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→‎Genetic studies: There are so many great papers that are not included on this Wikipedia page dealing with Ethiopians, we should include the latest ones. Same with the Somalis genetics section, there are so many to add.
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===Autosomal DNA===
===Autosomal DNA===


Studies of Ethiopians belonging to Semitic and Cushitic ethnic groups mostly from the north of the country (the Oromo, Amhara, Tigray, and Gurage) estimate approximately 40% of their autosomal ancestry to be derived from an ancient non-African back-migration from the near East, and about 60% to be of local native African origin (from a sub-Saharan population indigenous or "autochthonous" to the Horn of Africa).<ref name="Passarino1">{{cite journal |doi=10.1086/301702 |pmc=1376879 |title=Different genetic components in the Ethiopian population, identified by mtDNA and Y-chromosome polymorphisms |pmid=9463310 |year=1998 |last1=Passarino |first1=G |last2=Semino |first2=O |last3=Quintanamurci |first3=L |last4=Excoffier |first4=L |last5=Hammer |first5=M |last6=Santachiarabenerecetti |first6=A |journal=The American Journal of Human Genetics |volume=62 |issue=2 |pages=420–34}}</ref><ref>{{cite journal|author1=Jason A. Hodgson|author2=Connie J. Mulligan|author3=Ali Al-Meeri|author4=Ryan L. Raaum|date=12 June 2014|title=Early Back-to-Africa Migration into the Horn of Africa|journal=PLOS Genetics|volume=10|issue=6|pages=e1004393|doi=10.1371/journal.pgen.1004393|pmc=4055572|pmid=24921250}}; Supplementary Text S1: Affinities of the Ethio-Somali ancestry component {{doi|10.1371/journal.pgen.1004393.s017}}{{free access}}</ref>
Studies of Ethiopians belonging to Semitic and Cushitic ethnic groups mostly from the north of the country (the Oromo, Amhara, Tigray, and Gurage) estimate approximately 40% of their autosomal ancestry to be derived from an ancient non-African back-migration from the near East, and about 60% to be of local native African origin (from a sub-Saharan population indigenous or "autochthonous" to the Horn of Africa).<ref name="Passarino1">{{cite journal |doi=10.1086/301702 |pmc=1376879 |title=Different genetic components in the Ethiopian population, identified by mtDNA and Y-chromosome polymorphisms |pmid=9463310 |year=1998 |last1=Passarino |first1=G |last2=Semino |first2=O |last3=Quintanamurci |first3=L |last4=Excoffier |first4=L |last5=Hammer |first5=M |last6=Santachiarabenerecetti |first6=A |journal=The American Journal of Human Genetics |volume=62 |issue=2 |pages=420–34}}</ref><ref>{{cite journal|author1=Jason A. Hodgson|author2=Connie J. Mulligan|author3=Ali Al-Meeri|author4=Ryan L. Raaum|date=12 June 2014|title=Early Back-to-Africa Migration into the Horn of Africa|journal=PLOS Genetics|volume=10|issue=6|pages=e1004393|doi=10.1371/journal.pgen.1004393|pmc=4055572|pmid=24921250}}; Supplementary Text S1: Affinities of the Ethio-Somali ancestry component {{doi|10.1371/journal.pgen.1004393.s017}}{{free access}}</ref> Other studies, such as Pickrel et al (2014)<ref>{{Cite journal|last=Pickrell|first=Joseph K.|last2=Patterson|first2=Nick|last3=Loh|first3=Po-Ru|last4=Lipson|first4=Mark|last5=Berger|first5=Bonnie|last6=Stoneking|first6=Mark|last7=Pakendorf|first7=Brigitte|last8=Reich|first8=David|date=2014-02-18|title=Ancient west Eurasian ancestry in southern and eastern Africa|url=https://www.pnas.org/content/111/7/2632|journal=Proceedings of the National Academy of Sciences|language=en|volume=111|issue=7|pages=2632–2637|doi=10.1073/pnas.1313787111|issn=0027-8424|pmid=24550290}}</ref>, shows the West Eurasian ancestry to peak in the Tygray at 50%. In Pagani, Luca et al (2012)<ref>{{Cite journal|last=Pagani|first=Luca|last2=Kivisild|first2=Toomas|last3=Tarekegn|first3=Ayele|last4=Ekong|first4=Rosemary|last5=Plaster|first5=Chris|last6=Gallego Romero|first6=Irene|last7=Ayub|first7=Qasim|last8=Mehdi|first8=S. Qasim|last9=Thomas|first9=Mark G.|last10=Luiselli|first10=Donata|last11=Bekele|first11=Endashaw|date=2012-07-13|title=Ethiopian genetic diversity reveals linguistic stratification and complex influences on the Ethiopian gene pool|url=https://pubmed.ncbi.nlm.nih.gov/22726845/|journal=American Journal of Human Genetics|volume=91|issue=1|pages=83–96|doi=10.1016/j.ajhg.2012.05.015|issn=1537-6605|pmc=3397267|pmid=22726845}}</ref>, this non-African component, is estimated to have entered the Horn of Africa roughly ~3,000 years ago and was found to be similar to the populations in the Levant. The paper goes on to say that his coincides with the introduction of Ethio-Semitic languages into the region.


