East Caucasian tur
|East Caucasian tur|
|Subspecies:||C. c. cylindricornis|
|Capra caucasica cylindricornis
The East Caucasian tur or Daghestan tur (Capra caucasica cylindricornis) is a mountain-dwelling caprine found only in the eastern half of the Greater Caucasus mountains. East Caucasian turs live in rough mountainous terrain, where they eat mainly grasses and leaves and are preyed upon by wolves and lynxes. It is sometimes considered a subspecies of the West Caucasian tur, and sometimes as a full species in its own right.
East Caucasian turs are goat-like animals with large but narrow bodies and short legs, and show significant sexual dimorphism in overall size and horn development. Adult males stand about 105 cm (41 in) at the shoulder, measure 190 cm (75 in) in head-body length, and weigh around 140 kg (310 lb). The equivalent figures for adult females are 85 cm (33 in) for shoulder height, 138 cm (54 in) for head-body length, and just 56 kg (123 lb) for weight. Males have slightly lyre-shaped horns which reach 70 to 90 cm (28 to 35 in) in length, while in females they are typically only 20 to 22 cm (7.9 to 8.7 in) long.
The summer coat is short and sandy-yellow, with dirty white underparts. There are also dark brown stripes along the front surface of the legs and on the upper surface of the tail. In the winter, the coat of females and juvenile males becomes slightly greyish in colour, but otherwise remains similar. However, the winter coat of adult males is a solid dark brown, without visible stripes on the legs. Males develop a beard with their winter coat in their second year, reaching the full length of about 12 cm (4.7 in) by their fourth or fifth year. Compared with other goats, the beard of East Caucasian turs is relatively stiff, and projects somewhat forwards, rather than drooping down. The beard is small or entirely absent in females, and in males in their summer coats.
Distribution and habitat
The species range is restricted to the Greater Caucasus Mountains between 800 and 4,000 m (2,600 and 13,100 ft) above sea level, roughly extending from Mt. Shkhara (Georgia) in the west to Mt. Babadag (Azerbaijan) in the east. The western edge of the range of the East Caucasian tur remains unclear, as it overlaps with that of West Caucasian tur (Capra caucasica). Most of the species populations avoid human disturbance and occur in extremely rugged, open terrain around 3,000 m asl. In areas with no or little human disturbance, turs occur in gentler and much lower terrain. A fragment of a land with optimal terrain, climate, and degree of human disturbance for the species' occurrence is more likely to contain the species if the area of the fragment is larger and its distance to the species nearest source population is shorter.
Breeding occurs from late November to early January, with births taking place in May and June, after a gestation period of 160 to 165 days. Newborn turs weigh 3.4 to 4.1 kg (7.5 to 9.0 lb); they are usually singletons, although about 3% of pregnancies result in twin births. Young turs are extremely agile, being able to scamper about steep slopes after only a day of life. They generally start sampling grasses after one month, but continue to suckle until about December. Growth is relatively slow, with females not reaching their dull adult size for five years, and males at around ten or eleven years of age. Females reach sexual maturity at two years, but, in the wild, usually do not breed until the age of four.
Diet and behaviour
During the warm months, feeding occurs at intervals throughout the late afternoon, night, and morning, with the hottest hours of the day being spent resting in sheltered places. In winter, herds may remain in open pastures throughout the day, alternately grazing and resting. Daily movements may cover 15 to 20 km (9.3 to 12.4 mi). They eat almost all kinds of available vegetation, but prefer forbs in spring and summer, and grasses, trees, and shrubs in autumn and winter.
There is a seasonal migration covering a vertical distance of 1,500 to 2,000 m (4,900 to 6,600 ft), with an upward thrust in May and a retreat downwards in October. The adult males generally inhabit higher altitudes than females and their young, descending to join them in the breeding season. During the summer, the turs also make daily migrations, moving as much as 1,000 m (3,300 ft) vertically between feeding meadows and night-time resting spots.
During this rut, vigorous competitions arise as males vie for mating rights. Older males are dominant over younger individuals, which they drive away from females using threatening postures, rushing, and occasional clashes with their horns. Fights between equally sized males are fiercer, beginning with both animals rearing on their hind legs and butting each other, before vigorous horn-wrestling that often results in the combatants rolling down steep slopes until one submits and leaves the group. During the rut, males also mark their territory by de-barking and scent-marking tree trunks and heavy branches.
Outside of the rutting season, females live in stable groups with an average of seven individuals, often including a few juvenile males. Older males live in larger, single-sex groups, with an average of twelve members, while some younger males travel in groups of two or three. These male groups break up around November, when the rut begins and mixed-sex groups become the norm, re-forming again in January of February. In protected areas, the density of animals varies between 5 and 16/km2 (13 and 41/sq mi).
- Weinberg, P. (2008). Capra cylindricornis. In: IUCN 2008. IUCN Red List of Threatened Species. Retrieved 1 April 2009. Database entry includes a brief justification of why this species is of near threatened status.
- Weinberg, P.J. (2002). "Capra cylindricornis". Mammalian Species: Number 695: pp. 1–9. doi:10.1644/1545-1410(2002)695<0001:CC>2.0.CO;2.
- Gavashelishvili, A. (2004). "Habitat selection by East Caucasian tur (Capra cylindricornis)". Biological Conservation 120 (3): 391–398. doi:10.1016/j.biocon.2004.03.014.