2019 in paleoichthyology

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This list of fossil fishes described in 2019 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes of every kind that were described during the year 2019, as well as other significant discoveries and events related to paleoichthyology that occurred in 2019.

New taxa

Jawless vertebrates

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Archegonaspis bashkirica^[1]	Sp. nov	Valid	Märss	Silurian (Ludlow)		Russia ( Bashkortostan)	A member of the family Cyathaspididae.
Cyathaspis alexanderi^[1]	Sp. nov	Valid	Märss	Silurian	Tabuska Beds	Russia ( Chelyabinsk Oblast)	A member of the family Cyathaspididae.
Jiaoyu^[2]	Gen. et sp. nov	Valid	Liu et al.	Silurian	Kezirtag formation	China	A member of Galeaspida of uncertain phylogenetic placement. The type species is J. imperfectus.
Parathelodus liaokuoensis^[3]	Sp. nov	Valid	Cui et al.	Devonian (Lochkovian)	Xishancun Formation Xitun Formation	China	A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
Parathelodus wangi^[3]	Sp. nov	Valid	Cui et al.	Devonian (Lochkovian)	Xishancun Formation Xitun Formation	China	A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
Parathelodus xitunensis^[3]	Sp. nov	Valid	Cui et al.	Devonian (Lochkovian)	Xitun Formation	China	A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
Tethymyxine^[4]	Gen. et sp. nov	Valid	Miyashita et al.	Late Cretaceous (Cenomanian)	Hjoula Lagerstätte	Lebanon	A hagfish. The type species is T. tapirostrum.
Tolypelepis bedovensis^[1]	Sp. nov	Valid	Märss	Late Silurian	Krasnaya Bukhta Formation	Russia ( Krasnoyarsk Krai)	A member of the family Cyathaspididae.
Xiyuaspis^[2]	Gen. et comb. nov	Valid	Liu et al.	Silurian	Tataertag Formation	China	A member of Galeaspida belonging to the group Hanyangaspidida and to the family Xiushuiaspidae. The type species is "Nanjiangaspis" zhangi Lu et al. (2007).

Placoderms

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Asterolepis alticristata^[5]	Sp. nov	Valid	Downs et al.	Devonian (Frasnian)	Fram Formation	Canada ( Nunavut)	An antiarch in Asterolepis
Colombialepis^[6]	Gen. et sp. nov	Valid	Olive et al.	Late Devonian	Cuche Formation	Colombia	A member of Antiarchi, superfamily Asterolepidoidei. Genus includes new species C. villarroeli.
Colombiaspis^[6]	Gen. et sp. nov	Valid	Olive et al.	Late Devonian	Cuche Formation	Colombia	A member of Arthrodira, family Groenlandaspidae. Genus includes new species C. rinconensis.
Meeksiella^[7]	Gen. et sp. nov	Valid	Trinajstic et al.	Devonian (Frasnian)	Snetnya Gora Beds	Estonia Latvia Russia ( Pskov Oblast)	A member of Ptyctodontida. The type species is "Chelyophorus" pskovensis Obruchev (1947).

Acanthodians

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Cheiracanthus peachi^[8]	Sp. nov	Valid	Den Blaauwen, Newman & Burrow	Devonian (Givetian)	Mey Flagstone Formation Rousay Flagstone Formation	United Kingdom
Serradentus^[9]	Gen. et sp. nov		Newman, Burrow & Den Blaauwen	Devonian (Givetian)	Tordalen Formation	Germany Norway	A member of Ischnacanthiformes. The type species is S. armstrongi.

Cartilaginous fishes

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Antiquaobatis^[10]	Gen. et sp. nov	Valid	Stumpf & Kriwet	Early Jurassic (Pliensbachian)		Germany	A member of Rajiformes of uncertain phylogenetic placement. The type species is A. grimmenensis.
Asflapristis^[11]	Gen. et sp. nov	Valid	Villalobos-Segura et al.	Late Cretaceous (Turonian)	Akrabou Formation	Morocco	A member of Sclerorhynchoidei belonging to the family Ptychotrygonidae. Genus includes new species A. cristadentis.
Cantioscyllium clementsi^[12]	Sp. nov	Valid	Case et al.	Late Cretaceous (Campanian)	Bladen Formation	United States ( North Carolina)	A member of the family Ginglymostomatidae.
Carcharhinus mancinae^[13]	Sp. nov	Valid	Ebersole, Cicimurri & Stringer	Eocene (Bartonian)	Gosport Sand Lisbon Formation	United States ( Alabama)	A species of Carcharhinus.
Carcharhinus underwoodi^[14]	Sp. nov	Valid	Samonds et al.	Eocene	Mahajanga Basin	Madagascar	A species of Carcharhinus.
Carcharias dominguei^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A species of Carcharias.
Centrosqualus mustardi^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A member of the family Squalidae.
Chlamydoselachus balli^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A species of Chlamydoselachus.
Cretalamna feldmanni^[16]	Sp. nov	Valid	Hoganson, Erickson & Holland	Late Cretaceous (Maastrichtian)	Fox Hills Formation	United States ( North Dakota)
Cretodus houghtonorum^[17]	Sp. nov	Valid	Shimada & Everhart	Late Cretaceous (Turonian)	Carlile Shale	United States ( Kansas)	A mackerel shark belonging to the family Pseudoscapanorhynchidae.
Dasyatis northdakotaensis^[16]	Sp. nov	Valid	Hoganson, Erickson & Holland	Late Cretaceous (Maastrichtian)	Fox Hills Formation	United States ( North Dakota)	A species of Dasyatis.
Diprosopovenator^[18]	Gen. et sp. nov	Valid	Stumpf, Scheer & Kriwet	Late Cretaceous (Cenomanian)	Hesseltal Formation	Germany	A member of Carcharhiniformes belonging to the new family Pseudoscyliorhinidae. The type species is D. hilperti.
Dykeius^[15]	Gen. et sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A member of the family Chlamydoselachidae. The type species is D. garethi.
Florenceodon^[15]	Gen. et sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A member of Carcharhiniformes belonging to the new family Florenceodontidae. Genus includes new species F. johnyi.
Galagadon^[19]	Gen. et sp. nov	Valid	Gates, Gorscak & Makovicky	Late Cretaceous (Maastrichtian)	Hell Creek Formation	United States ( South Dakota)	A carpet shark. The type species is G. nordquistae.
Glencartius^[20]	Gen. et comb. nov	Valid	Ginter & Skompski	Carboniferous (Viséan)		Poland United Kingdom	A member of Ctenacanthiformes of uncertain phylogenetic placement. The type species is "Ctenacanthus" costellatus Traquair (1884).
Gydoselache^[21]	Gen. et sp. nov	Valid	Maisey et al.	Devonian (Emsian)	Gydo Formation	South Africa	A cartilaginous fish of uncertain phylogenetic placement, related to Pucapampella and assigned to the new family Pucapampellidae. The type species is G. oosthuizeni.
Hessinodon^[15]	Gen. et sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A member of the family Dalatiidae. Genus includes new species H. wardi.
Hypolophites beckeri^[22]	Sp. nov	Valid	Maisch	Paleocene	Midway Group	United States ( Arkansas)	A member of Myliobatiformes.
Isogomphodon aikenensis^[23]	Sp. nov	Valid	Cicimurri & Knight	Eocene (Priabonian)	Dry Branch Formation	United States ( South Carolina)	A relative of the daggernose shark.
Kimmerobatis^[24]	Gen. et sp. nov	Valid	Underwood & Claeson	Late Jurassic (Kimmeridgian)	Kimmeridge Clay	United Kingdom	A member of Batoidea related to Spathobatis. Genus includes new species K. etchesi.
Komoksodon^[15]	Gen. et sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A member of Hexanchiformes belonging to the new family Komoksodontidae. Genus includes new species K. kwutchakuth.
Lessiniabatis^[25]	Gen. et sp. nov	Valid	Marramà et al.	Eocene (Ypresian)	Monte Bolca Konservat-Lagerstätte	Italy	A member of Myliobatiformes belonging to the superfamily Dasyatoidea. The type species is L. aenigmatica.
Myledaphus araucanus^[26]	Sp. nov	Valid	Otero	Late Cretaceous (Maastrichtian)	Arauco Basin	Chile
"Myliobatis" foxhillsensis^[16]	Sp. nov	Valid	Hoganson, Erickson & Holland	Late Cretaceous (Maastrichtian)	Fox Hills Formation	United States ( North Dakota)	An eagle ray.
Odontorhytis bahariensis^[27]	Sp. nov	Valid	Salame & Asan	Eocene (probably Lutetian)		Egypt	A member of Neoselachii of uncertain phylogenetic placement.
Ottangodus^[28]	Gen. et sp. nov	Valid	Popov, Delsate & Felten	Middle Jurassic (Bajocian)		Luxembourg-French border	A chimaera belonging to the family Callorhinchidae. Genus includes new species O. lotharingiae.
Paraorthacodus rossi^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)
Pastinachus kebarensis^[29]	Sp. nov	Valid	Adnet et al.	Eocene (late Bartonian)		Tunisia	A cowtail stingray of Pastinachus. Announced in 2018; the final version of the article naming it was published in 2019.
Pristiophorus humboldti^[30]	Sp. nov	Valid	Villafaña et al.	Early Miocene		Chile	A species of Pristiophorus.
Pristiophorus pricei^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A species of Pristiophorus.
Pristiophorus smithi^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A species of Pristiophorus.
Protocentrophorus steviae^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A member of the family Centrophoridae.
Protoheptranchias^[15]	Gen. et sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A cow shark. Genus includes new species P. lowei.
Protolamna ricaurtensis^[31]	Sp. nov	Valid	Carrillo-Briceño, Parra & Luque	Early Cretaceous (late Barremian–early Aptian)	Paja Formation	Colombia	A mackerel shark belonging to the family Pseudoscapanorhynchidae.
Pseudabdounia^[13]	Gen. et comb. nov	Valid	Ebersole, Cicimurri & Stringer	Eocene	Gosport Sand Lisbon Formation Tallahatta Formation	Belgium United States ( Alabama)	A requiem shark. The type species is "Galeocerdo" recticonus Winkler (1874); genus also includes "Galeorhinus recticonus" claibornensis White (1956) (raised to the rank of a separate species Pseudabdounia claibornensis).
Ptychotrygon rostrispatula^[32]	Sp. nov	Valid	Villalobos‐Segura, Underwood & Ward	Late Cretaceous (Turonian)		Morocco
Rhinoscymnus clarki^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A member of the family Somniosidae.
Rolfodon^[15]	Gen. et sp. et comb. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous to Miocene	Angostura Formation Northumberland Formation Santa Marta Formation	Angola Antarctica Austria Canada ( British Columbia) Ecuador New Zealand	A member of the family Chlamydoselachidae. The type species is R. ludvigseni; genus also includes "Chlamydoselachus" bracheri Pfeil (1983), "Chlamydoselachus" fiedleri Pfeil (1983), "Chlamydoselachus" thomsoni Richter & Ward (1990), "Chlamydoselachus" goliath Antunes & Cappetta (2002), "Chlamydoselachus" keyesi Mannering & Hiller (2008), "Chlamydoselachus" tatere Consoli (2008) and "Chlamydoselachus" landinii Carrillo-Briceño, Aguilera & Rodriguez (2014).
Scymnodalatias kazenobon^[33]	Sp. nov	Valid	Nishimatsu & Ujihara	Middle Miocene	Yatsuo Group	Japan	A species of Scymnodalatias.
Squaliodalatias savoiei^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A member of the family Dalatiidae.
Squalus nicholsae^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A spurdog.
Synechodus dereki^[15]	Sp. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A member of the family Palaeospinacidae.
Tethytrygon^[34]	Gen. et comb. nov	Valid	Marramà et al.	Eocene (Ypresian)	Monte Bolca	Italy	A whiptail stingray belonging to the subfamily Neotrygoninae; a new genus for "Raja" muricata Volta (1796).
Tlalocbatos^[35]	Gen. et sp. nov	Valid	Brito, Villalobos-Segura & Alvarado-Ortega	Early Cretaceous	Tlayúa Formation	Mexico	A member of Batoidea related to the banjo rays. Genus includes new species T. applegatei.
Xampylodon^[15]	Gen. et comb. nov	Valid	Cappetta, Morrison & Adnet	Late Cretaceous (Campanian)	Northumberland Formation	Canada ( British Columbia)	A cow shark. Genus includes "Notidanus" dentatus Woodward (1886).

