2022 in arthropod paleontology

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2022 in arthropod paleontology is a list of new arthropod fossil taxa, including arachnids, crustaceans, insects, trilobites, and other arthropods that were announced or described, as well as other significant arthropod paleontological discoveries and events which occurred in 2022.

Arachnids

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acrometa gibbosa^[1]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider belonging to the family Synotaxidae.
Acrometa glomus^[1]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider belonging to the family Synotaxidae.
Acrometa longisetae^[1]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider belonging to the family Synotaxidae.
Annazomus^[2]	Gen. et sp. nov	Valid	De Francesco Magnussen & Müller in De Francesco Magnussen et al.	Cretaceous	Burmese amber	Myanmar	A member of Schizomida belonging to the family Hubbardiidae. The type species is A. parvulus De Francesco Magnussen.
Araneometa procera^[3]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Miocene	Mexican amber	Mexico	A long-jawed orb weaver.
Archaeocroton kaufmani^[4]	Sp. nov	Valid	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	Cretaceous	Burmese amber	Myanmar	A hard tick. Announced in 2022; the final article version was published in 2023.
Archaeofeaella^[5]	Gen. et sp. nov		Kolesnikov et al.	Late Triassic		Ukraine	A pseudoscorpion belonging to the family Feaellidae. Genus includes new species A. henderickxi.
Balticodromus^[1]	Gen. et sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider belonging to the family Philodromidae. The type species is B. porrectus.
Balticosynotaxus^[1]	Gen. et sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider belonging to the family Synotaxidae. The type species is B. angulatus.
Baltonychia^[6]	Gen. et sp. nov	Valid	Bartel, Derkarabetian & Dunlop	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of Opiliones belonging to the superfamily Travunioidea. The type species is B. obscura.
Betaburmesebuthus spinipedis^[7]	Sp. nov	In press	Xuan, Cai & Huang	Cretaceous	Burmese amber	Myanmar	A scorpion belonging to the family Palaeoburmesebuthidae.
Bicornoculus aungi^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Bicornoculus wunnai^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Bicornoculus yarzari^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Bothriocroton muelleri^[4]	Sp. nov	Valid	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	Cretaceous	Burmese amber	Myanmar	A hard tick. Announced in 2022; the final article version was published in 2023.
Burmalomanius^[9]	Gen. et sp. nov	In press	Bartel et al.	Cretaceous (Albian–Cenomanian)	Burmese amber	Myanmar	A member of Opiliones belonging to the family Podoctidae. Genus includes new species B. circularis.
Burmatheridion cetani^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Theridiidae.
Burmeochthonius^[10]	Gen. et 2 sp. nov		Johnson et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A pseudoscorpion belonging to the family Chthoniidae, subfamily Chthoniinae and the tribe Tyrannochthoniini. Genus includes new species B. kachinae and B. muelleri.
Caeculus fedrae^[11]	Sp. nov	Valid	Porta, Michalik & Ramírez	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A mite belonging to the family Caeculidae.
Chaerilobuthus brandti^[12]	Sp. nov	Valid	Lourenço in Lourenço & Velten	Cretaceous	Burmese amber	Myanmar	A scorpion belonging to family Chaerilobuthidae.
Chrosiothes chiapas^[3]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Miocene	Mexican amber	Mexico	A species of Chrosiothes.
Claspingblemma^[13]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae. The type species is C. duospinae.^[14]
Cretaceousbuthus^[15]	Gen. et sp. nov	Valid	Lourenço in Lourenço & Velten	Cretaceous	Burmese amber	Myanmar	A scorpion belonging to family Buthidae. The type species is C. fraaijeorum.
Cretaceoushormiops staxi^[16]	Sp. nov	Valid	Lourenço in Lourenço & Velten	Cretaceous	Burmese amber	Myanmar	A scorpion belonging to family Protoischnuridae.
Cretaceozomus^[2]	Gen. et 2 sp. nov	Valid	De Francesco Magnussen & Müller in De Francesco Magnussen et al.	Cretaceous	Burmese amber	Myanmar	A member of Schizomida. The type species is C. robustus De Francesco Magnussen; genus also includes C. angustocaudatus De Francesco Magnussen.
Cretotheridion champoi^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Theridiidae.
Cymbioblemma ohnmari^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Deinocroton copia^[17]	Sp. nov	Valid	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	Cretaceous	Burmese amber	Myanmar	A tick.
Doubravatarbus^[18]	Gen. et sp. nov	Valid	Hradská et al.	Carboniferous (Moscovian)	Kladno Formation	Czech Republic	A member of Trigonotarbida belonging to the family Aphantomartidae. The type species is D. krafti.
Electroblemma acuminataformis^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Eocryphoeca duplex^[1]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider related to Cryphoecina.
Eocryphoecara longtegap^[1]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider related to Cryphoecina.
Eogamasomorpha magnaseta^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Eotrechalea darrellubick^[1]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider, possibly a member or a relative of the family Zoropsidae.
Furcembolus inzaliae^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Pacullidae.
Groehnizomus^[2]	Gen. et 2 sp. nov	Valid	De Francesco Magnussen & Müller in De Francesco Magnussen et al.	Cretaceous	Burmese amber	Myanmar	A member of Schizomida. The type species is G. oculiferans De Francesco Magnussen & Müller; genus also includes G. rodrigoi Müller.
Hirsutisoma grimaldii^[19]	Sp. nov	Valid	Botero-Trujillo et al.	Cretaceous	Burmese amber	Myanmar	A member of Ricinulei belonging to the family Hirsutisomidae.
Ixodes antiquorum^[17]	Sp. nov	Valid	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	Cretaceous	Burmese amber	Myanmar	A tick, a species of Ixodes.
Kachintelema^[13]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Telemidae. The type species is K. calcarfemur.
Khimaira^[17]	Gen. et sp. nov	Valid	Chitimia-Dobler, Mans & Dunlop in Chitimia-Dobler et al.	Cretaceous	Burmese amber	Myanmar	A tick combining a body resembling that of a soft tick with a basis capitulum more like that of a hard tick, assigned to a new family Khimairidae as a possible transitional form between soft and hard ticks. Genus includes new species K. fossus.
Laccolithus petrunkevitchi^[1]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider belonging to the family Phrurolithidae.
Longissipalpus aliter^[13]	Nom. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the group Araneomorphae and the family Pholcochyroceridae; a replacement name for Longissipalpus cochlea Wunderlich (2021).
Mesodibunus^[9]	Gen. et sp. nov	In press	Bartel et al.	Cretaceous (Albian–Cenomanian)	Burmese amber	Myanmar	A member of Opiliones belonging to the family Epedanidae. Genus includes new species M. tourinhoae.
Microuloborus ater^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Uloboridae.
Muellerizomus^[2]	Gen. et 2 sp. nov	Valid	De Francesco Magnussen & Müller in De Francesco Magnussen et al.	Cretaceous	Burmese amber	Myanmar	A member of Schizomida. The type species is M. palicaudatus De Francesco Magnussen; genus also includes M. amandae De Francesco Magnussen & Müller.
Palaeoleptoneta acus^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Leptonetidae.
Palaeoleptoneta baculum^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Leptonetidae.
Palaeoleptoneta laticymbium^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Leptonetidae.
Palpalpaculla phyui^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Palptibiaap^[13]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and the family Zarqaraneidae. The type species is P. cochlear.
Parachimerarachne^[13]	Gen. et sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	An arachnid related to Chimerarachne. The type species is P. longiflagellum.
Petroburma^[9]	Gen. et sp. nov	In press	Bartel et al.	Cretaceous (Albian–Cenomanian)	Burmese amber	Myanmar	A member of Opiliones belonging to the family Petrobunidae. Genus includes new species P. tarsomeria.
Phycosoma icti^[20]	Sp. nov	Valid	García-Villafuerte et al.	Miocene	Mexican amber	Mexico	A species of Phycosoma.
Praeteraraneoides multidentatum^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the group Leptonetoidea and the family Protoaraneoididae.
Praetervetiator parvicirculus^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the group Araneomorphae and the family Vetiatoridae.
Priscaleclercera chimei^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae, a species of Priscaleclercera.
Priscaleclercera foshou^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae, a species of Priscaleclercera.
Priscaleclercera hlaingi^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae, a species of Priscaleclercera.
Priscaleclercera kani^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae, a species of Priscaleclercera.
Priscaleclercera kyawae^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae, a species of Priscaleclercera.
Priscaleclercera thanae^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae, a species of Priscaleclercera.
Priscaleclercera thaungi^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Psilodercidae, a species of Priscaleclercera.
Procerclypeus corniculatus^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Tetrablemmidae.
Propterkachin pygmaeus^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Uloboridae.
Propterkachin unispinatus^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Uloboridae.
Protochactas^[21]	Gen. et sp. nov	Valid	Fabio Magnani	Middle Triassic	Meride limestone	Switzerland	A basal scorpion.
Retrooecobius lwini^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Oecobiidae.
Sigillaricinuleus^[22]	Gen. et sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Ricinulei. The type species is S. tripares.
Spinicymbium curviparacymbium^[13]	Sp. nov	Valid	Wunderlich in Wunderlich & Müller	Cretaceous	Burmese amber	Myanmar	A spider belonging to the group Araneoidea and the family Zarqaraneidae.
?Sulcosynotaxus matrimonium^[1]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe (Baltic Sea region)	A spider belonging to the family Synotaxidae.
Taubaracna^[23]	Gen. et sp. nov		Martine et al.	Oligocene	Tremembé Formation	Brazil	A spider belonging to the group Araneomorphae. The type species is T. maculosa.
Thymoites carboti^[24]	Sp. nov		García-Villafuerte	Miocene	Mexican amber	Mexico	A species of Thymoites.
Triangulum^[8]	Gen. et sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Theridiidae. The type species is T. thutai.
Trichoribates roynortoni^[25]	Sp. nov		Ojeda & Rivas in Ojeda, Vega & Rivas	Miocene	La Quinta Formation (Mexican amber)	Mexico	A mite belonging to the family Ceratozetidae.
Zamilia arkari^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Oecobiidae.
Zamilia sheini^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Oecobiidae.
Zamilia shwayi^[8]	Sp. nov	Valid	Jiang & Li in Xin et al.	Late Cretaceous	Burmese amber	Myanmar	A spider belonging to the family Oecobiidae.