Tishkoff et al. (2009) identified fourteen ancestral population clusters which correlate with self-described ethnicity and shared cultural and/or linguistic properties in Africa in what was the largest autosomal study of the continent to date.{{failed verification|date=May 2018}}<!--the 14 clusters are not African but worldwide, read the study--><ref name="Out of Africa: Penn Geneticist Publishes Largest-Ever Study on African Genetics Revealing Origins, Migration">[http://www.upenn.edu/pennnews/news/out-africa-penn-geneticist-publishes-largest-ever-study-african-genetics-revealing-origins-migr Out of Africa: Penn Geneticist Publishes Largest-Ever Study on African Genetics Revealing Origins, Migration].</ref> The Burji, Konso and Beta Israel were sampled from Ethiopia. The Afroasiatic speaking Ethiopians sampled were cumulatively (Fig.5B) found to belong to: 71% in the "Cushitic" cluster, 6% in the "Saharan/Dogon" cluster, 5% in the "Niger Kordofanian" cluster, 3% each in the "Nilo-Saharan" and "Chadic Saharan" cluster, while the balance (12%) of their assignment was distributed among the remnant (9) Associated Ancestral Clusters (AAC's) found in Sub-Saharan Africa.<ref>{{cite journal |last1=Tishkoff |first1=SA |last2=Reed |first2=FA |last3=Friedlaender |first3=FR |last4=Ehret |first4=C |last5=Ranciaro |first5=A |last6=Froment |first6=A |last7=Hirbo |first7=JB |last8=Awomoyi |first8=AA |last9=Bodo |first9=JM |title=The Genetic Structure and History of Africans and African Americans|journal=Science |volume=324 |issue=5930 |pages=1035–1044 |date=May 2009 |pmid=19407144 |pmc=2947357 |doi=10.1126/science.1172257 |bibcode=2009Sci...324.1035T }}</ref> The "Cushitic" cluster was also deemed "closest to the non-African AACs, consistent with an East African migration of modern humans out of Africa or a back-migration of non-Africans into Saharan and Eastern Africa."<ref name="Tishkoff2009supl">{{cite journal |last1=Tishkoff |first1=SA |last2=Reed |first2=FA |last3=Friedlaender |first3=FR |last4=Ehret |first4=C |last5=Ranciaro |first5=A |last6=Froment |first6=A |last7=Hirbo |first7=JB |last8=Awomoyi |first8=AA |last9=Bodo |first9=JM |title=The Genetic Structure and History of Africans and African Americans|journal=Science |volume=324 |issue=5930 |pages=1035–1044 |date=May 2009 |pmid=19407144 |pmc=2947357 |doi=10.1126/science.1172257 |bibcode=2009Sci...324.1035T }}; [http://www.sciencemag.org/cgi/data/1172257/DC1/1 Supplementary Material]</ref>
Tishkoff et al. (2009) identified fourteen ancestral population clusters which correlate with self-described ethnicity and shared cultural and/or linguistic properties in Africa in what was the largest autosomal study of the continent to date.{{failed verification|date=May 2018}}<!--the 14 clusters are not African but worldwide, read the study--><ref name="Out of Africa: Penn Geneticist Publishes Largest-Ever Study on African Genetics Revealing Origins, Migration">[http://www.upenn.edu/pennnews/news/out-africa-penn-geneticist-publishes-largest-ever-study-african-genetics-revealing-origins-migr Out of Africa: Penn Geneticist Publishes Largest-Ever Study on African Genetics Revealing Origins, Migration].</ref> The Burji, Konso and Beta Israel were sampled from Ethiopia. The Afroasiatic speaking Ethiopians sampled were cumulatively (Fig.5B) found to belong to: 71% in the "Cushitic" cluster, 6% in the "Saharan/Dogon" cluster, 5% in the "Niger Kordofanian" cluster, 3% each in the "Nilo-Saharan" and "Chadic Saharan" cluster, while the balance (12%) of their assignment was distributed among the remnant (9) Associated Ancestral Clusters (AAC's) found in Sub-Saharan Africa.<ref>{{cite journal |last1=Tishkoff |first1=SA |last2=Reed |first2=FA |last3=Friedlaender |first3=FR |last4=Ehret |first4=C |last5=Ranciaro |first5=A |last6=Froment |first6=A |last7=Hirbo |first7=JB |last8=Awomoyi |first8=AA |last9=Bodo |first9=JM |title=The Genetic Structure and History of Africans and African Americans|journal=Science |volume=324 |issue=5930 |pages=1035–1044 |date=May 2009 |pmid=19407144 |pmc=2947357 |doi=10.1126/science.1172257 |bibcode=2009Sci...324.1035T }}</ref> The "Cushitic" cluster was also deemed "closest to the non-African AACs, consistent with an East African migration of modern humans out of Africa or a back-migration of non-Africans into Saharan and Eastern Africa."<ref name="Tishkoff2009supl">{{cite journal |last1=Tishkoff |first1=SA |last2=Reed |first2=FA |last3=Friedlaender |first3=FR |last4=Ehret |first4=C |last5=Ranciaro |first5=A |last6=Froment |first6=A |last7=Hirbo |first7=JB |last8=Awomoyi |first8=AA |last9=Bodo |first9=JM |title=The Genetic Structure and History of Africans and African Americans|journal=Science |volume=324 |issue=5930 |pages=1035–1044 |date=May 2009 |pmid=19407144 |pmc=2947357 |doi=10.1126/science.1172257 |bibcode=2009Sci...324.1035T }}; [http://www.sciencemag.org/cgi/data/1172257/DC1/1 Supplementary Material]</ref>

