=www.fao.org|publisher=FAO|accessdate=15 April 2017}}</ref> |
[[Borneo tallow nut oil]] is extracted from the egg-shaped winged fruit of ''Shorea'' species.<ref>{{cite web|title=Minor oil crops - Individual monographs (Balanites-Borneo tallow nut-Brazil nut-Caryocar spp)|url=http://www.fao.org/docrep/X5043E/x5043E05.htm#Borneo%20tallow%20nut|website=www.fao.org|publisher=FAO|accessdate=15 April 2017}}</ref> |
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[[Category:Shorea| ]] |
[[Category:Shorea| ]] |
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Revision as of 09:29, 9 August 2018
| Shorea | |
|---|---|
| |
| Shorea roxburghii | |
Scientific classification 
| |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Angiosperms |
| Clade: | Eudicots |
| Clade: | Rosids |
| Order: | Malvales |
| Family: | Dipterocarpaceae |
| Tribe: | Shoreae |
| Genus: | Shorea Roxb. ex C.F.Gaertn. |
| Sections | |
See List of Shorea species for complete taxonomy to species level. | |
Shorea is a genus of about 196 species of mainly rainforest trees in the family Dipterocarpaceae. The genus is named after Sir John Shore, the Governor-General of the British East India Company, 1793–1798. The timber of trees of the genus is sold under the common names lauan, luan, lawaan, meranti, seraya, balau, bangkirai and Philippine mahogany.[1]
Description
Shorea is native to southeast Asia, from Northern India to Malaysia, Indonesia and the Philippines. In west Malesia and the Philippines this genus dominates the skyline of the tropical forests. The tallest documented tropical angiosperm is an 88.3-metre-tall Shorea faguetiana in the Tawau Hills National Park, in Sabah on the island of Borneo, and in that park at least five other species of the genus have been measured to be over 80 m tall: S. argentifolia, S. gibbosa, S. johorensis, S. smithiana and S. superba.[2] Borneo is also the hotspot of Shorea diversity with 138 species, of which 91 are endemic to the island.[3]
Species
Reproductive biology
The majority of Shorea are general flowering species. General flowering is an event that occurs at irregular intervals of 3–10 yr, in which nearly all dipterocarp species together with species of other families bloom heavily.[4] It is thought that general flowering evolved to satiate seed predators[5] and/or to facilitate pollination.[4] It appears that both explanations hold merit.[6] Flowering is thought to be triggered by droughts that occur during transition periods from La Niña to El Niño.[7] It is suggested that the magnitude of a flowering event is dependent on the timing of the droughts associated with the El Niño southern oscillation (ENSO) cycle, with the largest events occurring after an interval of several years with no flowering.[7]
Shorea are insect pollinated and a variety of insects have been implicated, with species within the sections of Shorea sharing the same insect pollinators. Flowering within a section is sequential within one habitat and species association to prevent competition for pollinators.[8]
Uses
Many economically important timber trees belong to Shorea. They are sold under various trade names including "Lauan", "Lawaan", "Meranti", "Seraya", "Balau", "Bangkirai" and "Philippine mahogany". (For a list of species associated with each name, see the article on Dipterocarp timber classification). The "Philippine mahogany" sold in North America is not a true mahogany at all, but a mixture of woods from the genus Shorea.
Other products from Shorea spp. include dammar and Illepe. Dammar is a resin collected from a variety of species. It varies in colour among the different taxonomic groups. Shorea wiesneri is listed in many websites as an important source of dammar;[9] however, this appears to be either a trade name or a synonym.[10][11]
Borneo tallow nut oil is extracted from the egg-shaped winged fruit of Shorea species.[12]
Conservation status
One hundred and forty eight species of Shorea are currently listed on the IUCN Redlist, the majority of which are listed as being critically endangered.[13] There are some concerns regarding the IUCN's listing of Dipterocarps, as the criteria used to assess the level of threat are based mainly on animal population characteristics. This is thought to overstate the threat assessment, when applied to long-lived habitat-specific organisms such as trees.[3] Furthermore, one species reportedly extinct, on the IUCN Red list, Shorea cuspidata is reported to be common in the Bako National Park and also present in the Lambir National Park.[3] The Shorea species page gives threat classifications.
| IUCN red list category | Number of species |
|---|---|
| Extinct | 1 |
| Critically Endangered | 102 |
| Endangered | 34 |
| Vulnerable | 3 |
| Least concern | 6 |
| Data deficient | 2 |
| Not evaluated | ~48 |
References
- ^ Lauan - The Wood Database
- ^ "Borneo". Eastern Native Tree Society. Retrieved 2008-06-21.
- ^ a b c Ashton, P. S. "Dipterocarpaceae". In Tree Flora of Sabah and Sarawak, Volume 5, 2004. Soepadmo, E.; Saw, L. G. and Chung, R. C. K. eds. Government of Malaysia, Kuala Lumpur, Malaysia. ISBN 983-2181-59-3
- ^ a b Sakai, Shoko; K Momose; T Yumoto; T Nagamitsu; H Nagamasu; A A Hamid; T Nakashizuka (1999). "Plant reproductive phenology over four years including an episode of general flowering in a lowland dipterocarp forest, Sarawak, Malaysia". American Journal of Botany. 86 (10): 1414–36. doi:10.2307/2656924. JSTOR 2656924. PMID 10523283. Retrieved 2007-11-13.
- ^ Curran, Lisa M.; M. Leighton (2000). "Vertebrate responses to spatiotemporal variation in seed production of mast-fruiting Dipterocarpaceae". Ecological Monographs. 70 (1): 101–128. doi:10.1890/0012-9615(2000)070[0101:VRTSVI]2.0.CO;2. Retrieved 2007-11-13.
- ^ Maycock, Colin R.; R. N. Thewlis; J. Ghazoul; R. Nilus; David F. R. P. Burslem (2005). "Reproduction of dipterocarps during low intensity masting events in a Bornean rain forest". Journal of Vegetation Science. 16 (6): 635–46. doi:10.1658/1100-9233(2005)016[0635:RODDLI]2.0.CO;2. Retrieved 2007-11-13.
- ^ a b Sakai, Shoko; Rhett D. Harrison; Kuniyasu Momose; Koichiro Kuraji; Hidetoshi Nagamasu; Tetsuzo Yasunari; Lucy Chong; Tohru Nakashizuka (2006). "Irregular droughts trigger mass flowering in aseasonal tropical forests in Asia". American Journal of Botany. 93 (8): 1134–39. doi:10.3732/ajb.93.8.1134. PMID 21642179. Retrieved 2007-11-13.
- ^ LaFrankie, James V. Jr.; H. T. Chan (June 1991). "Confirmation of Sequential Flowering in Shorea (Dipterocarpaceae)". Biotropica. 23 (2): 200–203. doi:10.2307/2388308. JSTOR 2388308.
- ^ "Dammar". Encyclopædia Britannica. Retrieved 2007-11-14.
- ^ "Dipterocarpaceae Data Base—Taxonomic Information". Royal Botanical Gardens, Edinburgh. Retrieved 2007-11-14.
- ^ "Electronic Plant Information Centre". Royal Botanical Gardens, Kew. Retrieved 2007-11-14.
- ^ "Minor oil crops - Individual monographs (Balanites-Borneo tallow nut-Brazil nut-Caryocar spp)". www.fao.org. FAO. Retrieved 15 April 2017.
- ^ "IUCN Red List of Threatened Species, Shorea search results". IUCN. 2012. Retrieved 3 January 2013.