Arachnid research

Shanks & Selden (2022) describe the first trigonotarbid specimens from the Carboniferous Shelburn Formation of Indiana and Senora Formation of Oklahoma.^[26]
A study on the anatomy of Protoischnurus axelrodorum is published by Carvalho et al. (2022), who interpret this scorpion as a probable early member of the crown group of Iurida, possibly a stem hormurid.^[27]
Xuan, Cai & Huang (2022) describe two immature scorpions from the Cretaceous amber from Myanmar, assigned to the genus Chaerilus.^[28]
Revision of recluse spiders from the Dominican amber is published by Magalhaes et al. (2022), who transfer the species "Loxosceles" aculicaput to the family Drymusidae and to the genus Drymusa.^[29]

Crustaceans

Malacostracans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Afroapseudes^[30]	Gen. et sp. nov		Pasini, Vega & Garassino	Late Cretaceous (Cenomanian-Turonian)		Morocco	A member of Tanaidacea belonging to the group Apseudomorpha. The type species is A. cretacicus.
Anisaeger longirostrus^[31]	Sp. nov	Valid	Smith et al.	Early Triassic		United States	A member of the family Aegeridae.
Anisopagurus primigenius^[32]	Sp. nov	Valid	Ferratges et al.	Early Eocene	Serraduy Formation	Spain	A hermit crab.
Aquitainotlos^[33]	Gen. et sp. nov	Valid	Cluzaud & Ossó	Oligocene (Rupelian)		France	A crab belonging to the family Leucosiidae. The type species is A. gaasensis.
Asthenognathus fernandezi^[34]	Sp. nov		Ferratges, Zamora & Aurell	Eocene (Ypresian)	Roda Formation	Spain	A crab belonging to the family Varunidae.
Basidromilites^[35]	Gen. et sp. nov	Valid	Artal et al.	Early Eocene	Serraduy Formation	Spain	A dromioid crab. Genus includes new species B. glaessneri.
Bechleja brevirostris^[36]	Sp. nov	Valid	De Mazancourt, Wappler & Wedmann	Eocene	Messel pit	Germany	Possibly a member of the family Palaemonidae. Announced in 2022; the correction including evidence of registration in ZooBank was published in 2023.^[37]
Brunnaega labuttensis^[38]	Sp. nov	Valid	Wilson & Morel	Late Cretaceous (Cenomanian)		France	An isopod belonging to the family Cirolanidae.
Chiapasphaera^[39]	Gen. et sp. nov	In press	Vega & Bruce in Vega et al.	Early Cretaceous (Aptian)	Sierra Madre Formation	Mexico	An isopod belonging to the family Sphaeromatidae. The type species is C. cretacea.
Chronocancer^[40]	Gen. et sp. nov	Valid	Santana et al.	Early Cretaceous (Aptian-Albian)	Romualdo Formation	Brazil	A crab, probably a member of the family Orithopsidae. The type species is C. camilosantanai. Announced in 2022 in an online-only journal, and the publication did not include a ZooBank registration number; validated in 2023.^[41]
Clibanarius isabenaensis^[32]	Sp. nov	Valid	Ferratges et al.	Early Eocene	Serraduy Formation	Spain	A species of Clibanarius.
Daciapagurus szklarkaensis^[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Oxfordian)		Poland	A hermit crab.
Dardanus balaitus^[32]	Sp. nov	Valid	Ferratges et al.	Early Eocene	Serraduy Formation	Spain	A species of Dardanus.
Ebalia baldanzae^[43]	Sp. nov	Valid	Garassino, Pasini & Pizzolato	Early Pleistocene		Italy	A species of Ebalia.
Ebalia parva^[43]	Sp. nov	Valid	Garassino, Pasini & Pizzolato	Early Pleistocene		Italy	A species of Ebalia.
Eoacmaeopleura^[44]	Gen. et sp. nov	Valid	De Angeli & Bellin	Eocene		Italy	A crab belonging to the family Varunidae. Genus includes new species E. arzignanensis.
Eocalcinus veteris^[32]	Sp. nov	Valid	Ferratges et al.	Early Eocene	Serraduy Formation	Spain	A hermit crab.
Eolinurus^[45]	Gen. et comb. nov	Valid	Pasini et al.	Eocene (Ypresian)	Monte Bolca	Italy	A spiny lobster. The type species is "Palinurus" desmaresti Secrétan (1975).
Eomunidopsis texcalaensis^[46]	Sp. nov	Valid	Klompmaker et al.	Early Cretaceous (Barremian)	Zapotitlán Formation	Mexico	A member of the family Galatheidae.
Eostenetrium^[38]	Gen. et sp. nov	Valid	Wilson & Morel	Late Cretaceous (Cenomanian)		France	An isopod belonging to the family Stenetriidae. Genus includes new species E. guerangeri.
Eureotropis^[47]	Gen. et sp. nov	In press	Mendes, Santana & Carvalho	Early Cretaceous (Barremian)	Sergipe-Alagoas Basin	Brazil	A crab belonging to the family Hymenosomatidae. The type species is E. elongata.
Galatheites sforum^[46]	Sp. nov	Valid	Klompmaker et al.	Late Jurassic (Kimmeridgian)	Massenkalk Formation	Germany	A member of Galatheoidea belonging to the family Catillogalatheidae.
Galene dashtbani^[48]	Sp. nov	Valid	Khosravi et al.	Miocene (Langhian)	Mishan Formation	Iran	A species of Galene.
Huhatanka australis^[49]	Sp. nov	Valid	Schweitzer, Feldmann & Casadío	Early Cretaceous		Argentina	A member of Glypheoidea belonging to the family Mecochiridae.
Justitia confusa^[45]	Sp. nov	Valid	Pasini et al.	Eocene (Ypresian)	Monte Bolca	Italy	A spiny lobster, a species of Justitia.
Kromtitis isabenensis^[35]	Sp. nov	Valid	Artal et al.	Early Eocene	Serraduy Formation	Spain	A dromioid crab.
Ladinicaris^[50]	Gen. et sp. nov	Valid	Pasini et al.	Middle Triassic	Upper Meride Limestone	Switzerland	A member of the family Penaeidae. The type species is L. sceltrichensis.
Lantoceramia^[38]	Gen. et sp. nov	Valid	Wilson & Morel	Late Cretaceous (Cenomanian)		France	An isopod belonging to the group Cymothooidea and the new family Lantoceramiidae. Genus includes new species L. ooeida.
Lessinoachela^[45]	Gen. et sp. nov	Valid	Pasini et al.	Eocene (Ypresian)	Monte Bolca	Italy	A member of Achelata of uncertain affinities. The type species is L. scaligera.
Mainyozius^[44]	Gen. et sp. nov	Valid	De Angeli & Bellin	Eocene		Italy	A crab belonging to the family Pseudoziidae. Genus includes new species M. bituberculatus.
Mclaynotopus^[35]	Gen. et sp. nov	Valid	Artal et al.	Early Eocene	Serraduy Formation	Spain	A dromioid crab. Genus includes new species M. longispinosus.
Mesozoidotea^[38]	Gen. et sp. nov	Valid	Wilson & Morel	Late Cretaceous (Cenomanian)		France	An isopod belonging to the family Idoteidae. Genus includes new species M. gazonfierensis.
Necrocarcinus christinae^[51]	Sp. nov	In press	Van Bakel, Ossó & Jackson	Late Cretaceous (Cenomanian)	Woodbine Formation	United States ( Texas)	A crab belonging to the family Necrocarcinidae.
Paguristes perlatus^[32]	Sp. nov	Valid	Ferratges et al.	Early Eocene	Serraduy Formation	Spain	A species of Paguristes.
Palicus saenaensis^[52]	Sp. nov	Valid	Pasini, Garassino & Pizzolato	Early Pleistocene		Italy	A species of Palicus.
Paraglyphea simpsoni^[53]	Sp. nov	Valid	Charbonnier & Garassino	Middle Jurassic (Callovian)		United Kingdom	A member of Glypheidea
Parapetrochirus serratus^[32]	Sp. nov	Valid	Ferratges et al.	Early Eocene	Serraduy Formation	Spain	A hermit crab.
Paredonius^[54]	Gen. et comb. nov	Valid	Vega & Garassino	Late Cretaceous (Maastrichtian)	Potrerillos Formation	Mexico	A crab of uncertain affinities. Genus includes "Sodakus" mexicanus Vega, Feldmann & Villobos-Hiriart (1995).
Pemphix krumenackeri^[31]	Sp. nov	Valid	Smith et al.	Early Triassic		United States	A member of Glypheidea.
Protochaetilia^[38]	Gen. et sp. nov	Valid	Wilson & Morel	Late Cretaceous (Cenomanian)		France	An isopod belonging to the family Chaetiliidae. Genus includes new species P. delaunayi.
Protohymenosoma^[47]	Gen. et 2 sp. nov	In press	Mendes, Santana & Carvalho	Early Cretaceous (Barremian)	Sergipe-Alagoas Basin	Brazil	A crab belonging to the family Hymenosomatidae. Genus includes new species P. gondwanicum and P. hexagonale.
Protomunida bennickei^[46]	Sp. nov	Valid	Klompmaker et al.	Paleocene (Danian)		Denmark	A member of the family Munididae.
Protomunida eurekantha^[46]	Sp. nov	Valid	Klompmaker et al.	Paleocene (Danian)	Faxe Formation	Denmark	A member of the family Munididae.
Pylochelitergites cicatrix^[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Oxfordian)		Poland	A hermit crab.
Sienalia^[43]	Gen. et 2 sp. nov	Valid	Garassino, Pasini & Pizzolato	Early Pleistocene		Italy	A crab belonging to the family Leucosiidae. The type species is S. fredianii; genus also includes S. ristorii.
Sierradromia^[35]	Gen. et sp. nov	Valid	Artal et al.	Early Eocene	Serraduy Formation	Spain	A dromioid crab. Genus includes new species S. gladiator.
Stimdromia conternoi^[55]	Sp. nov	Valid	Beschin & De Angeli	Eocene (Bartonian)		Italy	A crab belonging to the family Dromiidae.
Teruzzicheles^[56]	Gen. et comb. nov	Valid	Audo & Charbonnier	Early Jurassic (Sinemurian)		Italy	A member of the family Polychelidae; a new genus for "Coleia" popeyei Teruzzi (1990).
Tethysgalathea^[46]	Gen. et comb. nov	Valid	Klompmaker et al.	Eocene (Ypresian)		Italy	A member of the family Galatheidae. The type species is "Eomunidopsis" prealpina Beschin et al. (2016).
Tomaricaris^[57]	Gen. et sp. nov	Valid	Garassino, Pasini & Nazarkin	Miocene (Serravallian-Tortonian)	Kurasi Formation	Russia ( Sakhalin Oblast)	A caridean shrimp belonging to the family Barbouriidae. The type species is T. ainuensis.
Torodromia^[35]	Gen. et sp. nov	Valid	Artal et al.	Early Eocene	Serraduy Formation	Spain	A dromioid crab. Genus includes new species T. elongata.
Vasconilia zapotitlanensis^[46]	Sp. nov	Valid	Klompmaker et al.	Early Cretaceous (Barremian)	Zapotitlán Formation	Mexico	A member of Galatheoidea belonging to the family Catillogalatheidae.
Viaia brinae^[58]	Sp. nov	In press	Gašparič et al.	Early Cretaceous (Aptian to Albian)		Slovenia	A crab belonging to the family Viaiidae.
Viaia tisae^[58]	Sp. nov	In press	Gašparič et al.	Early Cretaceous (Aptian to Albian)		Slovenia	A crab belonging to the family Viaiidae.
Zapalianassa^[49]	Gen. et comb. nov	Valid	Schweitzer, Feldmann & Casadío			Argentina	A new genus for "Pehuenchia" magna Rusconi (1948).

