List of ichthyosauromorph genera: Difference between revisions

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''[[Acamptonectes]]''<ref name=Acamptonectes>{{cite journal |authors=Fischer,V.; Maisch, M.W.; Naish, D.; Kosma, R.; Liston, J.; Joger, U.; Krüger, F.J.; Pérez,J.P.; Tainsh, J.; Appleby, R.M.|year=2012 |title=New Ophthalmosaurid Ichthyosaurs from the European Lower Cretaceous Demonstrate Extensive Ichthyosaur Survival across the Jurassic–Cretaceous Boundary |journal=PLoS ONE |volume=7 |issue=1 |pages=e29234 |doi=10.1371/journal.pone.0029234 }}</ref>
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Fischer<br/>
et al.<br/>
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[[2012 in paleontology|2012]]
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Valid.
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[[Lower Cretaceous|lK]]<ref name=Acamptonectes/><br/>
to</br>
[[Upper Cretaceous|uK]].<ref name=Acamptonectes/><br/>
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A strange [[Middle Triassic|mT]] [[mixosaurid]] with a high crest of bone on its head and a 5 meter body. It was thought to be a ''[[Mixosaurus]]'' species for some time until given its own genus in [[1998 in paleontology|1998]]. Some experts have felt that the splitting was unjustified.
A strange [[Middle Triassic|mT]] [[mixosaurid]] with a high crest of bone on its head and a 5 meter body. It was thought to be a ''[[Mixosaurus]]'' species for some time until given its own genus in [[1998 in paleontology|1998]]. Some experts have felt that the splitting was unjustified.
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''[[Cryopterygius]]''<ref name=DruckenmillerHurumKnutsenandNakrem>{{Cite journal|author=Patrick S. Druckenmiller, Jørn H. Hurum, Espen M. Knutsen and Hans Arne Nakrem |year=2012 |title=Two new ophthalmosaurids (Reptilia: Ichthyosauria) from the Agardhfjellet Formation (Upper Jurassic: Volgian/Tithonian), Svalbard, Norway |journal=Norwegian Journal of Geology |volume=92 |issue=2–3 |pages=311–339 |url=http://www.geologi.no/data/f/0/21/00/9_2401_0/NJG_2_3_2012_17_Druckenmiller_etal_Pr.pdf }}</ref>
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Druckenmiller<br/>
et al.</br>
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[[2012 in paleontology|2012]]<ref name=DruckenmillerHurumKnutsenandNakrem/>
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[[Upper Jurassic|uJ]].<ref name=DruckenmillerHurumKnutsenandNakrem/>
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Junior synonym of ''[[Brachypterygius]]''.
Junior synonym of ''[[Brachypterygius]]''.
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''[[Palvennia]]''
<ref name=DruckenmillerHurumKnutsenandNakrem/>
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Druckenmiller</br>
et al.</br>
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[[2012 in paleontology|2012]]
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Valid.
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[[Upper Jurassic|uJ]].<ref name=DruckenmillerHurumKnutsenandNakrem/>
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Non-ichthyosaur ichthyopterygian.
Non-ichthyosaur ichthyopterygian.
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|''[[Thalattoarchon]]''<ref name=thalattoarchon>{{cite journal |authors=Nadia B. Fröbisch, Jörg Fröbisch, P. Martin Sander, Lars Schmitz and Olivier Rieppel |year=2013 |title=Macropredatory ichthyosaur from the Middle Triassic and the origin of modern trophic networks |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=in press |issue= |pages= |doi=10.1073/pnas.1216750110 }}</ref>
|Fröbisch<br/>
et al.</br>
|[[2013 in paleontology|2013]]
|Valid.
|[[Middle Triassic|mT]].<ref name=thalattoarchon/>
|[[North America]].<ref name=thalattoarchon/>
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Revision as of 01:25, 12 January 2013

This list of ichthyosaurs is a comprehensive listing of all genera that have ever been included in the order Ichthyosauria or the parent clade Ichthyopterygia, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names, and genera that are no longer considered ichthyopterygian. Non-ichthyosaur ichthyopterygians shall be noted as such.

Scope and terminology

There is no official, canonical list of ichthyosaur genera but one of the most thorough attempts can be found at the "Ichthyosauromorpha" section of Mikko Haaramo's Phylogeny Archive.[1]

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

  • Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
  • Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen. In this case, these nomina nuda will be deleted from this list in favor of the published name.
  • Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
  • Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.