Revision as of 06:46, 28 December 2021

Ethiopians
Total population
c. 114 million[citation needed]
Regions with significant populations
 Ethiopia
 United States460,000[1]
 Israel155,300[2]
 Saudi Arabia124,347 (Ethiopian-born)[3]
 UAE90,000[1]
 United Kingdom90,000[4]
 South Africa44,891 (Ethiopian-born)[3]
 Canada44,065[5]
 Kenya36,889 (Ethiopian-born)[3]
 Lebanon30,000[1]
 Italy30,000[1]
 Sweden21,686[6]
 Germany20,465[7]
 Sudan19,000[8]
 Australia19,349[9][10]
 South Sudan12,786 (Ethiopian-born)[3]
 Norway12,380[11]
 Djibouti12,323 (Ethiopian-born)[3]
 Netherlands9,451 (Ethiopian-born)[3]
 France8,675 (Ethiopian-born)[3]
 Yemen5,740 (Ethiopian-born)[3]
  Switzerland5,211 (Ethiopian-born)[3]
 Spain3,713 (Ethiopian-born)[3]
 Kuwait3,595 (Ethiopian-born)[3]
 Greece2,420 (Ethiopian-born)[3]
 Finland2,366 (Ethiopian-born)[3]
 Denmark2,136 (Ethiopian-born)[3]
 Somalia2,079 (Ethiopian-born)[3]
 Libya1,831 (Ethiopian-born)[3]
 Qatar1,667 (Ethiopian-born)[3]
 Egypt1,457 (Ethiopian-born)[3]
 Austria1,276 (Ethiopian-born)[3]
 Belgium1,143 (Ethiopian-born)[3]
 Uganda1,070 (Ethiopian-born)[3]
Languages
Amharic, Oromo, Somali, Tigrinya, Wolaytta, Gurage, Sidamo and other Languages of Ethiopia
Religion
Christian 60.9% Muslim 35.6%, Traditional 2.6%. Jewish 1%,[12]

Ethiopians are the native inhabitants of Ethiopia, as well as the global diaspora of Ethiopia. Ethiopians constitute several component ethnic groups, many of which are closely related to ethnic groups in neighboring Eritrea and other parts of the Horn of Africa.

Ethiopia's population is highly diverse with different languages and ethnic groups. Most of its people speak Afro-Asiatic languages (Semitic, Cushitic, and Omotic) languages, while others speak Nilo-Saharan languages. The Oromo, Amhara, Somali and Tigrayans make up more than three-quarters (75%) of the population, but there are more than 80 different ethnic groups within Ethiopia. Some of these have as few as 10,000 members.

History

Ethiopia's population is highly diverse, containing over 80 different ethnic groups. Most people in Ethiopia speak Afro-Asiatic languages, mainly of the Cushitic and Semitic branches. Oromo, Somali, Tigrayans and Amharas make up three-quarters of the population. The country also has Omotic ethnic minorities who speak Afro-Asiatic languages of the Omotic branch. They inhabit the southern regions of the country, particularly the Southern Nations, Nationalities and Peoples' Region. Among these are the Welayta and Gamo.[13] The ancestry of Afroasiatic-speaking Ethiopians, Somalian and Eritreans is primarily that of indigenous East Africans, but it also has a significant contribution from Eurasian populations as the result of a back migration (1500-3500 years ago). Nilo-Saharan peoples do not exhibit this genetic similarity; instead, their DNA shows evidence of more recent admixture (less than 1200 years ago) with other African peoples.[14]

The Ethiopian national identity can be traced back to the Kingdom of Aksum. While kingdoms such as Dʿmt had been established in the area previously, Aksum was the first to unify the Afroasiatic peoples of modern-day Ethiopia into one sovereign nation. The use of the term "Ethiopia" for this region specifically also dates to the Aksumite period; Ezana of Axum appropriated the term in the 4th century.[15] The Ethiopian nation was further developed by the territorial expansions of the Solomonic dynasty in the Middle Ages.[16]

Ethnicity

Amhara people
Tigrayans
This photo represents the varieties of dressing and hairstyle of the Oromo culture. the kid sitting in front of the group dressed Guji Oromo clothes. the four girls at the back from left to right, dressed Harar, Kamise, Borena and Showa styles and all are Oromo style
Oromo people
People in Harar