Malacostracan research

Description of the fossil material of members of Pygocephalomorpha from the Carboniferous Piesberg quarry (Lower Saxony, Germany), including a record of Anthracaris gracilis which was previously reported from the Mazon Creek fossil beds (Illinois, United States) and from Bickershaw (Lancashire, United Kingdom), and a study on the distribution of eumalacostracan crustaceans from Carboniferous deposits of North America and Europe is published by Pazinato et al. (2022).^[59]
Redescription of Oncopareia bredai and a revision of other species previously referred to the genus Oncopareia is published by Tshudy et al. (2022).^[60]
A study on the evolution of carapace morphology of hermit crabs and changes of composition of their assemblages through time is published by Fraaije et al. (2022), who reinstate Probeebeidae as a distinct family, and name a new family Paguropsidae.^[61]
A study on sexual dimorphism and intersex specimens in the population of hundreds of specimens of the Cretaceous crab Dakoticancer overanus Jones, Schweitzer & Feldmann (2022).^[62]
Ossó et al. (2022) report the first known fossil material of Pleolobites from the Paleocene (Thanetian) of Togo, and reevaluate the phylogenetic affinities of this and other fossil portunoid crabs.^[63]
A concretion containing an adult and a juvenile individual of Trichopeltarion greggi, representing the first association of an adult female and a juvenile crab of the same species in the fossil record reported to date, is described from the Miocene Greta Siltstone (New Zealand) by Feldmann & Schweitzer (2022), who interpret this finding as possible evidence of maternal care of juveniles.^[64]