Ray-finned fishes

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acronuroides^[36]	Gen. et sp. nov.	Valid	Bannikov, Carnevale & Tyler	Eocene (late Ypresian)	Monte Bolca locality	Italy	A member of Percomorphacea of uncertain phylogenetic placement. The type species is A. eocaenicus.
Acropoma aurora^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan to Bortonian		New Zealand	A species of Acropoma.
Ambassis simesi^[37]	Sp. nov	Valid	Schwarzhans	Bortonian		New Zealand	A species of Ambassis.
Ampheristus pentlandensis^[37]	Sp. nov	Valid	Schwarzhans	Bortonian		New Zealand	A member of the family Ophidiidae.
Antigonia artata^[37]	Sp. nov	Valid	Schwarzhans	Waipawan-Heretaungan		New Zealand	A species of Antigonia.
Antimyctophum^[37]	Gen. et comb. nov	Valid	Schwarzhans	Clifdenian to Tongaporutuan		New Zealand	A lanternfish. The type species is "Scopelus" konganaruensis Frost (1933).
Archoplites langrellorum^[38]	Sp. nov	Valid	Van Tassell & Smith	Pliocene (Blancan)		United States ( Oregon)	A relative of the Sacramento perch.
Ardoreosomus^[39]	Gen. et sp. nov	Valid	Romano et al.	Early Triassic (Induan)	Candelaria Formation	United States ( Nevada)	A member of the family Ptycholepidae. Genus includes new species A. occidentalis.
Argyripnus oamaruensis^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Argyripnus.
Argyripnus rapahoensis^[37]	Sp. nov	Valid	Schwarzhans	Runangan		New Zealand	A species of Argyripnus.
Arnoglossus lautus^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian and Waitakian		New Zealand	A scaldfish.
Arnoglossus purus^[37]	Sp. nov	Valid	Schwarzhans	Lillburnian and Waiauan		New Zealand	A scaldfish.
Artediellichthys candelabrum^[40]	Sp. nov	Valid	Nazarkin	Miocene	Agnevo Formation	Russia ( Sakhalin Oblast)	A relative of the blackfin hookear sculpin.
Artediellus simplex^[40]	Sp. nov	Valid	Nazarkin	Miocene	Agnevo Formation	Russia ( Sakhalin Oblast)	A species of Artediellus.
Aseraggodes hudsoni^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Aseraggodes.
Auchenoceros simplex^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian		New Zealand	A relative of the ahuru.
Aulichthys miocaenicus^[41]	Sp. nov	Valid	Nazarkin	Miocene (Serravallian–Tortonian)	Agnevo Formation	Russia ( Sakhalin Oblast)	An extinct tubenose.
Ausonasynodus^[42]	Gen. et sp. nov	Valid	Carnevale et al.	Late Eocene		Spain	A lizardfish. Genus includes new species A. almerai.
Awamoa^[37]	Gen. et comb. et sp. nov	Valid	Schwarzhans	Otaian to Opoitian		New Zealand	A southern sandfish. The type species is "Leptoscopus" progressus Schwarzhans (1980); genus also includes "Leptoscopus" iocosus Schwarzhans (1980) and a new species A. kaawa.
Balistes vegai^[43]	Sp. nov	Valid	Viñola Lopez, Carr & Lorenzo	Miocene		Cuba	A species of Balistes.
Barathronus nielseni^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Barathronus.
Barbalepis^[44]	Gen. et comb. nov	Valid	Olive, Taverne & López-Arbarello	Early Cretaceous (Barremian–Aptian)		Belgium	A member the family Coccolepididae; a new genus for "Coccolepis" macroptera Traquair.
Bathycongrus waihaoensis^[37]	Sp. nov	Valid	Schwarzhans	Kaiatan		New Zealand	A member of the family Congridae. Originally described as a species of Bathycongrus, but subsequently transferred to the genus Smithconger.^[45]
Bathygadus waiohaensis^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Bathygadus.
Bembrops aequiformis^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian and Waitakian		New Zealand	A species of Bembrops.
Bembrops grenfelli^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Bembrops.
Benthosema minutum^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Benthosema.
Bidenichthys struthersi^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Bidenichthys.
Blachea semeniformis^[37]	Sp. nov	Valid	Schwarzhans	Waitakian to Altonian		New Zealand	A species of Blachea.
Bonapartia altidorsalis^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A member of the family Gonostomatidae.
Brosmophyciops tongarewae^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan-Heretaungan		New Zealand	A species of Brosmophyciops.
Burguklia minichorum^[46]	Sp. nov	Valid	Bakaev & Kogan	Permian (Wordian–Capitanian)		Russia	An early ray-finned fish. Announced in 2019; the final version of the article naming it was published in 2020.
Cabindachanos^[47]	Gen. et sp. nov	Valid	Taverne et al.	Paleocene (Danian or early Selandian)		Angola	A member of the family Chanidae. The type species is C. dartevellei.
Callionymus hakatarameaensis^[37]	Sp. nov	Valid	Schwarzhans	Waitakian		New Zealand	A species of Callionymus.
Callionymus triquetrus^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Callionymus.
Candelarialepis^[39]	Gen. et sp. nov	Valid	Romano et al.	Early Triassic (Induan)	Candelaria Formation	United States ( Nevada)	A stem-neopterygian belonging to the family Parasemionotidae. Genus includes new species C. argentus.
Cavinichthys^[48]	Gen. et sp. nov	Valid	Taverne & Capasso	Early Cretaceous (Albian)	Limestones of Pietraroja	Italy	A member of Crossognathiformes belonging to the family Pachyrhizodontidae. The type species is C. pachylepis.
Cephalopholis? aotearoa^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan to Bortonian		New Zealand	Possibly a species of Cephalopholis.
Cepola ambifaria^[37]	Sp. nov	Valid	Schwarzhans	Otaian to Clifdenian		New Zealand	A species of Cepola.
Ceratoscopelus richardsoni^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Ceratoscopelus.
Champsodon timaruensis^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Champsodon.
Chaunax bucculentus^[37]	Sp. nov	Valid	Schwarzhans	Waitakian		New Zealand	A species of Chaunax.
Chaychanus^[49]	Gen. et sp. nov	Valid	Cantalice Severiano, Alvarado Ortega & Bellwood	Paleocene		Mexico	A member of the family Pomacentridae. Genus includes new species C. gonzalezorum. Announced in 2019; the final version of the article naming it was published in 2020.
Chrionema salebrosa^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian to Altonian		New Zealand	A species of Chrionema.
Coelorinchus divulgatus^[37]	Sp. nov	Valid	Schwarzhans	Waitakian to Lillburnian		New Zealand	A species of Coelorinchus.
Coelorinchus pakaurangiensis^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian to Otaian		New Zealand	A species of Coelorinchus.
Coelorinchus preaustralis^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian to Altonian		New Zealand	A species of Coelorinchus.
Colobodus wushaensis^[50]	Sp. nov	Valid	Li et al.	Late Triassic (Carnian)	Falang Formation	China	A member of Perleidiformes.
Conger davidsmithi^[37]	Sp. nov	Valid	Schwarzhans	Waipawan to Bortonian		New Zealand	A species of Conger.
Conger tokoroa^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Conger.
Congopycnodus^[51]	Gen. et sp. nov	Valid	Taverne	Middle Jurassic	Stanleyville Formation	Democratic Republic of the Congo	A member of Pycnodontiformes belonging to the superfamily Coccodontoidea. The type species is C. cornutus.
Costapycnodus^[52]	Gen. et comb. nov	Valid	Taverne, Capasso & Del Re	Early Cretaceous (late Hauterivian-early Barremian)		Italy	A member of the family Pycnodontidae. The type species is "Coelodus" costae Heckel (1856).
Danaphos glomerosus^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Danaphos.
Diaphus audax^[37]	Sp. nov	Valid	Schwarzhans	Waitakian to Altonian		New Zealand	A species of Diaphus.
Diaphus caurus^[53]	Sp. nov	Valid	Schwarzhans & Ohe	Pleistocene (Calabrian)	Hijikata Formation	Japan	A species of Diaphus.
Diaphus endoi^[53]	Sp. nov	Valid	Schwarzhans & Ohe	Pliocene (Piacenzian)	Shinzato Formation	Japan	A species of Diaphus.
Diaphus exilis^[37]	Sp. nov	Valid	Schwarzhans	Otaian to Lillburnian		New Zealand	A species of Diaphus.
Diaphus grebneffi^[53]	Sp. nov	Valid	Schwarzhans & Ohe	Probably Piacenzian	Nakosi Formation	Fiji	A species of Diaphus.
Diaphus huatau^[37]	Sp. nov	Valid	Schwarzhans	Clifdenian to Lillburnian		New Zealand	A species of Diaphus.
Diaphus kaiparaensis^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Diaphus.
Diaphus kakegawaensis^[53]	Sp. nov	Valid	Schwarzhans & Ohe	Pleistocene (Gelasian)	Dainichi Formation	Japan	A species of Diaphus.
Diaphus manneringi^[37]	Sp. nov	Valid	Schwarzhans	Clifdenian to Waiauan		New Zealand	A species of Diaphus.
Diaphus mirus^[37]	Sp. nov	Valid	Schwarzhans	Tongaporutuan		New Zealand	A species of Diaphus.
Diaphus nafpaktitisi^[53]	Sp. nov	Valid	Schwarzhans & Ohe	Pliocene (Piacenzian)	Nobori Formation	Japan	A species of Diaphus.
Diaphus noboriensis^[53]	Sp. nov	Valid	Schwarzhans & Ohe	Pliocene (Piacenzian) and Pleistocene (Gelasian)	Dainichi Formation Nobori Formation Shinzato Formation	Japan	A species of Diaphus.
Diaphus tenax^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Diaphus.
Dibranchus kakahoensis^[37]	Sp. nov	Valid	Schwarzhans	Bortonian		New Zealand	A species of Dibranchus.
Diogenichthys rangiauriensis^[37]	Sp. nov	Valid	Schwarzhans	Mangapanian		New Zealand	A species of Diogenichthys.
Dipulus certus^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan-Heretaungan		New Zealand	A species of Dipulus.
Dolichopteryx iustus^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Dolichopteryx.
Echiodon teres^[37]	Sp. nov	Valid	Schwarzhans	Otaian to Clifdenian		New Zealand	A species of Echiodon.
Electrona subasperoides^[37]	Sp. nov	Valid	Schwarzhans	Waitakian to Altonian		New Zealand	A species of Electrona.
Ellimma longipectoralis^[54]	Sp. nov	Valid	Polck et al.	Early Cretaceous (Aptian)	Barra Velha Formation	Brazil	A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Paraclupeidae. Announced in 2019; the final version of the article naming it was published in 2020.
Emmelichthys tennysoni^[37]	Sp. nov	Valid	Schwarzhans	Waitakian		New Zealand	A species of Emmelichthys.
Encheliophis strigosus^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Encheliophis.
Eomyctophum broncus^[37]	Sp. nov	Valid	Schwarzhans	Waipawan to Porangan		New Zealand	A lanternfish.
Eomyctophum porokawa^[37]	Sp. nov	Valid	Schwarzhans	Bortonian		New Zealand	A lanternfish.