The List

Genus Authors Year Status Age Location Notes

Acamptonectes[2]

Fischer
et al.

2012

Valid.

lK[2]
to
uK.[2]

Europe[2]

Actiosaurus

Sauvage

1883

Disputed.

Europe

Aegirosaurus[3]

Bardet
Fernández

2000

Valid.

uJ.[3]

Europe.[3]

A 2 meter long[3] ophthalmosaurid[4] whose remains were discovered in the Solnhofen limestone and mistaken for a new Ichthyosaurus species. In 2000 scientists concluded that the fossil material should be assigned to a new genus.

Anhuisaurus[1]

Chen

1985

Jr. synonym.

N/A

N/A

Preoccupied by lizard genus Anhuisaurus Hou, 1974. Junior synonym of Chaohusaurus.

Arthropterygius

Maxwell

2010

Valid.

uJ.

North America

Athabascasaurus

Druckenmiller
Maxwell

2010

Valid.

lK.

North America

Barracudasauroides

Maisch

2010

Valid.

mT.

Asia

Besanosaurus[5]

Dal Sasso
Pinna

1996

Valid.

mT.[5]

Europe.[5]

Brachypterygius[6][7]

von Huene

1922

Valid.

uJ.[6]

Europe.[8][9]

A uJ Ophthalmosaurid closely related to Platypterygius and Caypullisaurus.[4] Fossils have been found in England and European Russia.[8][9]

Californosaurus[6][10]

Kuhn

1934

Valid.

uT.[6]

N. America.

A 3 meter teretocnemid who is also the basal-most known Euichthyosaurian. Its remains were first recovered from the Carnian Lower Hosselkus Limestone of California. Californosaurus has also been known as Shastasaurus perrini and Delphinosaurus perrini.

Callawayia[11]

Maische
Matzke

2000

Valid.

Caypullisaurus[6][7]

Fernández

1997

Valid.

uJ.[6]

to

lK.[citation needed]

S. America.

A large ophthalmosaurid closely related to Platypterygius and Brachypterygius.[4] It was discovered in strata dating to the uJ - lK of Argentina.

Chacaicosaurus[10]

Fernández

1994

Valid.

S. America.[9]

A Bajocian genus whose fossils have been found in Argentina.[9] Possibly a stenopterygiid.

Chaohusaurus[1][6]

Young
Dong

1972

Valid.

lT.[6]

Asia.

A primitive ichthyosaur that retained a lizard-like body discoverered in Chaohu, China. It was also one of the smallest ichthyosaurs, being from 70 to 180 cm long and an estimated weight of 10 kg.

Chensaurus[1][6]

Mazin
Suteethorn
Buffetaut
Jaeger
Helmcke-Ingavat

1991

Jr. synonym.

N/A

N/A

Junior synonym of Chaohusaurus.

Chonespondylus[6]

Leidy

1868

Valid.

mT.[6]

Contectopalatus[12]

Maische
Matzke

1998

Disputed.

mT.[12]

Europe.[12]

A strange mT mixosaurid with a high crest of bone on its head and a 5 meter body. It was thought to be a Mixosaurus species for some time until given its own genus in 1998. Some experts have felt that the splitting was unjustified.

Cryopterygius[13]

Druckenmiller
et al.

2012[13]

Valid.

uJ.[13]

Europe.[13]

Cymbospondylus[6][11]

Leidy

1868

Valid.

lT.[6]

to

mT.[6]

One of the largest ichthyosaurs, adults ranged in length from 6 m (18 ft) up to 10 m (30 ft) long. It was one of the least fish-like of the ichthyosaurs, lacking a dorsal fin and fluked tail. Consequently, Cymbospondylus probably swam by wriggling its body from side to side like a modern sea snake.

Delphinosaurus[6]

Merriam

1905

Jr. synonym.

N/A

N/A

Junior synonym of Californosaurus.

Eurhinosaurus[6][10]

Abel

1909

Valid.

lJ.[6]

Europe.

A genus exceeding 6 m in length[9] from the Sinemurian to Toarcian of England and Germany. Eurhinosaurus was built like a regular ichthyosaur, with a fish-like body including a large eyes, dorsal and caudal fins, but had one distinct feature that set it apart from other ichthyosaurs; its upper jaw was twice as long as the lower jaw and covered with sidewards-pointing 'teeth', like a sawfish's.