Ostracods

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Aaleniella franzi^[65]	Sp. nov	Valid	Tesakova	Middle Jurassic		Russia
Aaleniella? ovoidea^[65]	Sp. nov	Valid	Tesakova	Middle Jurassic		Russia
Aaleniella volganica^[65]	Sp. nov	Valid	Tesakova	Middle Jurassic		Russia
Acrocythere sokurensis^[66]	Sp. nov	Valid	Tesakova	Middle Jurassic		Russia
Alatobairdia? sohni^[67]	Sp. nov	Valid	Forel in Forel et al.	Late Triassic (Norian)	Aksala Formation	Canada ( Yukon)	A member of the family Bairdiidae.
Anticythereis dorsennus^[68]	Sp. nov	Valid	Puckett & Hunt	Late Cretaceous		United States
Anticythereis slipperi^[68]	Sp. nov	Valid	Puckett & Hunt	Late Cretaceous		United States
Asculdoracythereis^[68]	Gen. et 3 sp. nov	Valid	Puckett & Hunt	Late Cretaceous		United States	Genus includes new species A. asculdora, A. invicta and A. pseudoalabamensis.
Aurila diodoroi^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Aurila seguenzai^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Bairdia aksala^[67]	Sp. nov	Valid	Forel in Forel et al.	Late Triassic (Norian)	Aksala Formation	Canada ( Yukon)	A member of the family Bairdiidae.
Bairdia taan^[67]	Sp. nov	Valid	Forel in Forel et al.	Late Triassic (Carnian and Norian)	Aksala Formation	Canada ( Yukon) Italy	A member of the family Bairdiidae.
Bairdia yukonensis^[67]	Sp. nov	Valid	Forel in Forel et al.	Late Triassic (Norian)	Aksala Formation	Canada ( Yukon)	A member of the family Bairdiidae.
Bairdiacypris xainzaensis^[70]	Sp. nov	In press	Song et al.	Late Devonian	Chaguoluoma Formation	China
Bairdoppilata barremiana^[71]	Sp. nov		Mojon in Mojon & De Kaenel	Early Cretaceous (Barremian)		France Switzerland
Bradyleberis praecristatella^[72]	Sp. nov		McDonald & Warne	Miocene	Bookpurnong Formation	Australia
Callistocythere bookpurnongensis^[72]	Sp. nov		McDonald & Warne	Miocene	Bookpurnong Formation	Australia	A species of Callistocythere.
Callistocythere mchenryi^[72]	Sp. nov		McDonald & Warne	Miocene	Bookpurnong Formation	Australia	A species of Callistocythere.
Callistocythere zigzaga^[72]	Sp. nov		McDonald & Warne	Miocene	Bookpurnong Formation	Australia	A species of Callistocythere.
Camptocythere (Anabarocythere) muricata^[73]	Sp. nov	Valid	Gerke & Lev in Tesakova	Middle Jurassic (Bathonian–Callovian)		Russia
Camptocythere (Anabarocythere) triangula^[73]	Sp. nov	Valid	Tesakova	Middle Jurassic (Bajocian and Bathonian)		Russia ( Saratov Oblast)
Camptocythere (Camptocythere) quinta^[74]	Sp. nov	Valid	Tesakova in Tesakova & Seltser	Jurassic		Russia
Camptocythere (Palaeoloxoconcha) caudata^[74]	Sp. nov	Valid	Tesakova in Tesakova & Seltser	Jurassic		Russia
Camptocythere (Palaeoloxoconcha) ryazanica^[74]	Sp. nov	Valid	Tesakova in Tesakova & Seltser	Jurassic		Russia
Ceratopsis persicus^[75]	Sp. nov	Valid	Salas in Poursalehi et al.	Ordovician (Katian–Hirnantian)	Katkoyeh Formation	Iran	A member of Podocopa belonging to the order Beyrichiocopida and the family Tetradellidae.
Cimbaurila ramdohri^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Cistacythereis oertlii^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Clavofabella? lanshella^[76]	Sp. nov	Valid	Guillam & Forel in Guillam et al.	Devonian (Famennian)	Gelaohe Formation	China	A member of Palaeocopida belonging to the family Primitiopsidae.
Cornutobairdia yukonella^[67]	Sp. nov	Valid	Forel in Forel et al.	Late Triassic (Norian)	Aksala Formation	Canada ( Yukon)	A member of the family Bairdiidae.
Costa agyrina^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Cyprideis atalaiensis^[77]	Sp. nov	Valid	Linhares & Ramos	Early Miocene	Solimões Formation	Brazil	A member of the family Cytherideidae.
Cyprideis dictyon^[77]	Sp. nov	Valid	Linhares & Ramos	Middle to late Miocene	Solimões Formation	Brazil	A member of the family Cytherideidae.
Cyprideis indianaensis^[78]	Sp. nov		Sousa & Ramos	Miocene	Pebas Formation	Peru
Cyprideis santaelenae^[78]	Sp. nov		Sousa & Ramos	Miocene	Pebas Formation	Peru
Cyprideis soledadensis^[78]	Sp. nov		Sousa & Ramos	Miocene	Pebas Formation	Peru
Cytherellina caerulea^[76]	Sp. nov	Valid	Guillam & Forel in Guillam et al.	Devonian (Famennian)	Gelaohe Formation	China	A member of Podocopida belonging to the family Bairdiocyprididae.
Damonella pumila^[79]	Sp. nov		Guzmán et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil
Deefgella? pulchra^[80]	Sp. nov	Valid	Melnikova et al.	Ordovician (Sandbian)	Gryazno Formation	Russia ( Leningrad Oblast)
Echinocythereis hartmannii^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Electrocypria^[81]	Gen. et sp. nov	Valid	Wang, Matzke-Karasz & Horne	Cretaceous (Albian–Cenomanian)	Burmese amber	Myanmar	A member of the family Candonidae belonging to the subfamily Paracypridinae. The type species is E. burmitei.
Eucytherura sanctavenerae^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Fortistriginglymus^[72]	Gen. et comb. nov		McDonald & Warne	Miocene	Bookpurnong Formation	Australia	Genus includes "Cythere" postdeclivis Chapman (1914).
Frodocythereis^[68]	Gen. et sp. nov	Valid	Puckett & Hunt	Late Cretaceous		United States	Genus includes new species F. frodoi.
Glyptocythere bathonica^[82]	Sp. nov	Valid	Tesakova	Middle Jurassic (Bathonian)		Kazakhstan Russia
Hungarella limella^[67]	Sp. nov	Valid	Forel in Forel et al.	Late Triassic (Norian)	Aksala Formation	Canada ( Yukon)	A member of Podocopa belonging to the family Healdiidae.
Ilyocypris coimbrai^[79]	Sp. nov		Guzmán et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil
Karlingrella^[83]	Gen. et comb. nov	Valid	Groos-Uffenorde in Groos-Uffenorde et al.	Early Devonian		Germany Morocco Spain	A member of Podocopida, possibly a member of the family Ropolonellidae. The type species is "Euglyphella" granulosa Blumenstengel (1962).
Kirkbya panamintensis^[84]	Sp. nov	Valid	McMenamin	Carboniferous (Mississippian)	Tin Mountain Limestone	United States ( California)	A member of Palaeocopida belonging to the group Beyrichicopina and the family Kirkbyidae.
Klyasinella^[80]	Gen. nov	Valid	Melnikova et al.	Ordovician (Sandbian)	Gryazno Formation	Russia ( Leningrad Oblast)	The type species is K. bella.
Kozuria^[85]	Gen. et comb. nov	Junior homonym	Forel	Middle and Late Triassic		Hungary Romania Turkey	A new genus erected for Triassic ostracods formerly assigned to the genus Acanthoscapha, such as "Acanthoscapha" veghae Kozur (1970), "Acanthoscapha" mersinella Forel in Forel et al. (2017) and "Acanthoscapha" amphialata Kristan-Tollmann (1973). The generic name is preoccupied by the radiolarian genus Kozuria Zhang (1990).
Laevipellacythereis^[68]	Gen. et 2 sp. nov	Valid	Puckett & Hunt	Late Cretaceous		United States	Genus includes new species L. colossus and L. laevipellis.
Leviella alexi^[86]	Sp. nov		Forel, Kolar-Jurkovšek & Jurkovšek	Middle Triassic (Ladinian)		Slovenia
Leviella riedeli^[67]	Sp. nov	Valid	Forel in Forel et al.	Late Triassic (Norian)	Aksala Formation	Canada ( Yukon)	A member of the family Cytherellidae.
Lobobairdia whitella^[67]	Sp. nov	Valid	Forel in Forel et al.	Late Triassic (Norian)	Aksala Formation	Canada ( Yukon)	A member of the family Bairdiidae.
Longiscula? destorta^[80]	Sp. nov	Valid	Melnikova et al.	Ordovician (Sandbian)	Gryazno Formation	Russia ( Leningrad Oblast)
Loxoconcha pokornyi^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Mantelliana speculum^[87]	Sp. nov	Valid	Tomé et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil
Microceratina retangularis^[87]	Sp. nov	Valid	Tomé et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil
Mirabairdia canadia^[67]	Sp. nov	Valid	Forel in Forel et al.	Late Triassic (Norian)	Aksala Formation	Canada ( Yukon)	A member of the family Bairdiidae.
Mirabairdia slovenica^[86]	Sp. nov		Forel, Kolar-Jurkovšek & Jurkovšek	Middle Triassic (Ladinian)		Slovenia
Mongolianella aptianensis^[87]	Sp. nov	Valid	Tomé et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil
Nanacythere octum^[66]	Sp. nov	Valid	Tesakova	Middle Jurassic		Russia
Nikitinella songliangensis^[88]	Sp. nov	In press	Zhang	Ordovician	Huadan Formation	China
Paracandona rosaepraeceps^[89]	Sp. nov	Valid	Antonietto, Eaton & Park Boush	Paleogene	Claron Formation	United States ( Utah)
Paracypris undulareventralis^[87]	Sp. nov	Valid	Tomé et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil
Paracytheridea longicristata^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Parakeijia notoreticularis^[72]	Sp. nov		McDonald & Warne	Miocene	Bookpurnong Formation	Australia
Paranacythere^[90]	Gen. et sp. nov	Valid	Bergue et al.	Permian (Guadalupian)	Teresina Formation	Brazil	A possible member of the family Cytheruridae. The type species is P. nigripallus.
Pattersoncypris cucurves^[79]	Sp. nov		Guzmán et al.	Early Cretaceous (Aptian)	Barbalha Formation	Brazil
Pattersoncypris kroemmelbeini^[79]	Sp. nov		Guzmán et al.	Early Cretaceous (Aptian)		Brazil
Perissocytheridea florisdorsalis^[87]	Sp. nov	Valid	Tomé et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil
Perissocytheridea oculusutilis^[87]	Sp. nov	Valid	Tomé et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil
Perissocytheridea poruslinearis^[87]	Sp. nov	Valid	Tomé et al.	Early Cretaceous (Aptian)	Romualdo Formation	Brazil
Procytherura ippolitovi^[74]	Sp. nov	Valid	Tesakova in Tesakova & Seltser	Jurassic		Russia
Procytherura iyae^[66]	Sp. nov	Valid	Tesakova	Middle Jurassic		Russia
Quasillites (Beckjennites) gebeckeri^[83]	Sp. nov	Valid	Groos-Uffenorde in Groos-Uffenorde et al.	Early Devonian	Lower Yeraifa Formation	Morocco	A member of Podocopida belonging to the family Quasillitidae.
Rhinocypris spinata^[79]	Sp. nov		Guzmán et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil
Sansabella gelaohensis^[76]	Sp. nov	Valid	Guillam & Forel in Guillam et al.	Devonian (Famennian)	Gelaohe Formation	China	A member of Palaeocopida belonging to the family Sansabellidae.
Satiellina zarandensis^[75]	Sp. nov	Valid	Salas in Poursalehi et al.	Ordovician (Katian–Hirnantian)	Katkoyeh Formation	Iran	A member of Podocopa belonging to the order Beyrichiocopida and the superfamily Drepanelloidea.
Sulcella baisuzhena^[76]	Sp. nov	Valid	Guillam & Forel in Guillam et al.	Devonian (Famennian)	Gelaohe Formation	China	A member of Platycopida belonging to the family Cavellinidae.
Tegmenia mariasophiae^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Tenedocythere eleonorae^[69]	Sp. nov		Sciuto, Baldanza & Reitano	Early Pleistocene		Italy
Trachycythere peculiaris^[66]	Sp. nov	Valid	Tesakova	Middle Jurassic		Russia
Tumulocythereis^[68]	Gen. et 3 sp. nov	Valid	Puckett & Hunt	Late Cretaceous		United States	Genus includes new species T. incompta, T. tiberti and T. tumulus.
Velatomorpha pseudoaltilis^[90]	Sp. nov	Valid	Bergue et al.	Permian (Guadalupian)	Teresina Formation	Brazil	A member of Platycopida belonging to the family Carbonitidae.
Velatomorpha xavante^[90]	Sp. nov	Valid	Bergue et al.	Permian (Guadalupian)	Teresina Formation	Brazil	A member of Platycopida belonging to the family Carbonitidae.
Vendona spinifera^[88]	Sp. nov	In press	Zhang	Ordovician	Huadan Formation	China
Zonocypris berthoui^[91]	Sp. nov	Valid	Piovesan et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil
Zonocypris dorsoconvexa^[91]	Sp. nov	Valid	Piovesan et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil

Other crustaceans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Arcoscalpellum s.l. hamurorum^[92]	Sp. nov	Valid	Karasawa & Amano	Miocene	Higashibessho Formation	Japan	A barnacle belonging to the family Scalpellidae.
Cetopirus polysyrinx^[93]	Sp. nov		Collareta et al.	Pleistocene		United States ( California Oregon)	A whale barnacle.
Dabashanella longa^[94]	Sp. nov	In press	Zhang	Cambrian Stage 3	Shuijingtuo Formation	China	A member of Phosphatocopida.
Dabashanella semiorbiculata^[94]	Sp. nov	In press	Zhang	Cambrian Stage 3	Shuijingtuo Formation	China	A member of Phosphatocopida.
Dabashanella unispinata^[94]	Sp. nov	In press	Zhang	Cambrian Stage 3	Shuijingtuo Formation	China	A member of Phosphatocopida.
Dietericambria^[95]	Gen. et sp. nov	Valid	Peel	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A stem-group pentastomid. The type species is D. hensoniensis.
Heidiops^[96]	Gen. et comb. nov	Valid	Werneburg & Schneider	Permian (Kungurian)	Salagou Formation	France	A notostracan of uncertain affinities. The type species is "Triops cancriformis" permiensis Gand, Garric & Lapeyrie (1997), raised to the rank of the species Heidiops permiensis.
Koonwarrella^[97]	Gen. et sp. nov	In press	Van Houte, Hegna & Butler	Early Cretaceous (Aptian)	Koonwarra Fossil Bed	Australia	An anostracan, possibly a member of the family Thamnocephalidae. Genus includes new species K. peterorum.
Linglongtaestheria qinglongensis^[98]	Sp. nov		Teng, Li & Zhang	Late Jurassic	Tiaojishan Formation	China	A clam shrimp.
Magnitocyclus^[99]	Gen. et sp. nov	Valid	Mychko et al.	Carboniferous (Viséan–Serpukhovian)		Russia	A member of Cyclida. Genus includes new species M. struveae.
Pachyscalpellum saskatchewanensis^[100]	Comb. nov	valid	(Russell)	Upper Cretaceous (Maastrichtian)	Bearpaw Formation	Canada ( Saskatchewan)	A giant barnacle, formerly known as "Calantica" saskatchewanensis Russell (1967).
Protochelonibia starnesi^[101]	Sp. nov	Valid	Perreault, Collareta & Buckeridge	Oligocene (Rupelian)	Chickasawhay Formation	United States ( Mississippi)	A barnacle belonging to the family Chelonibiidae.
Rhabdostichus arcticensis^[102]	Sp. nov	Valid	Shen & Wu	Devonian	Melville Island Formation	Canada	A member of the clam shrimp or ostracod family Rhabdostichidae.
Thuringiops^[96]	Gen. et sp. nov	Valid	Werneburg & Schneider	Carboniferous (Gzhelian) to Permian (Sakmarian)	Upper Oberhof Formation	Germany	A calmanostracan branchiopod, possibly a member of Kazacharthra. The type species is T. nephroides.
Yanjiestheria huajiyingensis^[103]	Sp. nov		Li	Early Cretaceous (Valanginian–Hauterivian)	Huajiying Formation	China	A clam shrimp.

Other crustacean research

Collareta et al. (2022) describe borings on a sea turtle carapace from the Miocene (Tortonian) Pisco Formation (Peru), interpreted as probable attachment scars produced by turtle barnacles, and argue that sea turtles may have hosted barnacle symbionts as early as during the early Oligocene.^[104]

Insects

Radiodonts

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Innovatiocaris maotianshanensis^[105]	Gen. et sp. nov	Valid	Zeng et al.	Cambrian Stage 3	Maotianshan Shales	China	A radiodont of uncertain family. Formerly described as Anomalocaris sp. and whole body specimen of "Anomalocaris" saron.
"Innovatiocaris" multispiniformis^[105]	Sp. nov	Valid	Zeng et al.	Cambrian Stage 3	Maotianshan Shales	China	A radiodont of uncertain family.

Radiodont research

New information on the anatomy of Stanleycaris hirpex, based on data from 268 specimens from the Cambrian Burgess Shale (British Columbia, Canada), is presented by Moysiuk & Caron (2022), who report exquisite preservation of the brain of this radiodont and unexpected presence of a large median eye.^[106]