Eosemionotus diskosomus^[55]	Sp. nov	Valid	López-Arbarello et al.	Middle Triassic (Ladinian)	Meride Limestone	Italy Switzerland	A member of Semionotiformes.
Eosemionotus minutus^[55]	Sp. nov	Valid	López-Arbarello et al.	Middle Triassic (Ladinian)	Meride Limestone	Switzerland	A member of Semionotiformes.
Eosemionotus sceltrichensis^[55]	Sp. nov	Valid	López-Arbarello et al.	Middle Triassic (Ladinian)	Meride Limestone	Switzerland	A member of Semionotiformes.
Epaelops^[56]	Gen. et sp. nov	Valid	Alves, Alvarado-Ortega & Brito	Early Cretaceous (Albian)	Tlayúa Formation	Mexico	A member of Elopiformes. Genus includes new species E. martinezi. Announced in 2019; the final version of the article naming it was published in 2020.
Epigonus aquilonius^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Epigonus.
Epigonus opoitiensis^[37]	Sp. nov	Valid	Schwarzhans	Opoitian		New Zealand	A species of Epigonus.
Etrumeus carnatus^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Etrumeus.
Evynnis abax^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian to Altonian		New Zealand	A member of the family Sparidae.
Flagellipinna^[57]	Gen. et sp. nov	Valid	Cawley & Kriwet	Late Cretaceous (Cenomanian)	Sannine Formation	Lebanon	A member of Pycnodontiformes belonging to the family Pycnodontidae. The type species is F. rhomboides.
Forsterygion jawadi^[37]	Sp. nov	Valid	Schwarzhans	Nukumaruan		New Zealand	A species of Forsterygion.
Gadiculus adversus^[37]	Sp. nov	Valid	Schwarzhans	Waitakian to Lillburnian		New Zealand	A species of Gadiculus.
Gadiculus pereawa^[37]	Sp. nov	Valid	Schwarzhans	Tongaporutuan		New Zealand	A species of Gadiculus.
Galeichthys ohei^[37]	Sp. nov	Valid	Schwarzhans	Opoitian to Nukumaruan		New Zealand	A species of Galeichthys.
Gilmourella^[58]	Gen. et sp. nov.	Valid	Carnevale & Bannikov	Eocene (late Ypresian)	Monte Bolca locality	Italy	A member of Callionymoidei. The type species is G. minuta.
Gnathophis araroa^[37]	Sp. nov	Valid	Schwarzhans	Kapitean		New Zealand	A species of Gnathophis.
Gonostoma retunsum^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Gonostoma.
Grahamichthys frigophila^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A member of the family Thalasseleotrididae.
Guus^[59]	Gen. et comb. nov.	Valid	Bannikov	Eocene (late Ypresian)	Monte Bolca locality	Italy	A member of the family Tortonesidae. The type species is "Gobius" microcephalus Agassiz (1839).
Gyrodus huiliches^[60]	Sp. nov	Valid	Gouiric-Cavalli, Remírez & Kriwet	Early Cretaceous (Valanginian–early Hauterivian)	Agrio Formation	Argentina
Harpagifer? morgansi^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	Possibly a species of Harpagifer.
Hemerocoetes pukunati^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian to Altonian		New Zealand	A species of Hemerocoetes.
Hemerocoetes whiroki^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian to Altonian		New Zealand	A species of Hemerocoetes.
Hiascoactinus^[61]	Gen. et sp. nov	Valid	Kim et al.	Late Triassic	Amisan Formation	South Korea	A member of Redfieldiiformes. Genus includes new species H. boryeongensis. Announced in 2019; the final version of the article naming it was published in 2020.
Hintonia robertsi^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A lanternfish.
Howella monodens^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Howella.
Hygophum acutiventris^[37]	Sp. nov	Valid	Schwarzhans	Kapitean		New Zealand	A species of Hygophum.
Ichthyscopus pukeuriensis^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Ichthyscopus.
Klincigobius^[62]	Gen. et comb. et sp. nov	Valid	Bradić-Milinović, Ahnelt & Schwarzhans in Bradić-Milinović et al.	Early Miocene		Serbia	A member of the family Gobiidae. The type species is "Gobius" serbiensis Gaudant (1998); genus also includes new species K. andjelkovicae.
Kradimus^[63]	Gen. et sp. nov	Valid	Veysey, Brito & Martill	Late Cretaceous (Turonian)	Akrabou Formation	Morocco	A member of Crossognathiformes. Genus includes new species K. asflaensis. Announced in 2019; the final version of the article naming it was published in 2020.
Lactarius primigenius^[37]	Sp. nov	Valid	Schwarzhans	Bortonian to Kaiatan		New Zealand	A relative of the false trevally.
Lactarius pusillus^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A relative of the false trevally.
Laeops undulatus^[37]	Sp. nov	Valid	Schwarzhans	Lillburnian		New Zealand	A species of Laeops.
Lampanyctodes transitivus^[37]	Sp. nov	Valid	Schwarzhans	Kapitean		New Zealand	A lanternfish.
Lampanyctus popoto^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Lampanyctus.
Lampanyctus profestus^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Lampanyctus.
Lampanyctus waiohaensis^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Lampanyctus.
Lashanichthys^[64]	Gen. et comb. nov	Valid	Xu et al.	Middle Triassic (Anisian)		China	A member of Ginglymodi assigned to the group Kyphosichthyiformes and to the new family Lashanichthyidae. The type species is "Sangiorgioichthys" sui López-Arbarello et al. (2011); genus also includes "Sangiorgioichthys" yangjuanensis Chen et al. (2014).
Latellopsis^[65]	Gen. et comb. nov.	Valid	Bannikov & Zorzin	Eocene (late Ypresian)	Monte Bolca locality	Italy	A member of Perciformes of uncertain phylogenetic placement. The type species is "Psettopsis" latellai Bannikov (2005).
Lepidorhynchus frosti^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A member of the family Macrouridae.
Leptoscopus atavus^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A southern sandfish.
Lethrinus? crassiornatus^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan-Heretaungan		New Zealand	Possibly a species of Lethrinus.
Lindoeichthys^[66]	Gen. et sp. nov.	Valid	Murray et al.	Late Cretaceous (Maastrichtian)	Scollard Formation	Canada ( Alberta)	A member of Percopsiformes. Genus includes new species L. albertensis. Announced in 2019; the final version of the article naming it was published in 2020.
Liparis? hampdenensis^[37]	Sp. nov	Valid	Schwarzhans	Bortonian		New Zealand	Possibly a species of Liparis.
Loancorhynchus^[67]	Gen. et sp. nov	Valid	Otero	Middle Eocene	Millongue Formation	Chile	A relative of the swordfish. The type species is L. catrillancai.
Lophiodes hoi^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Lophiodes.
Lophius alveolus^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Lophius.
Lophius onokensis^[37]	Sp. nov	Valid	Schwarzhans	Mangapanian to Nukumaruan		New Zealand	A species of Lophius.
Lotella matata^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Lotella.
Lotella pfeili^[37]	Sp. nov	Valid	Schwarzhans	Altonian and Clifdenian		New Zealand	A species of Lotella.
Lotella spicata^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan-Heretaungan		New Zealand	A species of Lotella.
Macrurulus depressus^[37]	Sp. nov	Valid	Schwarzhans	Bortonian		New Zealand	A member of the family Merlucciidae.
Macrurulus fordycei^[37]	Sp. nov	Valid	Schwarzhans	Waipawan-Mangaorapan		New Zealand	A member of the family Merlucciidae.
Maoricottus^[37]	Gen. et comb. et 2 sp. nov	Valid	Schwarzhans	Altonian to Lillburnian		New Zealand	A member of the family Cottidae. The type species is "Cottidarum" impolitus Schwarzhans (1980); genus also includes new species M. calidophilus and M. ovatus.
Melamphaes leeae^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Melamphaes.
Moradebrichthys^[68]	Gen. et sp. nov	Valid	Cartanyà et al.	Middle Triassic (Ladinian)		Spain	A member of the family Perleididae. Genus includes new species M. vilasecae.
Muraenanguilla^[37]	Gen. et comb. et 2 sp. nov	Valid	Schwarzhans	Paleocene and Eocene		Belgium New Zealand	A member of Anguilloidei of uncertain phylogenetic placement. The type species is Otolithus (Trachini)" thevenini Priem (1906); genus also includes new species M. balegemensis and M. lacinata.
Myctophum bortonensis^[37]	Sp. nov	Valid	Schwarzhans	Bortonian		New Zealand	A species of Myctophum.
Myctophum tamumuensis^[37]	Sp. nov	Valid	Schwarzhans	Clifdenian to Waiauan		New Zealand	A species of Myctophum.
Myctophum tenellum^[37]	Sp. nov	Valid	Schwarzhans	Tongaporutuan		New Zealand	A species of Myctophum.
Mystocheilus^[38]	Gen. et sp. nov	Valid	Van Tassell & Smith	Pliocene (Blancan)		United States ( Oregon)	A member of the family Cyprinidae. The type species is M. fresti.
Nemadactylus trulliformis^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Nemadactylus.
Nemadactylus utoka^[37]	Sp. nov	Valid	Schwarzhans	Waitakian	Otekaike Limestone	New Zealand	A species of Nemadactylus.
Neobythites turpidus^[37]	Sp. nov	Valid	Schwarzhans	Runangan		New Zealand	A species of Neobythites.
Nezumia morgansi^[37]	Sp. nov	Valid	Schwarzhans	Bortonian and Kaiatan		New Zealand	A species of Nezumia.
Notoconger^[37]	Gen. et 2 sp. nov	Valid	Schwarzhans	Runangan to Altonian		New Zealand	A member of the family Congridae. The type species is N. hesperis; genus also includes N. devexus.
Notoscopelus effertus^[37]	Sp. nov	Valid	Schwarzhans	Kapitean		New Zealand	A species of Notoscopelus.
Notoscopelus praejaponicus^[53]	Sp. nov	Valid	Schwarzhans & Ohe	Pleistocene (Calabrian)	Chinen Formation Hijikata Formation	Japan	A species of Notoscopelus.
Opistognathus wharekuriensis^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian		New Zealand	A species of Opistognathus.
Oreochromimos^[69]	Gen. et sp. nov		Penk et al.	Miocene	Ngorora Formation	Kenya	A cichlid belonging to the tribe Oreochromini. The type species is O. kabchorensis.
Padangia^[70]	Gen. et comb. nov	Valid	Murray	Probably Eocene		Indonesia	A member of the family Cyprinidae. Genus includes "Sardinioides" amblyostoma von der Marck (1876).
Panturichthys grenfelli^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Panturichthys.
Paralabrus^[71]	Gen. et sp. nov.	Valid	Bannikov & Zorzin	Eocene (late Ypresian)	Monte Bolca locality	Italy	Possibly a wrasse. The type species is P. rossiae.
Parapercis bispicatus^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Parapercis.