Major ethnic groups

List

Ethnic
group
Language
family
Census
(1994) [18]
Census
(2007) [19]
Number % Number %
Aari Omotic 2 0 0 0
Afar Afro-Asiatic 979,367 1.84 1,276,374 1.73
Agaw-Awi Afro-Asiatic 397,491 0.75 631,565 0.85
Agaw-Hamyra Afro-Asiatic 158,231 0.30 267,851 0.36
Alaba Afro-Asiatic 125,900 0.24 233,299 0.32
Amhara Afro-Asiatic 16,007,933 30.13 19,870,651 26.89
Anuak Nilotic 45,665 0.09 85,909 0.12
Arbore Afro-Asiatic 6,559 0.01 6,840 0.01
Argobba Afro-Asiatic 62,831 0.12 140,134 0.19
Bacha Nilo-Saharan 2,632 < 0.01
Basketo Omotic 51,097 0.10 78,284 0.11
Bench Omotic 173,123 0.33 353,526 0.48
Berta Nilo-Saharan 183,259 0.25
Bodi Nilo-Saharan 4,686 0.01 6,994 0.01
Brayle ???? 5,002 0.01
Burji Afro-Asiatic 46,565 0.09 71,871 0.10
Bena ???? 27,022 0.04
Beta Israel Afro-Asiatic 2,321 <0.01
Chara Omotic 6,984 0.01 13,210 0.02
Daasanach Afro-Asiatic 32,099 0.06 48,067 0.07
Dawro Omotic 331,483 0.62 543,148 0.74
Debase/ Gawwada Afro-Asiatic 33,971 0.06 68,600 0.09
Dirashe Afro-Asiatic 30,081 0.04
Dime Omotic 6,197 0.01 891 <0.01
Dizi Omotic 21,894 0.04 36,380 0.05
Donga Afro-Asiatic 35,166 0.05
Fedashe ???? 7,323, 0.01 3,448 < 0.01
Gamo Omotic 719,847 1.35 1,107,163 1.50
Gebato ???? 75 <0.01 1,502 < 0.01
Gedeo Afro-Asiatic 639,905 1.20 986,977 1.34
Gedicho ???? 5,483 0.01
Gidole Afro-Asiatic 54,354 0.10 41,100 0.06
Goffa Omotic 241,530 0.45 363,009 0.49
Gumuz Nilo-Saharan 121,487 0.23 159,418 0.22
Gurage Afro-Asiatic 2,290,274 4.31 1,867,377 2.53
Silt'e Afro-Asiatic 940,766 1.27
Hadiya Afro-Asiatic 1,227,933 2.15 1,884,373 2.44
Hamar Omotic 42,466 0.08 46,532 0.06
Harari Afro-Asiatic 21,757 0.04 31,869 0.04
Irob Afro-Asiatic 33,372 0.05
Kafficho Omotic 599,188 1.13 870,213 1.18
Kambaata Afro-Asiatic 499,825 0.94 630,236 0.85
Konta Omotic 83,607 0.11
Komo Nilo-Saharan 1,526 <0.01 7,795 0.01
Konso Afro-Asiatic 153,419 0.29 250,430 0.34
Koore Omotic 107,595 0.20 156,983 0.21
Kontoma Afro-Asiatic 0.4 48,543 0.05
Kunama Nilo-Saharan 2,007 <0.01 4,860 0.01
Karo Omotic 1,464 < 0.01
Kusumie ???? 7,470 0.01
Kwegu Nilo-Saharan 4,407 0.01
Male Omotic 46,458 0.09 98,114 0.13
Mao Omotic 16,236 0.03 43,535 0.06
Mareqo Afro-Asiatic[20] 38,096 0.07 64,381 0.09
Mashola Afro-Asiatic 10,458 0.01
Mere people ???? 14,298 0.02
Me'en Nilo-Saharan 52,815 0.10 151,489 0.20
Messengo ???? 15,341 0.03 10,964 0.01
Majangir Nilo-Saharan 21,959 0.03
Mossiye Afro-Asiatic 9,207 0.02 19,698 0.03
Murle Nilo-Saharan 1,469 < 0.01
Mursi Nilo-Saharan 3,258 0.01 7,500 0.01
Nao Omotic 4,005 0.01 9,829 0.01
Nuer Nilotic 64,534 0.12 147,672 0.20
Nyangatom Nilotic 14,201 0.03 25,252 0.03
Oromo Afro-Asiatic 21,080,318 32.15 25,489,024 34.49
Oyda Omotic 14,075 0.03 45,149 0.06
Qebena Afro-Asiatic[21] 35,072 0.07 52,712 0.07
Qechem ???? 2,740 0.01 2,585 < 0.01
Qewama ???? 141 <0.01 298 < 0.01
She Omotic 13,290 0.03 320 < 0.01
Shekecho Omotic 53,897 0.10 77,678 0.11
Sheko Omotic 23,785 0.04 37,573 0.05
Shinasha Omotic 32,698 0.06 52,637 0.07
Shita/Upo Nilo-Saharan 307 <0.01 1,602 < 0.01
Sidama Afro-Asiatic 1,842,314 3.47 2,966,474 4.01
Somali Afro-Asiatic 6,785,266 6.18 4,581,794 6.21
Surma Nilo-Saharan 19,632 0.04 27,886 0.04
Tigrinya[22] Afro-Asiatic 3,284,568 6.18 4,483,892 6.07
Tembaro ???? 86,510 0.16 98,621 0.13
Tsamai Afro-Asiatic 9,702 0.02 20,046 0.03
Welayta Omotic 1,269,216 2.39 1,707,079 2.31
Werji Afro-Asiatic 20,536 0.04 13,232 0.02
Yem Omotic 165,184 0.31 160,447 0.22
Zeyese Omotic 10,842 0.02 17,884 0.02
Zelmam Nilo-Saharan 2,704 < 0.01
Other/unknown 155,972 0.29 178,799 0.24
Somalian (Somalis of Somalia) 200,227 0.9
Sudanese 2,035 <0.01 10,333 0.01
Eritrean (Ethiopian people of Eritrean descent) 61,857 0.12 9,736 0.01
Kenyan 134 <0.01 737 <0.01
Djiboutian 367 <0.01 733 <0.01
Other foreigners 15,550 0.02
Total 53,132,276 73,750,932

Ethiopian diaspora

Languages

Until the fall of the Derg, Amharic served as the sole official language in government administration, courts, church and even in primary school instruction. After 1991, Amharic has been replaced in many areas by other official government languages such as Oromo, Somali and Tigrinya.[23] English is the most widely spoken foreign language and is taught in all secondary schools.