Trilobites

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Agerina boygeorgei^[107]	Sp. nov	Valid	Karim & Adrain	Ordovician (Floian)		United States ( Nevada)	A member of the family Phillipsinellidae.
Aguaditaspis^[108]	Gen. et sp. nov	Valid	Randolfe, Rustán & Bignon	Devonian (Lochkovian-Pragian)	Talacasto Formation	Argentina	A member of the family Dalmanitidae. The type species is A. mediaspina.
Aitkenaspis^[109]	Gen. et sp. nov	Valid	Handkamer & Pratt in Handkamer, Pratt & MacNaughton	Cambrian		Canada	Genus includes new species A. keelensis.
Albertelloides eliasi^[109]	Sp. nov	Valid	Handkamer & Pratt in Handkamer, Pratt & MacNaughton	Cambrian		Canada
Amphitryon constrictus^[110]	Sp. nov	In press	Wei & Zhou in Wei et al.	Ordovician (Katian)	Daduhe Formation	China
Anebocephalus^[111]	Gen. et sp. nov	Valid	Sundberg & Webster	Cambrian Stage 4	Harkless Formation	United States ( Nevada)	Genus includes new species A. silverpeakensis.
Arndellaspis^[112]	Gen. et sp. nov	Valid	Holloway, Banks & Banks	Silurian (Rhuddanian)	Arndell Sandstone	Australia	A member of the family Encrinuridae. The type species is A. oryxis.
Auritolithus^[112]	Gen. et sp. nov	Valid	Holloway, Banks & Banks	Ordovician (Katian)	Arndell Sandstone	Australia	A member of the family Trinucleidae. The type species is A. corbetti.
Bignonops^[113]	Gen. et comb. nov	Valid	Van Viersen & Kloc	Devonian	Timnrhanrhart Formation	Morocco	Monotypic genus in Acastidae. The type species is B. tamnrhertus.
Bolbolenellus dodoensis^[109]	Sp. nov	Valid	Handkamer & Pratt in Handkamer, Pratt & MacNaughton	Cambrian		Canada
Brongniartella calveri^[112]	Sp. nov	Valid	Holloway, Banks & Banks	Silurian (Rhuddanian)	Arndell Sandstone	Australia	A member of the family Homalonotidae.
Cambrosaurura^[114]	Gen. et comb. et 2 sp. nov	Valid	Geyer in Cederström et al.	Cambrian	Jbel Wawrmast Formation	Morocco	A member of the family Ellipsocephalidae belonging to the subfamily Ellipsocephalinae. The type species is "Ornamentaspis" usitata Geyer (1990); genus also includes new species C. bommeli and C. robusta, and possibly also Ornamentaspis? todraensis Geyer (1990).
Catillicephala cifellii^[115]	Sp. nov	Valid	Westrop & Dengler	Cambrian (Guzhangian)	Shallow Bay Formation	Canada ( Newfoundland and Labrador)	A member of the family Catillicephalidae.
Chelediscus garzoni^[116]	Sp. nov	Valid	Collantes et al.	Cambrian Stage 4		Spain
Clarksonops^[117]	Gen. et sp. nov	Disputed	Crônier & Waters	Devonian (Famennian)	Hongguleleng Formation	China	A phacopid trilobite. Genus includes new species C. junggariensis. The genus Clarksonops was subsequently considered to be a junior synonym of the genus Omegops by Zong (2023), resulting in a new combination Omegops junggariensis.^[118]^[119]
Coenoides^[111]	Gen. et sp. nov	Valid	Sundberg & Webster	Cambrian Stage 4	Harkless Formation	United States ( Nevada)	Genus includes new species C. scholteni.
Cuneoaxiella^[114]	Gen. et comb. nov	Valid	Cederström et al.	Cambrian	Gislöv Formation	Morocco Sweden	A member of the family Ellipsocephalidae belonging to the subfamily Protoleninae. The type species is "Proampyx" grandis Ahlberg & Bergström (1978); genus also includes "Cambrunicornia" agdziensis Geyer (1990).
Dellingia^[114]	Gen. et comb. nov	Valid	Cederström et al.	Cambrian	Gislöv Formation	Sweden	A member of the family Ellipsocephalidae belonging to the subfamily Ellipsocephalinae. The type species is "Comluella" scanica Ahlberg & Bergström (1978).
Dodoella^[109]	Gen. et sp. nov	Valid	Handkamer & Pratt in Handkamer, Pratt & MacNaughton	Cambrian		Canada	Genus includes new species D. kobayashii.
Ellipsostrenua brevifrons^[114]	Sp. nov	Valid	Cederström et al.	Cambrian	Gislöv Formation	Sweden	A member of the family Ellipsocephalidae belonging to the subfamily Ellipsocephalinae.
Ellipsostrenua simrica^[114]	Sp. nov	Valid	Cederström et al.	Cambrian	Gislöv Formation	Sweden	A member of the family Ellipsocephalidae belonging to the subfamily Ellipsocephalinae.
Ellipsostrenua troedssoni^[114]	Sp. nov	Valid	Cederström et al.	Cambrian	Gislöv Formation	Sweden	A member of the family Ellipsocephalidae belonging to the subfamily Ellipsocephalinae.
Eobathyuriscus^[109]	Gen. et 2 sp. nov	Valid	Handkamer & Pratt in Handkamer, Pratt & MacNaughton	Cambrian		Canada	Genus includes new species E. mackenziensis and E. macqueeni.
Eosaukia anhuiensis^[120]	Sp. nov	Valid	Lei & Peng	Cambrian (Furongian)	Chaumitien Formation	China	A saukiid trilobite.
Epichalnipsus bergstroemi^[114]	Sp. nov	Valid	Cederström et al.	Cambrian	Gislöv Formation	Sweden	A member of the family Ellipsocephalidae belonging to the subfamily Strenuaevinae.
Fantasticolithus^[121]	Gen. et sp. nov		Fortey & Gutiérrez-Marco	Early Ordovician		Peru	A trinucleid trilobite. Genus includes new species F. isabelae.
Forteyaspis^[107]	Gen. et 2 sp. et comb. nov	Valid	Karim & Adrain	Ordovician		Canada United Kingdom	A member of the family Phillipsinellidae. The type species is F. idoli; genus also includes new species F. adamanti, as well as "Agerina" laurentica Ingham & Tripp (1991) and "Agerina" norrisi Ludvigsen (1980).
Geigibole kohleichensis^[122]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Viséan)		Germany
Glossopleura youngi^[109]	Sp. nov	Valid	Handkamer & Pratt in Handkamer, Pratt & MacNaughton	Cambrian		Canada
Gondwanaspis eisbornensis^[123]	Sp. nov	Valid	Helling & Becker	Devonian (probably early Frasnian)	Hagen-Balve Formation	Germany	A member of the family Odontopleuridae.
Gondwanaspis schloesseri^[123]	Sp. nov	Valid	Helling & Becker	Devonian (probably Givetian)	Hofermühle Formation	Germany	A member of the family Odontopleuridae.
Gravicalymene clarkei^[112]	Sp. nov	Valid	Holloway, Banks & Banks	Silurian (Rhuddanian)	Arndell Sandstone	Australia	A member of the family Calymenidae belonging to the subfamily Calymeninae.
Harklessaspis^[111]	Gen. et 2 sp. nov	Valid	Sundberg & Webster	Cambrian Stage 4	Harkless Formation	United States ( Nevada)	Genus includes new species H. rasettii and H. parvigranulosus.
Hollardops angustifrons^[113]	Sp. nov	Valid	Van Viersen & Kloc	Devonian	Timrhanrhart Formation	Morocco	A member of the family Acastidae.
Hollardops klugi^[113]	Sp. nov	Valid	Van Viersen & Kloc	Devonian	Seheb El Rhassel Group	Morocco	A member of the family Acastidae.
Hollardops kyriarchos^[113]	Sp. nov	Valid	Van Viersen & Kloc	Devonian	Khebchia Formation	Morocco	A member of the family Acastidae.
Hollardops luscus^[113]	Sp. nov	Valid	Van Viersen & Kloc	Devonian	El Otfal Formation	Morocco	A member of the family Acastidae.
Hollardops multatuli^[113]	Sp. nov	Valid	Van Viersen & Kloc	Devonian	Khebchia Formation	Morocco	A member of the family Acastidae.
Ivanites^[108]	Gen. et sp. nov	Valid	Randolfe, Rustán & Bignon	Devonian (Lochkovian-Pragian)	Talacasto Formation	Argentina	A member of the family Dalmanitidae. The type species is I. leonorae.
Japonoscutellum senectum^[112]	Sp. nov	Valid	Holloway, Banks & Banks	Silurian (Rhuddanian)	Arndell Sandstone	Australia	A member of the family Scutelluidae.
Kettneraspis plumula^[124]	Sp. nov	Valid	Flick & Flick	Devonian (Eifelian)		Germany	A member of the family Odontopleuridae.
Kongqiaoheia sarytumensis^[125]	Sp. nov	Valid	Ghobadi Pour	Ordovician		Kazakhstan
Liostracina fuluensis^[126]	Sp. nov	Valid	Yang et al.	Cambrian (Guzhangian)	Longha Formation	China	A member of Ptychopariida belonging to the family Liostracinidae.
Mackenzieaspis^[109]	Gen. et 2 sp. nov	Valid	Handkamer & Pratt in Handkamer, Pratt & MacNaughton	Cambrian		Canada	Genus includes new species M. parallelispinosa and M. divergens.
Malongullia sinensis^[110]	Sp. nov	In press	Wei & Zhou in Wei et al.	Ordovician (Katian)	Daduhe Formation	China
Malongullia zhenxiongensis^[110]	Sp. nov	In press	Wei & Zhou in Wei et al.	Ordovician (Katian)	Daduhe Formation	China
Mexicaspidella^[109]	Nom. nov	Not Valid	Handkamer & Pratt in Handkamer, Pratt & MacNaughton	Cambrian		Mexico United States	A replacement name for Mexicaspis Lochman (1948). Not valid because prior replacement: Caspimexis Özdikmen 2005 in: Paleontological Journal 39(5):563-564
Miaopopsis sokyrensis^[125]	Sp. nov	Valid	Ghobadi Pour	Ordovician		Kazakhstan
Minicryphaeus suavius^[113]	Sp. nov	Valid	Van Viersen & Kloc	Devonian	Seheb El Rhassel Group	Morocco	A member of the family Acastidae.
Mynaralaspis^[125]	Gen. et sp. nov	Valid	Ghobadi Pour	Ordovician		Kazakhstan	Genus includes new species M. perforata.
Needmorella^[127]	Gen. et sp. nov	Valid	Holloway & Scott	Devonian	Needmore Shale	United States (( Pennsylvania ( Virginia West Virginia)	A member of the family Dalmanitidae belonging to the subfamily Synphoriinae. The type species is N. simoni.
Niuchangella agastor^[112]	Sp. nov	Valid	Holloway, Banks & Banks	Silurian (Rhuddanian)	Arndell Sandstone	Australia	A member of the family Brachymetopidae.
Oenonella otherfellersorum^[128]	Sp. nov	Valid	Adrain & Fortey	Ordovician (Darriwilian)	Table Cove Formation	Canada ( Newfoundland and Labrador)
Oenonella wasisnamei^[128]	Sp. nov	Valid	Adrain & Fortey	Ordovician (Darriwilian)	Table Cove Formation	Canada ( Newfoundland and Labrador)
Pedinopariops? requadti^[124]	Sp. nov	Valid	Flick & Flick	Devonian (Eifelian)		Germany	A member of the family Phacopidae.
Peltura hutchinsoni^[129]	Sp. nov	Valid	Nguyen, Westrop & Landing	Cambrian (Furongian)	Chelsey Drive Group	Canada ( Nova Scotia)	An olenid trilobite.
Peltura undulata^[130]	Sp. nov		Nielsen et al.	Cambrian (Furongian)	Alum Shale Formation	Norway Sweden	An olenid trilobite.
Pepodes^[112]	Gen. et sp. nov	Valid	Holloway, Banks & Banks	Silurian (Rhuddanian)	Arndell Sandstone	Australia	A member of the family Illaenidae. The type species is P. agrestis.
Persiax^[131]	Nom. nov	Valid	Lerosey-Aubril & Deshmukh	Carboniferous (Tournaisian)		Iran	A replacement name for Persia Lerosey-Aubril (2012).
Platykardiapyge^[113]	Gen. et comb. nov	Valid	Van Viersen & Kloc	Devonian	Ihandar Formation	Morocco	Genus in Acastidae. The type species is P. maderensis; genus also includes P. aequisulcata.
Pricyclopyge keralensis^[125]	Sp. nov	Valid	Ghobadi Pour	Ordovician		Kazakhstan
Prosaukia xiaoxianensis^[120]	Sp. nov	Valid	Lei & Peng	Cambrian (Furongian)	Chaumitien Formation	China	A saukiid trilobite.
Sahtuia^[109]	Gen. et sp. nov	Valid	Handkamer & Pratt in Handkamer, Pratt & MacNaughton	Cambrian		Canada	Genus includes new species S. carcajouensis.
Salterocoryphe? bailliei^[112]	Sp. nov	Valid	Holloway, Banks & Banks	Ordovician (Katian)	Arndell Sandstone	Australia	A member of the family Calymenidae belonging to the subfamily Colpocoryphinae.
Sculptoproetus (Macroblepharum) bithynicus^[124]	Sp. nov	Valid	Flick & Flick	Devonian (Emsian)		Turkey	A member of the family Proetidae belonging to the subfamily Cornuproetinae.
Shumardia karasaiensis^[125]	Sp. nov	Valid	Ghobadi Pour	Ordovician		Kazakhstan
Taklamakania paucisegmentatus^[110]	Sp. nov	In press	Wei & Zhou in Wei et al.	Ordovician (Katian)	Daduhe Formation	China
Toxotiformis^[132]	Gen. et comb. et 5 sp. nov	Valid	Makarova	Cambrian		Russia	The type species is "Toxotis" venustus Lazarenko (1968); genus also includes new species T. artus, T. kotuyensis, T. nelegensis, T. tuberculosus and T. ventosus.
Triarthrus akkermensis^[125]	Sp. nov	Valid	Ghobadi Pour	Ordovician		Kazakhstan
Vietnamia hyron^[112]	Sp. nov	Valid	Holloway, Banks & Banks	Ordovician (Katian)	Arndell Sandstone	Australia	A member of the family Calymenidae belonging to the subfamily Reedocalymeninae.