Parapercis depressidorsalis^[37]	Sp. nov	Valid	Schwarzhans	Waitakian to Altonian		New Zealand	A species of Parapercis.
Parapercis fatoides^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Parapercis.
Parapercis pareoraensis^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Parapercis.
Parapercis richardsoni^[37]	Sp. nov	Valid	Schwarzhans	Waipawan to Porangan		New Zealand	A species of Parapercis.
Parapercis waiwaia^[37]	Sp. nov	Valid	Schwarzhans	Opoitian		New Zealand	A species of Parapercis.
Parascombrops giganteus^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan		New Zealand	A member of the family Acropomatidae.
Parascombrops schwarzhansi^[72]	Sp. nov	Valid	Van Hinsbergh & Helwerda	Late Pliocene to early Pleistocene		Philippines	A member of the family Acropomatidae.
Paraulopus rallus^[37]	Sp. nov	Valid	Schwarzhans	Bortonian		New Zealand	A species of Paraulopus.
Pelargorhynchus grandis^[73]	Sp. nov		Wallaard et al.	Late Cretaceous (Maastrichtian)	Maastricht Formation	Netherlands	A member of Aulopiformes belonging to the family Dercetidae.
Pempheris hurupiensis^[37]	Sp. nov	Valid	Schwarzhans	Tongaporutuan	Hurupi Formation	New Zealand	A species of Pempheris.
Phractocephalus yaguaron^[74]	Sp. nov	Valid	Bogan & Agnolín	Late Miocene		Argentina	A relative of the redtail catfish.
Physiculus beui^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Physiculus.
Platycephalus? iaiunus^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan to Bortonian		New Zealand	Possibly a species of Platycephalus.
Platycephalus? riremoana^[37]	Sp. nov	Valid	Schwarzhans	Runangan		New Zealand	Possibly a species of Platycephalus.
Platysiagum sinensis^[75]	Sp. nov	Valid	Wen et al.	Middle Triassic (Anisian)	Guanling Formation	China
Protomyctophum ahunga^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Protomyctophum.
Pseudaequalobythites^[37]	Gen. et sp. et comb. nov	Valid	Schwarzhans	Paleocene and Eocene		Belgium France Germany New Zealand United Kingdom	A member of the family Ophidiidae. The type species is P. biplex; genus also includes "Otolithus" hilgendorfi Koken (1891).
Pseudanthias multicrenatus^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan to Bortonian		New Zealand	A species of Pseudanthias.
Pseudocaranx? pertenuis^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	Possibly a species of Pseudocaranx.
Pseudophycis muringa^[37]	Sp. nov	Valid	Schwarzhans	Lillburnian		New Zealand	A species of Pseudophycis.
Pterygotrigla stewarti^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Pterygotrigla.
Pteronisculus nevadanus^[39]	Sp. nov	Valid	Romano et al.	Early Triassic (Induan)	Candelaria Formation	United States ( Nevada)	Possibly a member of the family Turseoidae.
Pycnodus multicuspidatus^[76]	Sp. nov	Valid	Vullo et al.	Paleocene (Thanetian)		Morocco
Quasinectes^[65]	Gen. et sp. nov.	Valid	Bannikov & Zorzin	Eocene (late Ypresian)	Monte Bolca locality	Italy	A member of Perciformes of uncertain phylogenetic placement. The type species is Q. durello.
Rebekkachromis^[77]	Gen. et 2 sp. nov	Valid	Kevrekidis, Valtl & Reichenbacher in Kevrekidis et al.	Miocene	Ngorora Formation	Kenya	A cichlid belonging to the subfamily Pseudocrenilabrinae and the tribe Oreochromini.^[78] The type species is R. ngororus; genus also includes R. kiptalami.
Rhamphogobius^[62]	Gen. et sp. et comb. nov	Valid	Bradić-Milinović, Ahnelt & Schwarzhans in Bradić-Milinović et al.	Early Miocene		Croatia Germany Serbia Switzerland	A member of the family Gobiidae. The type species is R. varidens; genus also includes "Gobius" doppleri Reichenbacher (1993), "Gobius" gregori Reichenbacher (1993) and "Gobius" helvetiae Salis (1967).
Rhynchoconger otaianus^[37]	Sp. nov	Valid	Schwarzhans	Waipawan-Mangaorapan		New Zealand	A species of Rhynchoconger.
Saccogaster parengarenga^[37]	Sp. nov	Valid	Schwarzhans	Otaian to Clifdenian		New Zealand	A species of Saccogaster.
Sardinella claviformis^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Sardinella.
Sardinella lintriculus^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Sardinella.
Schernfeldfuro^[79]	Gen. et sp. nov	Valid	Ebert	Late Jurassic		Germany	A member of Halecomorphi. Genus includes new species S. uweelleri.
Scleropages sanshuiensis^[80]	Sp. nov	Valid	Zhang	Early Eocene	Huachong Formation	China	A species of Scleropages. Announced in 2019; the final version of the article naming it was published in 2020.
Scopelarchoides neamticus^[81]	Sp. nov	Valid	Grădianu et al.	Oligocene	Lower Dysodilic Shales Formation	Romania	A pearleye, a species of Scopelarchoides.
Scopelosaurus? brevicauda^[37]	Sp. nov	Valid	Schwarzhans	Kaiatan		New Zealand	Possibly species of Scopelosaurus.
Sillaginodes albisaxosus^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A member of the family Sillaginidae.
Sillago maxwelli^[37]	Sp. nov	Valid	Schwarzhans	Bortonian		New Zealand	A species of Sillago.
Sprattus arewhana^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Sprattus.
Sundabarbus^[70]	Gen. et comb. nov	Valid	Murray	Probably Eocene	Sangkarewang Formation	Indonesia	A member of the family Cyprinidae. Genus includes "Barbus" megacephalus Günther (1876).
Symbolophorus moriguchii^[53]	Sp. nov	Valid	Schwarzhans & Ohe	Pliocene (Piacenzian) and Pleistocene (Gelasian)	Dainichi Formation Nobori Formation Shinzato Formation	Japan	A species of Symbolophorus.
Symbolophorus opononiensis^[37]	Sp. nov	Valid	Schwarzhans	Waitakian and Otaian		New Zealand	A species of Symbolophorus.
Symbolophorus tongaporutuensis^[37]	Sp. nov	Valid	Schwarzhans	Tongaporutuan		New Zealand	A species of Symbolophorus.
Symphysanodon inamata^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan to Bortonian		New Zealand	A species of Symphysanodon.
Thalasseleotris whatua^[37]	Sp. nov	Valid	Schwarzhans	Lillburnian		New Zealand	A species of Thalasseleotris.
Tharsis elleri^[82]	Sp. nov	Valid	Arratia, Schultze & Tischlinger	Late Jurassic (Tithonian)	Altmühltal Formation	Germany	A teleost belonging to the family Ascalaboidae.
Thiollierepycnodus^[83]	Gen. et comb. nov	Valid	Ebert	Late Jurassic		France Germany	A member of the family Pycnodontidae; a new genus for "Pycnodus wagneri Thiollière (1852).
Thryptodus loomisi^[84]	Sp. nov	Valid	Shimada	Late Cretaceous (Cenomanian–Turonian)	Britton Formation	United States ( Texas)	A member of the family Plethodidae.
Tonganago wharenga^[37]	Sp. nov	Valid	Schwarzhans	Waipawan to Bortonian		New Zealand	A member of the family Congridae.
Tongarewa^[37]	Gen. et 2 sp. nov	Valid	Schwarzhans	Duntroonian to Altonian		New Zealand	A threefin blenny. The type species is T. waihaoensis; genus also includes T. clementsi.
Toroatherina^[37]	Gen. et sp. et comb. nov	Valid	Schwarzhans	Eocene		New Zealand United States	An Old World silverside. The type species is T. toroa; genus also includes "Otolithus (Mugilidarum)" debilis Koken (1891).
Toxopyge^[62]	Gen. et sp. et comb. nov	Valid	Bradić-Milinović, Ahnelt & Schwarzhans in Bradić-Milinović et al.	Early Miocene		Serbia Switzerland	A member of the family Gobiidae. The type species is T. campylus; genus also includes "Gobius" longus Salis (1967).
Trachyrincus aulax^[37]	Sp. nov	Valid	Schwarzhans	Otaian to Tongaporutuan		New Zealand	A species of Trachyrincus.
Tranawuen^[60]	Gen. et comb. nov	Valid	Gouiric-Cavalli, Remírez & Kriwet	Early Cretaceous (Valanginian–early Hauterivian)	Agrio Formation	Argentina	A member of Pycnodontiformes. Genus includes "Macromesodon" agrioensis.
Trawdenia^[85]	Gen. et comb. nov	Valid	Coates & Tietjen	Carboniferous (Pennsylvanian)		United Kingdom	An early ray-finned fish. The type species is "Rhadinichthys" planti Traquair (1888); genus also includes "Mesopoma" carricki Coates (1993) and "Mesopoma" pancheni Coates (1993).
Tunisiaclupea^[86]	Gen. et sp. nov	Valid	Boukhalfa et al.	Early Cretaceous (Barremian)	Chotts Basin	Tunisia	A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Paraclupeidae. Genus includes new species T. speratus.
Uranoscopus rudis^[37]	Sp. nov	Valid	Schwarzhans	Duntroonian	Chatton Formation	New Zealand	A species of Uranoscopus.
Uranoscopus tectiformis^[37]	Sp. nov	Valid	Schwarzhans	Altonian		New Zealand	A species of Uranoscopus.
Valenciennellus fastigatus^[37]	Sp. nov	Valid	Schwarzhans	Otaian and Altonian		New Zealand	A species of Valenciennellus.
Veridagon^[87]^[88]	Gen. et sp. nov	Valid	Díaz-Cruz, Alvarado-Ortega & Carbot-Chanona	Late Cretaceous (Cenomanian)	Cintalapa Formation	Mexico	A member of Aulopiformes belonging to the family Enchodontidae. Genus includes new species V. avendanoi.
Vinciguerria orientalis^[89]	Sp. nov	Valid	Nam, Ko & Nazarkin	Middle Miocene	Duho Formation	South Korea	A species of Vinciguerria.
Waihaoclupea^[37]	Gen. et sp. nov	Valid	Schwarzhans	Mangaorapan to Bortonian		New Zealand	A member of the family Clupeidae. The type species is W. pinguis.
Waitahana^[37]	Gen. et comb. nov	Valid	Schwarzhans	Otaian to Waiauan		New Zealand	A southern sandfish. The type species is "Trachinoideorum" sagittiformis Schwarzhans (1980); genus also includes "Citharus" latisulcatus Frost (1924) and "Trachinoideorum" ultimus Schwarzhans (1980).
Waitakia aho^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan to Porangan		New Zealand	A member of the family Hemerocoetidae.
Waitakia proclinens^[37]	Sp. nov	Valid	Schwarzhans	Mangaorapan to Bortonian		New Zealand	A member of the family Hemerocoetidae.
Waitakia profunda^[37]	Sp. nov	Valid	Schwarzhans	Kaiatan to Runangan		New Zealand	A member of the family Hemerocoetidae.
Xenocephalus otaianus^[37]	Sp. nov	Valid	Schwarzhans	Otaian		New Zealand	A species of Xenocephalus.
Yudaiichthys^[64]	Gen. et sp. nov	Valid	Xu et al.	Middle Triassic (Anisian)	Guanling Formation	China	A member of Ginglymodi assigned to the group Kyphosichthyiformes and to the new family Lashanichthyidae. The type species is Y. eximius.
Zandtfuro^[79]	Gen. et sp. nov	Valid	Ebert	Late Jurassic		Germany	A member of Halecomorphi. Genus includes new species Z. tischlingeri.
Zonobythites cornifer^[37]	Sp. nov	Valid	Schwarzhans	Porangan and Bortonian		New Zealand	A member of the family Ophidiidae.