Eurypterygius

Jaekel

1904

Jr. synonym.

N/A

N/A

Junior synonym of Ichthyosaurus.

Excalibosaurus

McGowan

1986

Valid.

lJ.[6]

Lived during the Sinemurian stage in what is now England. It is characterized by the extreme elongation of the rostrum, giving the animal a swordfish-like look. Excalibosaurus is known from two skeletons. The holotype was from a juvenile with an estimated body length of 4 m. The estimated body length of the second specimen, discovered in 1996, is 7 m.

Grendelius[6]

McGowan

1976

Jr. synonym.

N/A

N/A

Junior synonym of Brachypterygius.

Grippia[1][6]

Wiman

1930

Valid.

lT.[6]

Asia.
Greenland.
Japan.
N. America.

A small ichthyosaur only 1-1.5 m long, its fossils have been found along the coasts of Greenland, China, Japan, and Canada.

Guizhouichthyosaurus

Cao
Luo

2000

Jr. synonym.

N/A

N/A

Junior synonym of Shastasaurus.[14]

Himalayasaurus

Young
Dong

1972

Jr. synonym.

N/A

N/A

Junior synonym of Shonisaurus.

Hudsonelpidia

McGowan

1995

Valid.

uT.[6]

Hupehsuchus[1]

Carroll
Dong

1991

Valid.

mT.

Asia.

Non-ichthyosaur ichthyopterygian from the mT of China. The type species is H. nanchangensis.

Ichthyosaurus[6][10]

De la Beche
Conybeare

1821

Valid.

lJ.[6]

Isfjordosaurus

Motani

1999

Valid.

Non-ichthyosaur ichthyopterygian.

Leptocheirus

Merriam

1903

Valid. [dubious ]

Leptonectes

McGowan

1996

Valid.

Europe.[9]

Lived from the Rhaetian to Pliensbachian in what is now Belgium, England and Germany.[9]

Leptopterygius

Huene

1922

Valid.

uT.[6] to lJ.[6]

Macgowania

Motani

1999

Valid.

Macropterygius

Huene

1923

Jr. synonym.

N/A

N/A

Junior synonym of Aegirosaurus.

Maiaspondylus[7]

Maxwell
Caldwell

2006

Valid.

uK.[7]

N. America.[7]

Merriamia[6]

Boulenger

1904

Jr. synonym. N/A N/A

Junior synonym of Toretocnemus.

Metashastasaurus

Nicholls
Manabe

2001

Jr. synonym. N/A N/A

Junior synonym of Callawayia.

Mikadocephalus[11][15]

Maisch
Matzke

1998

Valid.

Europe.[15]

Mixosaurus[6][11]

Baur

1887

Valid.

mT.[6]

Mollesaurus

Fernandez

1999

Jr. synonym. N/A N/A

Junior synonym of Ophthalmosaurus.

Myopterygius

von Huene

1922

Syn.

Synonymy of Platypterygius

Nanchangosaurus

Wang

1959

Valid.

mT.[6]

Nannopterygius

Huene

1922

Valid.

uJ.[6]

Europe.

Nannopterygius is a genus of ophthalmosaurid ichthyosaur that lived in from the Kimmeridgian to Tithonian. Fossils have been found in England and Germany.[9]

Omphalosaurus

Merriam

1906

Valid.

lT.

Europe.

An extinct genus of marine reptile from Spitsbergen. Is has been considered an ichthyopterygian, however a study by Motani in 2000 showed that it lacks the basal synapomorphies of Ichthyopterygia. As such, its exact position in the diapsid tree is currently unknown.

Ophthalmosaurus[7]

Seeley

1874

Valid.

mJ.[6]

to

uJ.[6]

Europe.[citation needed]
N. America.[citation needed]
S. America.[7]

Otschevia

Efimov

1997

Jr. synonym. N/A N/A

Junior synonym of Brachypterygius.

Palvennia [13]

Druckenmiller
et al.

2012

Valid.

uJ.[13]

Europe.[13]

Panjiangsaurus

Chen
Chang

2003

Jr. synonym.

N/A

N/A

Junior synonym of Shastasaurus.[14]

Paraophthalmosaurus

Arkhangelsky

1997

Jr. synonym. N/A N/A

Junior synonym of Ophthalmosaurus.

Parvinatator

Nicholls
Brinkman

1995

Valid.