Trilobite research

A study on patterns of segment allocation and expression in the bodies of trilobites throughout their evolutionary history is published by Hopkins & To (2022), who argue that neither taxonomic turnover nor enrolment behaviour of trilobites can sufficiently explain the studied changes of segmentation patterns.^[133]
A study on the early evolutionary history of trilobites is published by Holmes & Budd (2022), who argue that the first appearance datum of trilobites in the fossil record closely reflects their evolutionary origins, and that there is no compelling evidence to suggest an extended cryptic evolutionary history for this group.^[134]
Bicknell et al. (2022) describe malformed trilobite specimens from the Cambrian Beetle Creek Formation (Australia), Chisholm Formation (Nevada, United States) and Wheeler Formation (Utah, United States) and from the Ordovician Llanfawr Mudstones (Wales, United Kingdom), and attempt to determine the origin of the studied injuries.^[135]
A study on the injured specimens of Redlichia takooensis and Redlichia rex from the Cambrian Emu Bay Shale (Australia) is published by Bicknell et al. (2022), who argue that R. rex was likely the chief producer of the injuries in the studied specimen and of large shelly coprolites in the Emu Bay Shale biota, and represents one of the earliest cannibalistic trilobites.^[136]
Losso & Ortega-Hernández (2022) report evidence of the presence of significantly modified and reduced endopodites underneath the seventh thoracic and first pygidial tergites of Olenoides serratus and interpret these appendages as likely functional analogs to claspers.^[137]
A study on the phylogenetic relationships of members of the olenid group Hypermecaspidinae is published by Monti, Tortello & Confalonieri (2022).^[138]
Revision of Ordovician trilobite collections from Shan State (Myanmar) and Yunnan (China), first described by F.R.C. Reed, is published by Fortey, Wernette & Hughes (2022).^[139]
A study on the growth and mortality of Triarthrus eatoni, reevaluating the data presented by Cisne (1973),^[140] is published by Pauly & Holmes (2022).^[141]
Edgecombe & Fortey (2022) describe a specimen of Asaphellus tataensis from the Fezouata Formation (Morocco) preserved with antennae bearing a series of round, dome-shaped organs of uncertain homology and function, larger than sensilla on the antennae of other arthropods.^[142]
A study on the degree and structure of modularity in the heads of Calyptaulax annulata and Cloacaspis senilis is published by Vargas-Parra & Hopkins (2022), who consider the best modularity models to be those in which the eyes and anteriormost cranidium formed a single module, or belonged to two modules that highly covaried relative to other modules.^[143]
Bicknell & Smith (2022) seven new abnormal specimens of Odontopleura (Sinespinaspis) markhami from the Silurian (Telychian) Cotton Formation (Australia), interpreting their abnormalities as teratological developments through genetic malfunctions, and evaluate likely causes of abnormalities in Silurian trilobite specimens in general.^[144]
A study on the distribution patterns of Devonian trilobites from Morocco and northwestern Algeria through time and space is published by Bault, Crônier & Bignon (2022).^[145]
A study on the morphological diversity and possible relationship between morphology and environmental and/or ecological factors in Devonian trilobites from North Africa is published by Bault, Crônier & Monnet (2022).^[146]
A study on changes in global distribution of trilobites during the late Paleozoic is published by Brezinski (2022).^[147]
A study on functional morphology, coaptation and palaeoecology of selected acastid trilobites was published by Van Viersen & Kloc (2022).^[113]
A study on the morphological diversity of cephalic sclerites of asteropygine acastids throughout their evolutionary history is published by Martin et al. (2022).^[148]

Other arthropods

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acheronauta^[149]	Gen. et sp. nov	Valid	Pulsipher et al.	Silurian	Waukesha Lagerstätte	United States ( Wisconsin)	A vermiform arthropod, possibly a basal member of Mandibulata. Genus includes new species A. stimulapis.
Astutuscaris^[150]	Gen. et sp. nov	Valid	Jiao & Du	Cambrian Stage 4	Wulongqing Formation	China	A member of Euarthropoda of uncertain affinities. The type species is A. bispinifer.
Balhuticaris^[151]	Gen. et sp. nov	Valid	Izquierdo-López & Caron	Cambrian	Burgess Shale	Canada ( British Columbia)	A bivalved arthropod belonging to the group Hymenocarina. The type species is B. voltae.
Cyrtoctenus bambachi^[152]	Sp. nov	Valid	Plotnick & Lamsdell	Carboniferous (Tournaisian)	Price Formation	United States ( Virginia)	A hibbertopterid eurypterid.
Erettopterus qujingensis^[153]	Sp. nov	Valid	Ma et al.	Silurian (Pridolí)	Yulongsi Formation	China	A pterygotid eurypterid.
Erratus^[154]	Gen. et sp. nov		Fu et al.	Cambrian Stage 3	Helinpu Formation	China	An early carapace-bearing arthropod. The type species is E. sperare.
Fengzhengia^[155]	Gen. et sp. nov	Valid	O'Flynn et al.	Cambrian Stage 3	Chiungchussu Formation	China	A member of Euarthropoda of uncertain affinities. The type species is F. mamingae.
Guangyuanolimulus^[156]	Gen. et sp. nov		Hu et al.	Permian-Triassic transition		China	A member of Xiphosura, possibly an early horseshoe crab. Genus includes new species G. shangsiensis.
Hibbertopterus lamsdelli^[157]	Sp. nov	In press	Braddy, Lerner & Lucas	Late Carboniferous		United States ( New Mexico)	A eurypterid.
Lunataspis borealis^[158]	Sp. nov	In press	Lamsdell et al.	Ordovician (Sandbian)	Gull River Formation	Canada ( Ontario)	A member of Xiphosurida.
Malongia^[159]	Gen. et sp. nov		Wang et al.	Devonian (Lochkovian)	Xiaxishancun Formation	China	A eurypterid belonging to the family Dolichopteridae. Genus includes new species M. mirabilis.
Masticaris^[160]	Gen. et sp. nov	Valid	Harvey & Butterfield	Cambrian Stage 4	Mount Clark Formation	Canada ( Northwest Territories)	An arthropod described on the basis of mandibles co-occurring with small carbonaceous fossils. The type species is M. fimbriata.
Monstrodesmus grimaldii^[161]	Sp. nov	Valid	Su, Cai & Huang	Cretaceous	Burmese amber	Myanmar	A millipede belonging to the family Trichopolydesmidae.
Paleolimulus mazonensis^[162]	Sp. nov	Valid	Bicknell, Naugolnykh & McKenzie	Carboniferous (Moscovian)	Carbondale Formation	United States ( Illinois)	A member of Xiphosura belonging to the family Paleolimulidae.
Parahughmilleria fuea^[163]	Sp. nov		Ma et al.	Devonian (Lochkovian)	Xiaxishancun Formation	China	An adelophthalmid eurypterid.
Pterygotus wanggaii^[163]	Sp. nov		Ma et al.	Devonian (Lochkovian)	Xiaxishancun Formation	China	A pterygotid eurypterid.
Tomlinsonus^[164]	Gen. et sp. nov	Valid	Moysiuk et al.	Ordovician (Katian)	Kirkfield Formation	Canada ( Ontario)	A marrellomorph arthropod. The type species is T. dimitrii.
Vermontcaris^[165]	Gen. et sp. nov	Valid	Pari, Briggs & Gaines	Cambrian Stage 4		United States ( Vermont)	A bivalved arthropod. Genus includes new species V. montcalmi.