Lobe-finned fishes

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Anchidipterus^[90]	Gen. et sp. nov	Valid	Krupina in Lebedev, Krupina & Linkevich	Devonian (Famennian)		Russia ( Tver Oblast)	A lungfish. Genus includes new species A. dariae.
Clackodus^[91]	Gen. et comb. nov	Valid	Smithson, Challands & Smithson	Carboniferous (Serpukhovian)	Clackmannan Group	United Kingdom	A lungfish; a new genus for "Ctenodus" angustulus Traquair.
Ctenodus boudariensis^[92]	Sp. nov	Valid	Kemp	Carboniferous (Viséan)	Ducabrook Formation	Australia	A lungfish.
Isityumzi^[93]	Gen. et sp. nov	Valid	Gess & Clement	Devonian (Famennian)	Witpoort Formation	South Africa	A lungfish. The type species is I. mlomomde.
Limanichthys^[94]	Gen. et sp. nov	Valid	Challands et al.	Carboniferous (Tournaisian)	Ballagan Formation	United Kingdom	A lungfish. Genus includes new species L. fraseri.
Rubrognathus^[95]	Gen. et sp. nov	Valid	Lebedev & Clément	Devonian (late Givetian–early Frasnian)	Yam-Tesovo Formation	Russia ( Leningrad Oblast)	A member of Elpistostegalia. The type species is R. kuleshovi.
Selenodus^[96]	Gen. et sp. nov	Valid	Mondéjar-Fernández	Devonian (Eifelian)		Morocco	An onychodont sarcopterygian. Genus includes new species S. aquesbiae.
Whiteia uyenoteruyai^[97]	Sp. nov	Valid	Yabumoto et al.	Early Triassic	Middle Sakamena Group	Madagascar

Other fishes

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aberrosquama^[98]	Gen. et sp. nov	Valid	Burrow in Burrow et al.	Silurian (probably Gorstian)	Hamelin Formation	Australia	A stem-gnathostome of uncertain phylogenetic placement. The type species is A. occidens.