Non-ichthyosaur ichthyopterygian.

Pessopteryx

Wiman

1910

Valid.

Pessosaurus

Wiman

1910

Valid.

mT.[6]

Phalarodon

Merriam

1910

Valid.

mT.[6]

Asia.[16]

Phantomosaurus[6]

Maisch
Matzke

2000

Valid.

Platypterygius

Kuhn

1946

Valid.

uK.[6]

Plutonisaurus

Efimov

1997

Jr. synonym. N/A N/A

Junior synonym of Platypterygius.

Proteosaurus

Home

1819

Jr. synonym. N/A N/A

Junior synonym of Ichthyosaurus.

Protoichthyosaurus

Appleby

1979

Jr. synonym. N/A N/A

Junior synonym of Ichthyosaurus.

Qianichthyosaurus

Li

1999

Valid.

Sangiorgiosaurus

Brinkmann

1998

Valid.

Shastasaurus

Merriam

1895

Valid.

mT[14] to uT.[6]

Asia.
Europe.
N. America.

Shastasaurus is a genus of ichthyosaur from whose remains have been found in United States, Canada, Mexico, Austria, Italy, Switzerland, Germany, Spitsbergen and China.[14]

Shonisaurus

Camp

1976

Valid.

uT.[6]

?Asia.
N. America.

The largest genus of ichthyosaur that has yet been found. Fossils of Shonisaurus were first found in Nevada in 1920. It lived during the Norian stage of the late Triassic period and had a long pointed mouth that contained teeth only at the front end. S. popularis specimens reached a length of 15 meters (50 feet). A second species, S. sikanniensis, was later discovered in British Columbia and has an estimated length of 21 meters (70 feet).

Simbirskia

Otschev
Efimov

1985

Valid.

Simbirskiasaurus

Otschev
Efimov

1985

Valid.

lK.[6]

Stenopterygius

Jaekel

1904

Valid.

lJ.[6]

Europe.

Lived from the Toarcian to Aalenian from England, France, Germany and Luxembourg. Maximum length was 4 m.[9] One famous fossil is that of a mother and baby that died in childbirth. This proved that ichthyosaur infants were born tail-first, just like cetaceans, to prevent them from drowning before fully clearing the birth canal.

Suevoleviathan

Maisch

1998

Valid.

Svalbardosaurus

Mazin

1981

Nomen dubium.[1]

lT.[6]

Sveltonectes

Fischer
Masure
Arkhangelsky
Godefroit

2011

Valid.

lK.

Europe.

Temnodontosaurus

Lydekker

1889

Valid.

lJ.[6]

Europe.

Lived from the Hettangian to Toarcian in England and Germany. It was a large genus, exceeded 12 meters (30 ft) in length.[9] Temnodontosaurus eyes were approximately 20 cm (8 in) in diameter making them the largest of any known vertebrate[citation needed].

Thaisaurus

Mazin
Sutetthorn
Buffetaut
Jaeger
Helmcke-Ignavat

1991

Valid.

Non-ichthyosaur ichthyopterygian.

Thalattoarchon[17] Fröbisch

et al.

2013 Valid. mT.[17] North America.[17]

Toretocnemus

Merriam

1902

Valid.

uT.[6]

Undorosaurus

Efimov

1997

Jr. synonym. N/A N/A

Junior synonym of Ophthalmosaurus.

Utatsusaurus[1][6]

Shikoma

1978

Valid.

lT.[6]

Non-ichthyosaur ichthyopterygian.

Wimanius

Maisch
Matzke

1988

Valid.
Besanosaurus
Californosaurus.
Caypullisaurus.
Chaohusaurus.
Cymbospondylus.
Eurhinosaurus.
Excalibosaurus.
Grippia.
Hupehsuchus.
Ichthyosaurus.
Leptonectes.
Mixosaurus.
Ophthalmosaurus.
Platypterigius.
Shastasaurus.
Shonisaurus.
Stenopterygius.
Temnodontosaurus.
Utatsusaurus.