According to the 2007 Ethiopian census and the CIA World Fact Book, the largest first languages are: Oromo 24,929,567 speakers or 33.8% of the total population; Amharic 21,631,370 or 29.3% (federal working language); Somali 4,609,274 or 6.2%; Tigrinya 4,324,476 or 5.9%; Sidamo 4,981,471 or 4%; Wolaytta 1,627,784 or 2.2%; Gurage 1,481,783 or 2%; and Afar 1,281,278 or 1.7%.[17][24] Widely-spoken foreign languages include Arabic, English (major foreign language taught in schools), and Italian (spoken by an Italian minority).[17]

Religion

According to the CIA Factbook the religious demography of Ethiopia is as follows; Ethiopian Orthodox 43.8%, Muslim 31.3%, Protestant 22.8%, Catholic 0.7%, traditional 0.6%, and other 0.8%.[17]

Diaspora

The largest diaspora community is found in the United States. According to the U.S. Census Bureau,[25] 250,000 Ethiopian immigrants lived in the United States as of 2008. An additional 30,000 U.S.-born citizens reported Ethiopian ancestry.[26] According to Aaron Matteo Terrazas, "if the descendants of Ethiopian-born migrants (the second generation and up) are included, the estimates range upwards of 460,000 in the United States (of which approximately 350,000 are in the Washington, DC Metropolitan Area; 96,000 in Los Angeles; and 10,000 in New York)."[1]

A large Ethiopian community is also found in Israel, where Ethiopians make up almost 1.9% of the population.[citation needed] Almost the entire community are members of the Beta Israel community. There are also large number of Ethiopian emigrants in Saudi Arabia, Italy, Lebanon, United Kingdom, Canada, Sweden and Australia.[citation needed].

Genetic studies

Autosomal DNA

Studies of Ethiopians belonging to Semitic and Cushitic ethnic groups mostly from the north of the country (the Oromo, Amhara, Tigray, and Gurage) estimate approximately 40% of their autosomal ancestry to be derived from an ancient non-African back-migration from the near East, and about 60% to be of local native African origin (from a sub-Saharan population indigenous or "autochthonous" to the Horn of Africa).[27][28] Other studies, such as Pickrel et al (2014)[29], shows the West Eurasian ancestry to peak in the Tygray at 50%. In Pagani, Luca et al (2012)[30], this non-African component, is estimated to have entered the Horn of Africa roughly ~3,000 years ago and was found to be similar to the populations in the Levant. The paper goes on to say that his coincides with the introduction of Ethio-Semitic languages into the region.

Tishkoff et al. (2009) identified fourteen ancestral population clusters which correlate with self-described ethnicity and shared cultural and/or linguistic properties in Africa in what was the largest autosomal study of the continent to date.[failed verification][31] The Burji, Konso and Beta Israel were sampled from Ethiopia. The Afroasiatic speaking Ethiopians sampled were cumulatively (Fig.5B) found to belong to: 71% in the "Cushitic" cluster, 6% in the "Saharan/Dogon" cluster, 5% in the "Niger Kordofanian" cluster, 3% each in the "Nilo-Saharan" and "Chadic Saharan" cluster, while the balance (12%) of their assignment was distributed among the remnant (9) Associated Ancestral Clusters (AAC's) found in Sub-Saharan Africa.[32] The "Cushitic" cluster was also deemed "closest to the non-African AACs, consistent with an East African migration of modern humans out of Africa or a back-migration of non-Africans into Saharan and Eastern Africa."[33]

Wilson et al. (2001), an autosomal DNA study based on cluster analysis that looked at a combined sample of Amhara and Oromo examining a single enzyme variants: drug metabolizing enzyme (DME) loci, found that 62% of Ethiopeans fall into the same cluster of Ashkenazi Jews, Norwegians and Armenians based on that gene. Only 24% of Ethiopians cluster with Bantus and Afro-Caribbeans, 8% with Papua New Guineans, and 6% with Chinese.[34]

Paternal lineages

Y DNA Haplogroups of Ethiopia alt text
Y DNA Haplogroups of Ethiopia[35][36][37][38][39]

A composite look at most YDNA studies done so far[36][37][35][38][39] reveals that, out of a total of 459 males sampled from Ethiopia, approximately 58% of Y-chromosome haplotypes were found to belong to Haplogroup E, of which 71% (41% of total) were characterized by one of its further downstream sub lineage known as E1b1b, while the remainder were mostly characterized by Haplogroup E1b1(x E1b1b,E1b1a), and to a lesser extent Haplogroup E2. With respect to E1b1b, some studies have found that it exists at its highest level among the Oromo, where it represented 62.8% of the haplotypes, while it was found at 35.4% among the Amhara,[37] other studies however have found an almost equal representation of Haplogroup E1b1b at approximately 57% in both the Oromo and the Amhara.[40] The haplogroup (as its predecessor E1b1) is thought to have originated in Ethiopia or elsewhere in the Horn of Africa. About one half of E1b1b found in Ethiopia is further characterized by E1b1b1a (M78), which arose later in north-eastern Africa and then back-migrated to eastern Africa.[41]

Haplogroup J has been found at a frequency of approximately 18% in Ethiopians, with a higher prevalence among the Amhara, where it has been found to exist at levels as high as 35%, of which about 94% (17% of total) is of the type J1, while 6% (1% of total) is of J2 type.[42] On the other hand, 26% of the individuals sampled in the Arsi control portion of Moran et al. (2004) were found to belong to Haplogroup J.[38]

Another fairly prevalent lineage in Ethiopia belongs to Haplogroup A, occurring at a frequency of about 17% within Ethiopia, it is almost all characterized by its downstream sub lineage of A3b2 (M13). Restricted to Africa, and mostly found along the Rift Valley from Ethiopia to Cape Town, Haplogroup A represents the deepest branch in the Human Y- Chromosome phylogeny.[43]

Finally, Haplogroup T at approximately 4% and Haplogroup B at approximately 3%, make up the remainder of the Y-DNA Haplogroups found within Ethiopia.