General research

Revision of Early Devonian psammosteids described by Beverly Halstead from the so-called "Placoderm Sandstone" (Świętokrzyskie Mountains, Poland) is published by Dec (2019).^[99]
A study on the locomotion of psammosteids, focusing on the influence of the shape of the psammosteid body form on hydrodynamic performance, is published by Dec (2019).^[100]
A study on the anatomy of dermal plates of Astraspis is published by Lemierre & Germain (2019), who report possible evidence of presence of proliferative cartilage in this taxon.^[101]
A study on the anatomy of the circulatory system of the head of Shuyu is published by Gai, Zhu & Donoghue (2019).^[102]
Redescription of Sinogaleaspis shankouensis, based on data from 11 new specimens from the Silurian Xikeng Formation (Jiangxi, China), is published online by Gai et al. (2019).^[103]
A study on the anatomy of the dermal skeleton of Tremataspis mammillata is published by O'Shea, Keating & Donoghue (2019).^[104]
Redescription of Asterolepis orcadensis based on newly collected fossil material is published by Newman, den Blaauwen & Leather (2019).^[105]
A study on the morphology of the jaw elements of a buchanosteoid placoderm specimen ANU V244 from the Early Devonian limestones (~400 Ma) at Burrinjuck, near Canberra (Australia) is published by Hu et al. (2019).^[106]
A specimen of Dunkleosteus terrelli preserving vertebrae fused into a structure known as the synarcual is described from the Devonian Cleveland Shale Member of the Ohio Shale Formation (Ohio, United States) by Johanson et al. (2019).^[107]
A redescription and a study on the phylogenetic relationships of a putative antiarch Silurolepis platydorsalis is published by Zhu, Lu & Zhu (2019), who reinterpret this species as a maxillate placoderm close to Qilinyu.^[108]
New body fossils of Cheiracanthus intricatus, including the first known articulated specimen of this species, are described from the Givetian Tordalen Formation (Spitsbergen, Norway) by Newman, Burrow & den Blaauwen (2019).^[109]
A study on the anatomy of the skeletal elements of the pharynx of Ptomacanthus anglicus, and on its implications for the knowledge of the evolution of the pharynx of jawed vertebrates, is published by Dearden, Stockey & Brazeau (2019).^[110]
Description of fossils of Carboniferous (Mississippian) cartilaginous fishes from the area of Krzeszowice (Poland) is published by Ginter & Złotnik (2019).^[111]
A study on microwear on teeth of Edestus minor, and on its implications for the knowledge of function of teeth of this fish, is published by Itano (2019).^[112]
A revision of species belonging to the genus Edestus is published by Tapanila & Pruitt (2019).^[113]
Discovery of a cast of the holotype of Petalodus ohioensis in the collections of the Yale Peabody Museum of Natural History is reported by Carpenter & Itano (2019), who consider the species Petalodus alleghaniensis to be a junior synonym of P. ohioensis.^[114]
Restudy of a putative bill of an ibis-like bird from the Eocene La Meseta Formation (Antarctica) described by Jadwiszczak, Gaździcki & Tatur (2008)^[115] is published by Agnolin, Bogan & Rozadilla (2019), who consider this specimen to be more likely to be a dorsal spine of a chimaeroid cartilaginous fish.^[116]
A study on the anatomy of the skull of Tristychius arcuatus, providing evidence of adaptations for suction feeding, is published by Coates et al. (2019).^[117]
Description of the first skeletal remains of Phoebodus from the Famennian of the Maïder region of Morocco, providing new information on the anatomy of this species, and a study on the phylogenetic affinities of Phoebodus is published by Frey et al. (2019).^[118]
Teeth of a hybodont shark belonging to the genus Asteracanthus, with anatomy indicative of a crushing feeding behaviour, are described from the Upper Jurassic deposits of the Monte Nerone Pelagic Carbonate Platform, in the Umbria‐Marche‐Sabina Palaeogeographic Domain (Italy) by Citton et al. (2019).^[119]
An association of 58 teeth of Ptychodus anonymus, representing the first occurrence of an associated dentition of this species, is described from the Cenomanian Jetmore Member of the Greenhorn Formation (Kansas, United States) by Hamm (2019).^[120]
Description of new associated skeletal remains of Ischyrhiza mira from the Upper Cretaceous of Tennessee and Alabama and a study on the paleobiology of this species is published by Sternes & Shimada (2019).^[121]
A study on the anatomy and phylogenetic relationships of Promyliobatis gazolai is published by Marramà et al. (2019).^[122]
Fossils of members of the genus Aetomylaeus are described from localities in Peru and Chile by Villafaña et al. (2019), representing the first unambiguous fossil record of this genus from the Neogene of the southeastern Pacific.^[123]
A specimen of the whiptail stingray species Tethytrygon muricatus preserving the uterus bearing four eggs is described from the Eocene of the Monte Bolca locality (Italy) by Fanti, Mazzuferi & Marramà (2019).^[124]
Description of Langhian ray fossils from the Lower Tagus Basin (Portugal) is published by Fialho, Balbino & Antues (2019).^[125]
A study on the taxonomic status and geological age of large shark remains from the Upper Cretaceous of the Castellavazzo locality (Italy) discovered in the 19th century is published by Conte et al. (2019).^[126]
15 partial skeletons of lamniform sharks, including the largest specimen of Cretoxyrhina mantelli known to date, are described from the Upper Cretaceous Scaglia Rossa Formation (Italy) by Amalfitano et al. (2019), who also review the taxonomic history of C. mantelli.^[127]
A study on teeth histology and mineralization pattern in lamniform sharks, based on data from extant and fossil taxa (including enigmatic galeomorph shark Palaeocarcharias stromeri), is published by Jambura et al. (2019).^[128]
A study on changes of diversity of lamniform sharks throughout their evolutionary history, aiming to determine the causes of their decline in the last 20 million years, is published by Condamine, Romieu & Guinot (2019).^[129]
A study on changes in the presence or absence of lateral cusplets on teeth of members of the genus Carcharocles from the Calvert, Choptank, and St. Marys formations, and on their implications for the transition of shark populations from these formations from a Carcharocles chubutensis-dominated population to one dominated by C. megalodon, is published by Perez et al. (2019).^[130]
A revision and a reevaluation of the reliability of all post-Messinian occurrences of Otodus megalodon in marine strata from western North America, and a study on the timing of extinction of this species, is published by Boessenecker et al. (2019).^[131]
A study on the body size of Otodus megalodon, as inferred from the relationship between the ontogenetic development of teeth and total body length in the great white shark, is published by Shimada (2019).^[132]
Partial forelimb of a rorqual with several shark bite marks is described from the Pliocene Burica Formation (Panama) by Cortés et al. (2019).^[133]
A study on the anatomy of the holotype specimen of a putative Paleocene shark Platyacrodus unicus is published by Bogan, Agnolin & Ezcurra (2019), who reinterpret this specimen as a carapace of a small retroplumid crab belonging to the genus Costacopluma.^[134]
An assemblage of well-preserved isolated teeth of elasmobranchs is described from the late Oligocene of the North Alpine Foreland Basin (Austria) by Feichtinger et al. (2019).^[135]
Description of the deep-sea elasmobranch fauna from the Miocene Yatsuo Group in central Japan, including the first fossil occurrences of the genera Arhynchobatis and Pseudoraja, will be published by Nishimatsu & Ujihara (2019).^[136]
Description of the deep-sea elasmobranch fossils from the Miocene Makino Formation in southwest Japan, including the first fossil occurrences of the genera Springeria and Narke, is published by Nishimatsu (2019).^[137]
A study on Paleocene cartilaginous fish fossils from the Lower Clayton Limestone Unit of the Midway Group near Malvern, Arkansas, evaluating the implications of these fossils for the knowledge of cartilaginous fish diversity across the Cretaceous-Paleogene boundary in the Malvern region and Gulf Coastal Plain of southwestern Arkansas, is published online by Maisch, Becker & Griffiths (2019).^[138]
A study on the morphology of scales and squamation pattern in Guiyu oneiros is published by Cui, Qiao & Zhu (2019).^[139]
A study on the anatomy and phylogenetic relationships of Brazilichthys macrognathus is published by Figueroa, Friedman & Gallo (2019).^[140]
Permian species Palaeothrissum inaequilobum Blainville (1818) and P. parvum Blainville (1818) are found to be senior synonyms of the widely used species name Aeduella blainvillei (Agassiz, 1833) by Brignon (2019). Conditions exist for reversal of precedence and Aeduella blainvillei is declared nomen protectum.^[141]
A study on the anatomy of the jaws, palate and teeth of Eurynotus crenatus is published by Friedman et al. (2019).^[142]
New specimen of Birgeria liui, representing the most complete and articulated postcranial skeleton of an adult specimen of Birgeria reported so far, is described from the Ladinian of South China by Ni et al. (2019).^[143]
Fossil remains of a spiral valve are reported in a specimen of Peipiaosteus pani from the Lower Cretaceous Yixian Formation (China) by Capasso (2019).^[144]
Teeth of members of the genus Pycnodus are described from the Upper Cretaceous Cap de Naze marine formation (Senegal) by Capasso (2019).^[145]
Description of a partial skeleton of Micropycnodon kansasensis from the Smoky Hill Chalk Member of the Niobrara Chalk (Kansas, United States), providing new information on the anatomy of this species, is published by Cronin & Shimada (2019).^[146]
A study on the paleobiology of specimens of Nursallia gutturosum from the Cenomanian-Turonian platy limestone deposit of Vallecillo (north-eastern Mexico) is published by Stinnesbeck, Rust & Herder (2019).^[147]
A study on the anatomy and phylogenetic relationships of Robustichthys luopingensis is published by Xu (2019).^[148]
Fossil teeth representing the first evidence of Late Jurassic ginglymodians from Mt. Nerone in the Umbria-Marche-Sabina Domain (Italy) are described by Romano et al. (2019).^[149]
A study on the anatomy and phylogenetic relationships of "Lepidotes" bernissartensis is published online by Cavin, Deesri & Olive (2019), who transfer this species to the genus Scheenstia.^[150]
A study on the internal anatomy of a left jaw of a member of the genus Scheenstia from the Kimmeridgian Reuchenette Formation (Switzerland) is published online by Leuzinger et al. (2019), who describe a peculiar tooth replacement mode in this specimen.^[151]
A study on the stomach contents of two specimens of Lepidotes from the Lower Jurassic of Germany is published online by Thies, Stevens & Stumpf (2019).^[152]
Six new occurrences of Belonostomus, documenting the biogeographic and biostratigraphic range of this genus in North America, are reported from the Upper Cretaceous of Texas, Alabama and Mississippi by Van Vranken, Fielitz & Ebersole (2019).^[153]
A study on the diversity of pectoral fin shape amongst members of Pachycormiformes is published by Liston et al. (2019).^[154]
A study on the anatomy of the skull of Martillichthys renwickae is published online by Dobson et al. (2019).^[155]
Fossil remains of a member or a relative of the genus Asthenocormus are described from the Upper Jurassic of the Ameghino (= Nordenskjöld) Formation of the Antarctic Peninsula by Gouiric-Cavalli et al. (2019), representing the first record of a suspension-feeding pachycormid from the Upper Jurassic of the Antarctic Peninsula and the oldest pachycormid yet recovered from Antarctica.^[156]
New, three-dimensionally preserved specimens of Pachycormus are described from the Toarcian of Strawberry Bank at Ilminster (Somerset, United Kingdom) by Cawley et al. (2019).^[157]
A study on the anatomy of Pleuropholis decastroi from the Lower Cretaceous (Albian) limestones of Pietraroja (Province of Benevento, Italy) and on the phylogenetic relationships of the family Pleuropholidae is published by Taverne & Capasso (2019).^[158]
A study on the anatomy and phylogenetic relationships of the teleost species Majokia brasseuri from the Middle Jurassic Stanleyville Formation (Democratic Republic of the Congo) is published by Taverne (2019), who names a new order Majokiiformes.^[159]
A study on the anatomy, jaw mechanics and phylogenetic relationships of Dugaldia emmilta is published by Cavin & Berrell (2019).^[160]
A juvenile specimen of Xiphactinus audax, representing the smallest specimen of this species reported so far, will be described from the Cretaceous Niobrara Chalk (Kansas, United States) by King & Super (2019).^[161]
A study on the internal structure of a fossil specimen of Notelops brama, investigated by neutron tomography, is published by Pugliesi et al. (2019).^[162]
A study on the anatomy and phylogenetic relationships of Cavenderichthys talbragarensis and Waldmanichthys koonwarri is published online by Bean & Arratia (2019).^[163]
A study on fossils of members of the genus Capoeta from the Pliocene locality Çevirme (Turkey) and on the evolutionary history of this genus is published by Ayvazyan, Vasilyan & Böhme (2019).^[164]
Evidence of presence of managed aquaculture of the common carp by around 6000 BC is reported from the Early Neolithic Jiahu site (China) by Nakajima et al. (2019).^[165]
A study on the morphology of fossil catfish spines from the Upper Cretaceous Adamantina and Marilia formations (Brazil) is published by Alves, Bergqvist & Brito (2019).^[166]
A study on positions, heading directions and possible behavioural rules used in a group of fossil specimens of Erismatopterus levatus from the Eocene Green River Formation is published by Mizumoto, Miyata & Pratt (2019).^[167]
Description of gadiform fossils from the Eocene sediments of the Sverdlovsk and Tyumen regions (Ural and Western Siberia, Russia), including fossils of members of the family Merlucciidae, is published by Marramà et al. (2019).^[168]
Description of an incomplete percomorph specimen from the Miocene Ixtapa Formation (Mexico), representing the oldest primary freshwater percomorph fish from Mexico reported so far, is published by Cantalice & Alvarado-Ortega (2019).^[169]
A study on the evolutionary history of the fish clade Pelagiaria is published by Friedman et al. (2019).^[170]
A revision of the nomenclature of extant and fossil barracudas is published by Ballen (2019).^[171]
A review of the published fossil record of the family Labridae is published by Bellwood et al. (2019).^[172]
A study on the feeding habits of the percomorph fish Rhenanoperca minuta and other fishes from the Eocene Messel pit (Germany) is published by Micklich, Baranov & Wappler (2019).^[173]
A study on the phylogenetic relationships of "psarolepid" bony fishes, evaluating which characters cause their different placements in analyses utilizing different methods for reconstructing the tree of life, is published by King (2019).^[174]
A study on the ontogeny of the neurocranium and brain in the West Indian Ocean coelacanth, and on its implications for the knowledge of the evolution of the head of lobe-finned fishes, is published by Dutel et al. (2019).^[175]
Redescription of Axelrodichthys araripensis and a comparative study of several other members of Mawsoniidae is published by Fragoso, Brito & Yabumoto (2019), who transfer the species Mawsonia lavocati to the genus Axelrodichthys.^[176]
A study on the phylogenetic relationships and evolutionary history of mawsoniid coelacanths is published by Cavin et al. (2019).^[177]
A study on the skull anatomy of Arquatichthys porosus, focusing on a newly-discovered postparietal shield, is published by Lu & Zhu (2019).^[178]
Description of the posterior part of the skull of Tungsenia paradoxa is published by Lu et al. (2019).^[179]
New fossil material of "Holoptychius" radiatus Newberry (1889) from the Devonian (Famennian) Catskill Formation (Pennsylvania, United States), providing new information on the anatomy of this species, is described by Daeschler, Downs & Matzko (2019), who transfer this species to the tristichopterid genus Langlieria.^[180]
A study on the bone histology of the humerus of Hyneria lindae is published by Kamska et al. (2019).^[181]
Description of new fossil material of Edenopteron from the Devonian (Famennian) Worange Point Formation (Australia) and a study on the phylogenetic relationships of this taxon is published online by Young et al. (2019).^[182]
A study on the anatomy of the shoulder girdle and opercular series of Gogonasus andrewsae is published by Hu, Young & Lu (2019).^[183]
A historical review of the fossil record of Devonian tetrapods and basal tetrapodomorphs from East Gondwana (Australasia, Antarctica) is published by Long, Clement & Choo (2019), who also present preliminary findings on the anatomy of the canowindrid Koharalepis jarviki based on synchrotron scan data.^[184]
A study on the anatomy of fossil coelacanth lungs, on accessory air-breathing structures in fossil fishes and stem-tetrapods, and on the evolution of air breathing is published by Cupello, Clément & Brito (2019).^[185]
A study on patterns of tooth replacement in Onychodus jandemarrai, Eusthenopteron foordi, Tiktaalik roseae and in extant West Indian Ocean coelacanth is published by Doeland et al. (2019).^[186]
A study on changes of the skeletal anatomy of the pelvic and pectoral appendages during the transition from fins to limbs in vertebrate evolution, as indicated by data from fossil lobe-finned fishes and early tetrapods, is published by Esteve-Altava et al. (2019).^[187]
A study on the anatomy of dermal rays in pectoral fins of Sauripterus taylori, Eusthenopteron foordi and Tiktaalik roseae, evaluating its implications for the knowledge of the evolution of dermal rays in early members of Tetrapodomorpha prior to the origin of digits, is published online by Stewart et al. (2019).^[188]
A study on the evolution of the branchiostegal ray series in the skull of bony fishes, as indicated by data from extant and fossil taxa, is published by Ascarrunz et al. (2019).^[189]
A study on the origin and evolution of acellular bone (bone without osteocytes) in fossil and extant actinopterygian fishes is published by Davesne et al. (2019).^[190]
A diverse fish assemblage is reported from the Carboniferous (upper Pennsylvanian) Horquilla Formation (New Mexico, United States) by Ivanov & Lucas (2019).^[191]
A study on the diversity and ecology of Triassic fish assemblages from the Villány Hills (Hungary) is published by Szabó, Botfalvai & Osi (2019).^[192]
Cretaceous (probably Barremian–Aptian) fish fossils with strong affinities with Early Cretaceous faunas of Thailand are described from Pahang (Malaysia) by Teng et al. (2019).^[193]
Description of a freshwater fish assemblage from the Cretaceous (Albian-Cenomanian) Açu Formation (Brazil) is published by Veiga, Bergqvist & Brito (2019).^[194]
Description of Eocene (Bartonian) fish fauna from the Luna de Sus locality (Romania) is published by Trif, Codrea & Arghiuș (2019).^[195]
A study on the composition of the otolith assemblage from the Santa Barbara Basin near the coast of California over the preceding two millennia is published by Jones & Checkley (2019).^[196]
A study on the evolution of the herbivorous coral reef fishes, as indicated by data from extant and fossil species, is published by Siqueira, Bellwood & Cowman (2019).^[197]