See also

References

  1. ^ a b c d e f g h Haaramo, Mikko (2004). "Ichthyosauromorpha". Mikko's Phylogeny Archive. Retrieved 9 October 2008. {{cite web}}: Unknown parameter |month= ignored (help)
  2. ^ a b c d "New Ophthalmosaurid Ichthyosaurs from the European Lower Cretaceous Demonstrate Extensive Ichthyosaur Survival across the Jurassic–Cretaceous Boundary". PLoS ONE. 7 (1): e29234. 2012. doi:10.1371/journal.pone.0029234. {{cite journal}}: Cite uses deprecated parameter |authors= (help)CS1 maint: unflagged free DOI (link)
  3. ^ a b c d Bardet N, Fernández M. 2000. A new ichthyosaur from the Upper Jurassic lithographic limestones of Bavaria. Journal of Paleontology 74 (3): 503-511.
  4. ^ a b c Fernández, M. (2007). "Redescription and phylogenetic position of Caypullisaurus (Ichthyosauria: Ophthalmosauridae)". Journal of Paleontology. 81 (2): 368–375. doi:10.1666/0022-3360(2007)81[368:RAPPOC]2.0.CO;2.
  5. ^ a b c Dal Sasso, Cristiano and Giovanni Pinna, 1996. Besanosaurus leptorhynchus n. gen. n. sp., a new shastasaurid ichthyosaur from the Middle Triassic of Besano (Lombardy, N. Italy). Paleontologia Lombarda, Nuova seire Volume IV: 1-23.
  6. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at Sepkoski, Jack (2002). "A compendium of fossil marine animal genera (entry on Reptilia)". Bulletins of American Paleontology. 364: p.560. Retrieved 2008-09-28. {{cite journal}}: |pages= has extra text (help)
  7. ^ a b c d e f g Haaramo, Mikko (2008). "Ophthalmosauria". Mikko's Phylogeny Archive. Retrieved 9 October 2008. {{cite web}}: Unknown parameter |month= ignored (help)
  8. ^ a b Efimov, VM. (1998). "An Ichthyosaur, Otschevia pseudoscythica gen. et sp. nov. from the Upper Jurassic Strata of the Ulyanovsk Region (Volga Region)". Paleontologicheskii Zhurnal. 32 (2): 187–191.
  9. ^ a b c d e f g h i j Maisch, MW; Matzke, AT. (2000). "The Ichthyosauria". Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie). 298: 1–159.
  10. ^ a b c d Haaramo, Mikko (2004). "Euichthyosauria". Mikko's Phylogeny Archive. Retrieved 9 October 2008. {{cite web}}: Unknown parameter |month= ignored (help)
  11. ^ a b c d Haaramo, Mikko (2008). "Ichthyosauria". Mikko's Phylogeny Archive. Retrieved 9 October 2008. {{cite web}}: Unknown parameter |month= ignored (help)
  12. ^ a b c Maisch & Matzke, A. T. 2000. The mixosaurid ichthyosaur Contectopalatus from the Middle Triassic of the German Basin. Lethaia 33: 71-74.
  13. ^ a b c d e f g Patrick S. Druckenmiller, Jørn H. Hurum, Espen M. Knutsen and Hans Arne Nakrem (2012). "Two new ophthalmosaurids (Reptilia: Ichthyosauria) from the Agardhfjellet Formation (Upper Jurassic: Volgian/Tithonian), Svalbard, Norway" (PDF). Norwegian Journal of Geology. 92 (2–3): 311–339.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  14. ^ a b c d Shang Qing-Hua, Li Chun (2009). "On the occurrence of the ichthyosaur Shastasaurus in the Guanling Biota (Late Triassic), Guizhou, China" (PDF). Vertebrata PalAsiatica. 47 (3): 178–193.
  15. ^ a b Maisch, M. W,. and Matzke A. T., 1998, Mikadocephalus gracilirostris n. gen., n. sp., a new ichthyosaur from the Grenzbitumenzone (Anisian-Ladinian) of Mont San Giorgio (Switzerland): Palaontologische Zeitschrift, v. 71, n. ¾, p. 267-289.
  16. ^ Jiang, D.-Y.; Schmitz, L.; Motani, R.; Hao, W.-C.; Sun, Y.-L. (2007). "The mixosaurid ichthyosaur Phalarodon cf. P. fraasi from the Middle Triassic of Guizhou Province, China". Journal of Paleontology. 81 (3): 602–605. doi:10.1666/05092.1.
  17. ^ a b c "Macropredatory ichthyosaur from the Middle Triassic and the origin of modern trophic networks". Proceedings of the National Academy of Sciences of the United States of America. in press. 2013. doi:10.1073/pnas.1216750110. {{cite journal}}: Cite uses deprecated parameter |authors= (help)

External links