Maternal lineages

mtDNA Haplogroups of Ethiopia alt text
mtDNA Haplogroups of Ethiopia[44]

The maternal ancestry of Ethiopians is similarly diverse. About half (52.2%) of Ethiopians belongs to mtdna Haplogroups L0, L1, L2, L3, L4, L5, or L6. These haplogroups are generally confined to the African continent. They also originated either in Ethiopia or very near. The other portion of the population belong to Haplogroup N (31%) and Haplogroup M1 (17%).[44] There is controversy surrounding their origins as either native or a possible ancient back migration into Ethiopia from Asia.

Passarino et al. (1998) suggested that:

Caucasoid gene flow into the Ethiopian gene pool occurred predominantly through males. Conversely, the Niger–Congo contribution to the Ethiopian population occurred mainly through females.[27]

While there is debate among the scientific community of what exactly constitutes "Caucasoid gene flow",[45][46] the same study further stated:

Indeed, Ethiopians do not seem to result only from a simple combination of proto-Niger–Congo and Middle Eastern genes. Their African component cannot be completely explained by that of present-day Niger–Congo speakers, and it is quite different from that of the Khoisan. Thus, a portion of the current Ethiopian gene pool may be the product of in situ differentiation from an ancestral gene pool."[27]

Scott et al. (2005) similarly observed that the Ethiopian population is almost equally divided between individuals that carry Eurasian maternal lineages, and those that belong to African clades. They describe the presence of Eurasian clades in the country as sequences that "are thought to be found in high numbers in Ethiopia either as a result of substantial gene flow into Ethiopia from Eurasia (Chen et al., 2000; Richards et al., 2003), or as a result of having undergone several branching events in demic diffusion, acting as founder lineages for non-African populations". The researchers further found no association between regional origin of subjects or language family (Semitic/Cushitic) and their mitochondrial type:

The haplogroup distribution amongst all subjects (athletes and controls) from different geographical regions of Ethiopia is displayed in Table 3. As can be seen graphically in Fig. 3, the mtDNA haplogroup distribution of each region is similar, with all regions displaying similar proportions of African 'L' haplogroups (Addis Ababa: 59%, Arsi: 50%, Shewa: 44%, Other: 57%). No association was found between regional origin of subjects and their mitochondrial type (v2=8.5, 15 df, P=0.9). Similarly, the mtDNA haplogroup distribution of subjects (athletes and controls) speaking languages from each family is shown in Table 3. Again there was no association between language family and mitochondrial type (v2=5.4, 5 df, P=0.37). As can be seen in Fig. 4, the haplogroup distributions of each language family are again very similar.[47]

In addition, Musilová et al. (2011) observed significant maternal ties between its Ethiopian and other Horn African samples with its Western Asian samples; particularly in terms of the HV1b mtDNA haplogroup. The authors noted:

"Detailed phylogeography of HV1 sequences shows that more recent demographic upheavals likely contributed to their spread from West Arabia to East Africa, a finding concordant with archaeological records suggesting intensive maritime trade in the Red Sea from the sixth millennium BC onwards."[48]

According to Černý et al. (2008), many Ethiopians also share specific maternal lineages with areas in Yemen and other parts of Northeast Africa. The authors indicate that:

"The most frequent haplotype in west coastal Yemen is 16126–16362, which is found not only in the Ethiopian highlands but also in Somalia, lower Egypt and at especially high frequency in the Nubians. The Tihama share some West Eurasian haplotypes with Africans, e.g. J and K with Ethiopians, Somali and Egyptians."[49]