References

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^ Michał Ginter; Stanisław Skompski (2019). "The squamation of "Ctenacanthus" costellatus (Chondrichthyes: Ctenacanthiformes) from the Carboniferous of Lublin area, south-eastern Poland". Acta Geologica Polonica. 69 (4): 571–582. doi:10.24425/agp.2019.126440.
^ John G. Maisey; Philippe Janvier; Alan Pradel; John S. S. Denton; Allison Bronson; Randall Miller; Carole J. Burrow (2019). "Doliodus and pucapampellids. Contrasting perspectives on stem chondrichthyan morphology". In Zerina Johanson; Charlie Underwood; Martha Richter (eds.). Evolution and development of fishes. Cambridge University Press. pp. 87–109. doi:10.1017/9781316832172.006. ISBN 978-1-107-17944-8. S2CID 92398530.
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[EJES683cyathaspidids-1] Tiiu Märss (2019). "Silurian cyathaspidid heterostracans of Northern Eurasia". Estonian Journal of Earth Sciences. 68 (3): 113–146. doi:10.3176/earth.2019.11. S2CID 202181325.

[Jiaoyu-2] Yu-Hai Liu; Min Zhu; Xiang-Hong Lin; Li-Wu Lu; Zhi-Kun Gai (2019). "A reappraisal of the Silurian galeaspids (stem-Gnathostomata) from Tarim Basin, Xinjiang". Vertebrata PalAsiatica. 57 (4): 253–273. doi:10.19615/j.cnki.1000-3118.190524.

[VPthelodonts-3] Xin-Dong Cui; Qiang Li; Tuo Qiao; Min Zhu (2020). "New material of thelodonts from Lochkovian (Lower Devonian) of Qujing, Yunnan, China". Vertebrata PalAsiatica. 58 (1): 1–15. doi:10.19615/j.cnki.1000-3118.190612.

[4] Tetsuto Miyashita; Michael I. Coates; Robert Farrar; Peter Larson; Phillip L. Manning; Roy A. Wogelius; Nicholas P. Edwards; Jennifer Anné; Uwe Bergmann; A. Richard Palmer; Philip J. Currie (2019). "Hagfish from the Cretaceous Tethys Sea and a reconciliation of the morphological–molecular conflict in early vertebrate phylogeny". Proceedings of the National Academy of Sciences of the United States of America. 116 (6): 2146–2151. Bibcode:2019PNAS..116.2146M. doi:10.1073/pnas.1814794116. PMC 6369785. PMID 30670644.

[5] Jason P. Downs; Edward B. Daeschler; Nathanael Lo; Emily N. Carey; Neil H. Shubin (2019). "Asterolepis alticristata n. sp. (Antiarchi) from the Upper Devonian (Frasnian) of Nunavut, Canada, and a report on the antiarch diversity of the Fram Formation". Geodiversitas. 41 (19): 679–698. doi:10.5252/geodiversitas2019v41a19. S2CID 210624573.

[Colombialepis-6] Sébastien Olive; Alan Pradel; Carlos Martinez-Pérez; Philippe Janvier; James C. Lamsdell; Pierre Gueriau; Nicolas Rabet; Philippe Duranleau-Gagnon; Andres L. Cardenas-Rozo; Paula A. Zapata Ramirez; Héctor Botella (2019). "New insights into Late Devonian vertebrates and associated fauna from the Cuche Formation (Floresta Massif, Colombia)". Journal of Vertebrate Paleontology. 39 (3): e1620247. doi:10.1080/02724634.2019.1620247. S2CID 198237241.

[7] Kate Trinajstic; John A. Long; Alexander O. Ivanov; Elga Mark-Kurik (2019). "A new genus of ptyctodont (Placodermi) from the Late Devonian of Baltic area". Palaeontologia Electronica. 22 (2): Article number 22.2.23. doi:10.26879/890. S2CID 181769139.

[8] Jan L. den Blaauwen; Michael J. Newman; Carole J. Burrow (2019). "A new cheiracanthid acanthodian from the Middle Devonian (Givetian) Orcadian Basin of Scotland and its biostratigraphic and biogeographical significance". Scottish Journal of Geology. 55 (2): 166–177. doi:10.1144/sjg2018-023. S2CID 189990496.

[9] Michael J. Newman; Carole J. Burrow; Jan L. den Blaauwen (2019). "A new species of ischnacanthiform acanthodian from the Givetian of Mimerdalen, Svalbard". Norwegian Journal of Geology. 99 (4): 619–631. doi:10.17850/njg99-4-05. S2CID 214220714.

[10] Sebastian Stumpf; Jürgen Kriwet (2019). "A new Pliensbachian elasmobranch (Vertebrata, Chondrichthyes) assemblage from Europe, and its contribution to the understanding of late Early Jurassic elasmobranch diversity and distributional patterns". PalZ. 93 (4): 637–658. doi:10.1007/s12542-019-00451-4. S2CID 181782998.

[11] Eduardo Villalobos-Segura; Charlie J. Underwood; David J. Ward; Kerin M. Claeson (2019). "The first three-dimensional fossils of Cretaceous sclerorhynchid sawfish: Asflapristis cristadentis gen. et sp. nov., and implications for the phylogenetic relations of the Sclerorhynchoidei (Chondrichthyes)" (PDF). Journal of Systematic Palaeontology. 17 (21): 1847–1870. doi:10.1080/14772019.2019.1578832. S2CID 145940997.

[12] Gerard R. Case; Todd D. Cook; Taylor Kightlinger; Paul D. Borodin (2019). "Middle Campanian euselachian diversity of the southern region of the Atlantic Coastal Plain of North America". Vertebrate Anatomy Morphology Palaeontology. 7: 69–82. doi:10.18435/vamp29345. S2CID 201576213.

[EJT585-13] Jun A. Ebersole; David J. Cicimurri; Gary L. Stringer (2019). "Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths". European Journal of Taxonomy (585): 1–274. doi:10.5852/ejt.2019.585. S2CID 213997330.

[14] Karen E. Samonds; Tsiory H. Andrianavalona; Lane A. Wallett; Iyad S. Zalmout; David J. Ward (2019). "A middle - late Eocene neoselachian assemblage from nearshore marine deposits, Mahajanga Basin, northwestern Madagascar". PLOS ONE. 14 (2): e0211789. Bibcode:2019PLoSO..1411789S. doi:10.1371/journal.pone.0211789. PMC 6392239. PMID 30811516.