See also

Footnotes

  1. ^ a b c d e Terrazas, Aaron Matteo (June 2007). "Beyond Regional Circularity: The Emergence of an Ethiopian Diaspora". Migration Policy Institute. Retrieved 2011-11-25.
  2. ^ Israel Central Bureau of Statistics: The Ethiopian Community in Israel
  3. ^ a b c d e f g h i j k l m n o p q r s t u "Trends in International Migrant Stock: Migrants by Destination and Origin (United Nations database, POP/DB/MIG/Stock/Rev.2015)" (XLS). United Nations, Department of Economic and Social Affairs. 2015. Retrieved 16 June 2021.
  4. ^ "Members of Ethiopian Diaspora Gather at British Home of Former Emperor". Voice of America. 14 January 2020. Retrieved 2021-12-24.
  5. ^ "Ethnic origin population". Statistics Canada. Retrieved 18 September 2019.
  6. ^ "Befolkning efter födelseland, ålder, kön och år". Statistics Sweden. Retrieved 2020-02-25.
  7. ^ https://www.destatis.de/DE/Themen/Gesellschaft-Umwelt/Bevoelkerung/Migration-Integration/Publikationen/Downloads-Migration/auslaend-bevoelkerung-2010200207004.pdf?__blob=publicationFile
  8. ^ "Members of Ethiopian Diaspora Gather at British Home of Former Emperor". Voice of America. 14 January 2020. Retrieved 2021-12-24.
  9. ^ ABS 2006a, Country of Birth
  10. ^ "Ancestry - Australia - Community profile". ABS. 2016. Retrieved 18 June 2021.
  11. ^ "Immigrants and Norwegian-born to immigrant parents, 1 January 2021". Statistics Norway (in Norwegian).
  12. ^ Berhanu Abegaz, "Ethiopia: A Model Nation of Minorities" (accessed 6 April 2006)
  13. ^ Ethiopia, 2 November 2021
  14. ^ Hellenthal, Garrett; Bird, Nancy; Morris, Sam (2021). "Structure and ancestry patterns of Ethiopians in genome-wide autosomal DNA". Human Molecular Genetics. 30 (R1): R42–R48. doi:10.1093/hmg/ddab019. PMC 8242491. PMID 33547782. Retrieved 21 October 2021.
  15. ^ Munro-Hay, Stuart (1991). Aksum: An African Civilization of Late Antiquity. Edinburgh: Edinburgh University Press. p. 15-16. ISBN 978-0748601066.
  16. ^ Henze, Paul B. (2000). Layers of Time: A History of Ethiopia. Leiden: Brill. p. 64. ISBN 978-1-137-11786-1.
  17. ^ a b c d [1]. The World Factbook. Central Intelligence Agency.
  18. ^ United nations Demographic Yearbook Special Census Topics. Volume 2b Table 4 [2]
  19. ^ 2007 Ethiopian census, first draft Archived 2012-02-14 at the Wayback Machine, Ethiopian Central Statistical Agency (accessed 6 May 2009)
  20. ^ Ulrich Braukamper 2012: 189
  21. ^ Ulrich Braukamper 2012: 165
  22. ^ [3], Ethiopian Government Portal.
  23. ^ Bulcha, Mekuria (July 1997), "The Politics of Linguistic Homogenization in Ethiopia and the Conflict over the Status of "Afaan Oromoo"", African Affairs, 96 (384): 325–352, doi:10.1093/oxfordjournals.afraf.a007852, JSTOR 723182
  24. ^ "Statistical Tables for the 2007 Population and Housing Census of Ethiopia: Country Level". Central Statistical Agency. 2007. pp. 91–92. Retrieved 2011-08-12.
  25. ^ "Ethiopia 5th largest source of Black Immigrants in America". Retrieved 16 May 2017.
  26. ^ Giorgis, Tedla W. (2011-06-29). "Potential into Practice: The Ethiopian Diaspora Volunteer Program". Migration Policy Institute. Retrieved 25 November 2011.
  27. ^ a b c Passarino, G; Semino, O; Quintanamurci, L; Excoffier, L; Hammer, M; Santachiarabenerecetti, A (1998). "Different genetic components in the Ethiopian population, identified by mtDNA and Y-chromosome polymorphisms". The American Journal of Human Genetics. 62 (2): 420–34. doi:10.1086/301702. PMC 1376879. PMID 9463310.
  28. ^ Jason A. Hodgson; Connie J. Mulligan; Ali Al-Meeri; Ryan L. Raaum (12 June 2014). "Early Back-to-Africa Migration into the Horn of Africa". PLOS Genetics. 10 (6): e1004393. doi:10.1371/journal.pgen.1004393. PMC 4055572. PMID 24921250.{{cite journal}}: CS1 maint: unflagged free DOI (link); Supplementary Text S1: Affinities of the Ethio-Somali ancestry component doi:10.1371/journal.pgen.1004393.s017Free access icon
  29. ^ Pickrell, Joseph K.; Patterson, Nick; Loh, Po-Ru; Lipson, Mark; Berger, Bonnie; Stoneking, Mark; Pakendorf, Brigitte; Reich, David (2014-02-18). "Ancient west Eurasian ancestry in southern and eastern Africa". Proceedings of the National Academy of Sciences. 111 (7): 2632–2637. doi:10.1073/pnas.1313787111. ISSN 0027-8424. PMID 24550290.
  30. ^ Pagani, Luca; Kivisild, Toomas; Tarekegn, Ayele; Ekong, Rosemary; Plaster, Chris; Gallego Romero, Irene; Ayub, Qasim; Mehdi, S. Qasim; Thomas, Mark G.; Luiselli, Donata; Bekele, Endashaw (2012-07-13). "Ethiopian genetic diversity reveals linguistic stratification and complex influences on the Ethiopian gene pool". American Journal of Human Genetics. 91 (1): 83–96. doi:10.1016/j.ajhg.2012.05.015. ISSN 1537-6605. PMC 3397267. PMID 22726845.
  31. ^ Out of Africa: Penn Geneticist Publishes Largest-Ever Study on African Genetics Revealing Origins, Migration.
  32. ^ Tishkoff, SA; Reed, FA; Friedlaender, FR; Ehret, C; Ranciaro, A; Froment, A; Hirbo, JB; Awomoyi, AA; Bodo, JM (May 2009). "The Genetic Structure and History of Africans and African Americans". Science. 324 (5930): 1035–1044. Bibcode:2009Sci...324.1035T. doi:10.1126/science.1172257. PMC 2947357. PMID 19407144.
  33. ^ Tishkoff, SA; Reed, FA; Friedlaender, FR; Ehret, C; Ranciaro, A; Froment, A; Hirbo, JB; Awomoyi, AA; Bodo, JM (May 2009). "The Genetic Structure and History of Africans and African Americans". Science. 324 (5930): 1035–1044. Bibcode:2009Sci...324.1035T. doi:10.1126/science.1172257. PMC 2947357. PMID 19407144.; Supplementary Material
  34. ^ Wilson, James F.; Weale, Michael E.; Smith, Alice C.; Gratrix, Fiona; Fletcher, Benjamin; Thomas, Mark G.; Bradman, Neil; Goldstein, David B. (2001). "Population genetic structure of variable drug response". Nature Genetics. 29 (3): 265–9. doi:10.1038/ng761. PMID 11685208. S2CID 25627134.
  35. ^ a b Underhill PA; Shen P; Lin AA; et al. (November 2000). "Y chromosome sequence variation and the history of human populations". Nature Genetics. 26 (3): 358–61. doi:10.1038/81685. PMID 11062480. S2CID 12893406.
  36. ^ a b Cruciani F; Santolamazza P; Shen P; et al. (May 2002). "A Back Migration from Asia to Sub-Saharan Africa Is Supported by High-Resolution Analysis of Human Y-Chromosome Haplotypes". American Journal of Human Genetics. 70 (5): 1197–214. doi:10.1086/340257. PMC 447595. PMID 11910562.
  37. ^ a b c Semino O, Santachiara-Benerecetti AS, Falaschi F, Cavalli-Sforza LL, Underhill PA (January 2002). "Ethiopians and Khoisan Share the Deepest Clades of the Human Y-Chromosome Phylogeny". American Journal of Human Genetics. 70 (1): 265–8. doi:10.1086/338306. PMC 384897. PMID 11719903.
  38. ^ a b c Moran et al. (2004)[4] Y chromosome haplogroups of elite Ethiopian endurance
  39. ^ a b Shenn et al. (2004)[5] Reconstruction of Patrilineages and Matrilineages of Samaritans and Other Israeli Populations From Y-Chromosome and Mitochondrial DNA Sequence Variation
  40. ^ Cruciani et al. (2004)[6] Archived 2006-05-22 at the Wayback Machine Phylogeographic Analysis of Haplogroup E3b (E-M215) Y Chromosomes Reveals Multiple Migratory Events Within and Out Of Africa
  41. ^ Cruciani F; La Fratta R; Trombetta B; et al. (June 2007). "Tracing past human male movements in northern/eastern Africa and western Eurasia: new clues from Y-chromosomal haplogroups E-M78 and J-M12". Molecular Biology and Evolution. 24 (6): 1300–11. doi:10.1093/molbev/msm049. PMID 17351267.
  42. ^ Semino O; Magri C; Benuzzi G; et al. (May 2004). "Origin, Diffusion, and Differentiation of Y-Chromosome Haplogroups E and J: Inferences on the Neolithization of Europe and Later Migratory Events in the Mediterranean Area". American Journal of Human Genetics. 74 (5): 1023–34. doi:10.1086/386295. PMC 1181965. PMID 15069642.
  43. ^ Chiaroni J, Underhill PA, Cavalli-Sforza LL (December 2009). "Y chromosome diversity, human expansion, drift and cultural evolution". Proceedings of the National Academy of Sciences of the United States of America. 106 (48): 20174–9. Bibcode:2009PNAS..10620174C. doi:10.1073/pnas.0910803106. PMC 2787129. PMID 19920170.
  44. ^ a b Kivisild T; Reidla M; Metspalu E; et al. (November 2004). "Ethiopian Mitochondrial DNA Heritage: Tracking Gene Flow Across and Around the Gate of Tears". American Journal of Human Genetics. 75 (5): 752–70. doi:10.1086/425161. PMC 1182106. PMID 15457403.
  45. ^ Hunley, Keith L.; Healy, Meghan E.; Long, Jeffrey C. (2009). "The global pattern of gene identity variation reveals a history of long-range migrations, bottlenecks, and local mate exchange: Implications for biological race". American Journal of Physical Anthropology. 139 (1): 35–46. doi:10.1002/ajpa.20932. hdl:2027.42/62159. PMID 19226641. S2CID 1757977.
  46. ^ Relethford, John H. (2009). "Race and global patterns of phenotypic variation". American Journal of Physical Anthropology. 139 (1): 16–22. doi:10.1002/ajpa.20900. PMID 19226639.
  47. ^ ScottRA (Mar 2005). "Mitochondrial DNA lineages of elite Ethiopian athletes". Comp Biochem Physiol B. 140 (3): 497–503. doi:10.1016/j.cbpc.2004.11.014. PMID 15694598.
  48. ^ Musilová; et al. (2011). "Population history of the Red Sea—genetic exchanges between the Arabian Peninsula and East Africa signaled in the mitochondrial DNA HV1 haplogroup". American Journal of Physical Anthropology. 145 (4): 592–598. doi:10.1002/ajpa.21522. PMID 21660931.
  49. ^ Černý et al. (2008), Regional differences in the distribution of the sub-Saharan, West Eurasian and South Asian mtDNA lineages in Yemen, Volume 136, Issue 2, pages 128–137, June 2008.

References

  • E. Sylvia Pankhurst, The Ethiopian People. Their Rights and Progress. Woodford Green, Essex: New Times & Ethiopia News 1946.
  • Edward Ullendorff, The Ethiopians: an introduction to country and people. London: Oxford University Press 1960, ²1965, ³1973 (ISBN 0-19-285061-X), ⁴1990 (Wiesbaden: F. Steiner; ISBN 3-515-05693-9).