[Dykeius-15] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r Henri Cappetta; Kurt Morrison; Sylvain Adnet (2019). "A shark fauna from the Campanian of Hornby Island, British Columbia, Canada: an insight into the diversity of Cretaceous deep-water assemblages". Historical Biology: An International Journal of Paleobiology. 33 (8): 1121–1182. doi:10.1080/08912963.2019.1681421. S2CID 212878837.

[BAP398-16] John W. Hoganson; J. Mark Erickson; F. D. Holland, Jr. (2019). "Chondrichthyan and osteichthyan paleofaunas from the Cretaceous (Late Maastrichtian) Fox Hills Formation of North Dakota, USA: Paleoecology, paleogeography, and extinction" (PDF). Bulletins of American Paleontology. 398: 1–94. doi:10.32857/bap.2019.398.

[17] Kenshu Shimada; Michael J. Everhart (2019). "A New Large Late Cretaceous Lamniform Shark from North America, with Comments on the Taxonomy, Paleoecology, and Evolution of the Genus Cretodus". Journal of Vertebrate Paleontology. 39 (4): e1673399. doi:10.1080/02724634.2019.1673399. S2CID 209439997.

[18] Sebastian Stumpf; Udo Scheer; Jürgen Kriwet (2019). "A new genus and species of extinct ground shark, †Diprosopovenator hilperti, gen. et sp. nov. (Carcharhiniformes, †Pseudoscyliorhinidae, fam. nov.), from the Upper Cretaceous of Germany". Journal of Vertebrate Paleontology. 39 (2): e1593185. doi:10.1080/02724634.2019.1593185. S2CID 155785248.

[19] Terry A. Gates; Eric Gorscak; Peter J. Makovicky (2019). "New sharks and other chondrichthyans from the latest Maastrichtian (Late Cretaceous) of North America". Journal of Paleontology. 93 (3): 512–530. doi:10.1017/jpa.2018.92. S2CID 198159821.

[20] Michał Ginter; Stanisław Skompski (2019). "The squamation of "Ctenacanthus" costellatus (Chondrichthyes: Ctenacanthiformes) from the Carboniferous of Lublin area, south-eastern Poland". Acta Geologica Polonica. 69 (4): 571–582. doi:10.24425/agp.2019.126440.

[21] John G. Maisey; Philippe Janvier; Alan Pradel; John S. S. Denton; Allison Bronson; Randall Miller; Carole J. Burrow (2019). "Doliodus and pucapampellids. Contrasting perspectives on stem chondrichthyan morphology". In Zerina Johanson; Charlie Underwood; Martha Richter (eds.). Evolution and development of fishes. Cambridge University Press. pp. 87–109. doi:10.1017/9781316832172.006. ISBN 978-1-107-17944-8. S2CID 92398530.

[22] Harry M. Maisch (2019). "A new species of Hypolophites (Chondrichthyes, Myliobatiformes) from the Lower Clayton Limestone Unit of the Midway Group (Paleocene), near Malvern, Arkansas, USA". Journal of Paleontology. 94 (3): 548–556. doi:10.1017/jpa.2019.99. S2CID 214211012.

[23] David J. Cicimurri; James L. Knight (2019). "Late Eocene (Priabonian) elasmobranchs from the Dry Branch Formation (Barnwell Group) of Aiken County, South Carolina, USA". PaleoBios. 36: ucmp_paleobios_43964.

[24] Charlie J. Underwood; Kerin M. Claeson (2019). "The Late Jurassic ray Kimmerobatis etchesi gen. et sp. nov. and the Jurassic radiation of the Batoidea" (PDF). Proceedings of the Geologists' Association. 130 (3–4): 345–354. doi:10.1016/j.pgeola.2017.06.009. S2CID 90691006.

[25] Giuseppe Marramà; Giorgio Carnevale; Luca Giusberti; Gavin J. P. Naylor; Jürgen Kriwet (2019). "A bizarre Eocene dasyatoid batomorph (Elasmobranchii, Myliobatiformes) from the Bolca Lagerstätte (Italy) reveals a new, extinct body plan for stingrays". Scientific Reports. 9 (1): Article number 14087. Bibcode:2019NatSR...914087M. doi:10.1038/s41598-019-50544-y. PMC 6773687. PMID 31575915.

[26] Rodrigo A. Otero (2019). "Myledaphus araucanus sp. nov. (Batomorphi, Rajiformes incertae sedis), a new Late Cretaceous ray from the austral Pacific, and first occurrence of the genus in the Southern Hemisphere". Cretaceous Research. 100: 82–90. doi:10.1016/j.cretres.2019.03.025. S2CID 134755626.

[27] Iman Salame; Anhar Asan (2019). "A new Odontorhytis species (Chondrichthyes) from the Middle Eocene of ElGedida mine, Bahariya oasis, Egypt". Egyptian Journal of Geology. 63: 407–415.

[28] Evgeny V. Popov; Dominique Delsate; Roland Felten (2019). "A new callorhinchid genus (Holocephali, Chimaeroidei) from the early Bajocian of Ottange-Rumelange, on the Luxembourg-French border". Paleontological Research. 23 (3): 220–230. doi:10.2517/2018PR021. S2CID 198423356.

[29] Sylvain Adnet; Medhi Mouana; Anne-Lise Charruault; El Mebrouk Essid; Hayet Khayati Ammar; Wissem Marzougui; Gilles Merzeraud; Rodolphe Tabuce; Monique Vianey-Liaud; Laurent Marivaux (2019). "Teeth, fossil record and evolutionary history of the cowtail stingray Pastinachus Rüppell, 1829" (PDF). Historical Biology: An International Journal of Paleobiology. 31 (9): 1213–1222. doi:10.1080/08912963.2018.1431779. S2CID 90521628.

[30] Jaime A. Villafaña; Sven N. Nielsen; Stefanie Klug; Jürgen Kriwet (2019). "Early Miocene cartilaginous fishes (Chondrichthyes: Holocephali, Elasmobranchii) from Chile: Diversity and paleobiogeographic implications". Journal of South American Earth Sciences. 96: Article 102317. Bibcode:2019JSAES..9602317V. doi:10.1016/j.jsames.2019.102317. S2CID 202190340.

[31] Jorge D. Carrillo-Briceño; Juan Parra; Javier Luque (2019). "A new lamniform shark Protolamna ricaurtensis sp. nov. from the Lower Cretaceous of Colombia" (PDF). Cretaceous Research. 95: 336–340. doi:10.1016/j.cretres.2018.12.007. S2CID 134390711.

[32] Eduardo Villalobos‐Segura; Charlie J. Underwood; David J. Ward (2019). "The first skeletal record of the enigmatic Cretaceous sawfish genus Ptychotrygon (Chondrichthyes, Batoidea) from the Turonian of Morocco" (PDF). Papers in Palaeontology. 7 (1): 353–376. doi:10.1002/spp2.1287. S2CID 210302939.

[33] Kouki Nishimatsu; Atsushi Ujihara (2019). "A new deep-sea shark Scymnodalatias kazenobon (Squaliformes, Somniosidae) from the Miocene Yatsuo Group in Central Japan". Paleontological Research. 23 (1): 23–29. doi:10.2517/2018PR006. S2CID 135131261.

[34] Giuseppe Marramà; Giorgio Carnevale; Gavin J. P. Naylor; Jürgen Kriwet (2019). "Reappraisal of the Eocene whiptail stingrays (Myliobatiformes, Dasyatidae) of the Bolca Lagerstätte, Italy". Zoologica Scripta. 48 (2): 168–184. doi:10.1111/zsc.12330. PMC 6446806. PMID 30983683.

[35] Paulo M. Brito; Eduardo Villalobos-Segura; Jésus Alvarado-Ortega (2019). "A new Early Cretaceous guitarfish (Chondrichthyes, Batoidea) from the Tlayúa Formation, Puebla, Mexico". Journal of South American Earth Sciences. 90: 155–161. Bibcode:2019JSAES..90..155B. doi:10.1016/j.jsames.2018.12.005. S2CID 135426155.

[36] Alexandre F. Bannikov; Giorgio Carnevale; James C. Tyler (2019). "Acronuroides eocaenicus gen. et sp. nov., a new percomorph fish from the Eocene of Bolca, Italy" (PDF). Studi e Ricerche Sui Giacimenti Terziari di Bolca, XIX - Miscellanea Paleontologica. 16: 29–37.

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[Mystocheilus-38] Gerald R. Smith; Jay Van Tassell (2019). "Fishes of the Mio-Pliocene Western Snake River Plain and vicinity. V. Keating, Always Welcome Inn, and Imbler Fish paleofaunas, NE Oregon: Tests of Miocene-Pliocene drainage connections". Miscellaneous Publications. Museum of Zoology, University of Michigan. 204 (5): 1–33. hdl:2027.42/150494.

[Ardoreosomus-39] Carlo Romano; Adriana López-Arbarello; David Ware; James F. Jenks; Winand Brinkmann (2019). "Marine Early Triassic Actinopterygii from the Candelaria Hills (Esmeralda County, Nevada, USA)". Journal of Paleontology. 93 (5): 971–1000. doi:10.1017/jpa.2019.18. S2CID 155564297.

[PJ532-40] M. V. Nazarkin (2019). "Hooked-horn sculpins (Pisces: Cottidae, Artediellinae) from the Miocene of Sakhalin Island, Russia". Paleontological Journal. 53 (2): 172–182. doi:10.1134/S0031030119020084. S2CID 195302343.

[41] M. V. Nazarkin (2019). "Tubesnout Aulichthys miocaenicus sp. nov. from the Miocene of Sakhalin Island (Russia): the first reliable fossil record for the family Aulorhynchidae". Paleontological Journal. 53 (5): 522–529. doi:10.1134/S0031030119040105. S2CID 203848519.

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[43] Lazaro W. Viñola Lopez; Richard Carr; Logel Lorenzo (2019). "First occurrence of fossil Balistes (Tetradontiformes: Balistidae) from the Miocene of Cuba with the description of a new species and a revision of fossil Balistes". Historical Biology: An International Journal of Paleobiology. 32 (9): 1290–1299. doi:10.1080/08912963.2019.1580278. S2CID 92003143.

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[92] Anne Kemp (2019). "Ctenodus boudariensis nov. sp., a ctenodont lungfish from the mid-Viséan Ducabrook Formation of Queensland, Australia". Alcheringa: An Australasian Journal of Palaeontology. 43 (3): 423–429. doi:10.1080/03115518.2019.1587786. S2CID 149997652.

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