Redescription of Chuandianella ovata, based on data from new specimens from the Yu’anshan Member of the Chiungchussu Formation (Cambrian Stage 3; Yunnan, China) preserving unprecedented details of their soft anatomy, is published by Zhai et al. (2022).^[166]
A study on the ventral aspect of head organization of Jianfengia multisegmentalis, and on its evolutionary significance, is published by Zhang et al. (2022).^[167]
Redescription of Triopus draboviensis is published by Van Roy, Rak & Fatka (2022), who also provide a revised diagnosis for Cheloniellida, and exclude Parioscorpio venator from this clade.^[168]
Redescription of the ventral morphology of Retifacies abnormalis and a study on the implications of this taxon for the knowledge of the relationships and evolution of Cambrian artiopods is published by Zhang et al. (2022).^[169]
A study on the appendicular organization in Pygmaclypeatus daziensis and on its ecological and evolutionary implications is published by Schmidt et al. (2022).^[170]
Description of the organization of the central nervous system of a specimen of Mollisonia symmetrica from the Burgess Pass (Burgess Shale; British Columbia, Canada) is published by Ortega-Hernández et al. (2022).^[171]
A study on the evolutionary stability in the history of fossil and living xiphosurids is published by Bicknell et al. (2022).^[172]
Revision of Australian xiphosurids Austrolimulus fletcheri, Dubbolimulus peetae, Tasmaniolimulus patersoni and Victalimulus mcqueeni, and a study on the temporal range of these taxa is published by Bicknell et al. (2022), who reinterpret T. patersoni as living in the Triassic rather than Permian.^[173]
New specimen of Vaderlimulus tricki, providing new information on the anatomy of this xiphosuran and representing the first record of muscles in an austrolimulid reported to date, is described from the Olenekian Thaynes Group (Idaho, United States) by Lerner & Lucas (2022).^[174]
A study on the ontogenetic stages, allometry and ecology of Paleolimulus kunguricus is published by Naugolnykh & Bicknell (2022).^[175]
A study on the anatomy of the chelicerae of Slimonia acuminata, based on data from a new specimen, is published by Lamsdell (2022).^[176]
Braddy & Gass (2022) redescribe tracks from the Ordovician Martinsburg Formation (New York, United States) assigned to the ichnotaxon Palmichnium gallowayi, attribute these tracks to a medium-sized stylonurid eurypterid, and interpret them as the earliest trace fossil evidence for mass migrations of eurypterids into nearshore environments to molt and mate.^[177]
Biomechanical study of the chelicerae of pterygotid eurypterids is published by Bicknell et al. (2022), who argue that pterygotid chelicerae were functionally analogous to scorpion chelae, and that Erettopterus bilobus and Pterygotus anglicus had a generalised diet and were apex predators of their ecosystems, while Acutiramus bohemicus was adapted to piercing and slicing the cuticle of other eurypterids, and Jaekelopterus rhenaniae was adapted to capturing large, highly mobile, armoured prey.^[178]
New fossil material of Tuzoia with exceptionally preserved soft tissues is described from the Cambrian Burgess Shale (Canada) by Izquierdo-López & Caron (2022), who interpret this arthropod as adapted to predation or scavenging while swimming along the seafloor, and interpret it as an early member of Hymenocarina.^[179]
A study on the functional morphology of Ercaicunia multinodosa, aiming to determine the posture used by this arthropod to overcome resistance and to obtain most lift while sliding in the water column, is published by Li et al. (2022).^[180]

General research

Review of the paleontological, phylogenomic and molecular clock evidence pertaining to the possibly Cambrian terrestrialization of the arthropods is published Tihelka et al. (2022).^[181]

References

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^ ^a ^b ^c ^d De Francesco Magnussen, I.; Müller, S. P.; Hammel, J. U.; Kotthoff, U.; Harms, D. (2022). "Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane". Zoological Journal of the Linnean Society. 196 (2): 792–844. doi:10.1093/zoolinnean/zlac034.
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^ Kolesnikov, V. B.; Turbanov, I. S.; Eskov, K. Yu.; Propistsova, E. A.; Bashkuev, A. S. (2022). "First non-amber Mesozoic pseudoscorpion from Upper Triassic deposits of eastern Europe, with a description of two new fossil subfamilies (Arachnida, Pseudoscorpiones, Feaellidae)". Papers in Palaeontology. 8 (5): e1466. doi:10.1002/spp2.1466. S2CID 253137909.
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^ Xuan, Q.; Cai, C.; Huang, D. (2022). "A new Palaeoburmesebuthidae scorpion from mid-Cretaceous Burmese amber (Arachnida: Scorpiones: Buthoidea)". Cretaceous Research. 134: Article 105165. Bibcode:2022CrRes.13405165X. doi:10.1016/j.cretres.2022.105165. S2CID 246664669.
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[34] Ferratges, F. A.; Zamora, S.; Aurell, M. (2022). "Systematics and paleoecology of a new species of Varunidae H. Milne Edwards, 1853 (Decapoda: Brachyura) from the lower Eocene of Spain". Journal of Crustacean Biology. 42 (2): ruac013. doi:10.1093/jcbiol/ruac013.

[Basidromilites-35] Artal, P.; Ferratges, F. A.; van Bakel, B. W. M.; Zamora, S. (2022). "A highly diverse dromioid crab assemblage (Decapoda, Brachyura) associated with pinnacle reefs in the lower Eocene of Spain". Journal of Paleontology. 96 (3): 591–610. Bibcode:2022JPal...96..591A. doi:10.1017/jpa.2021.114. S2CID 246068493.

[36] Mazancourt, V.; Wappler, T.; Wedmann, S. (2022). "Exceptional preservation of internal organs in a new fossil species of freshwater shrimp (Caridea: Palaemonoidea) from the Eocene of Messel (Germany)". Scientific Reports. 12 (1). 18114. Bibcode:2022NatSR..1218114D. doi:10.1038/s41598-022-23125-9. PMC 9613706. PMID 36302944.

[37] Mazancourt, V.; Wappler, T.; Wedmann, S. (2023). "Author Correction: Exceptional preservation of internal organs in a new fossil species of freshwater shrimp (Caridea: Palaemonoidea) from the Eocene of Messel (Germany)". Scientific Reports. 13 (1). 5943. Bibcode:2023NatSR..13.5943D. doi:10.1038/s41598-023-32986-7. PMC 10097716. PMID 37046034.

[Eostenetrium-38] Wilson, G. D. F.; Morel, N. (2022). "Isopod crustacean fossils from the Cenomanian stratotype: five new species in suborders Cymothoida, Asellota and Valvifera". Annales de Paléontologie. 108 (1): Article 102538. Bibcode:2022AnPal.10802538W. doi:10.1016/j.annpal.2022.102538. S2CID 249114110.

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[41] Santana, W.; Tavares, M.; Martins, C. A. M.; Melo, J. P. P.; Pinheiro, A. P. (2023). "Validation of Chronocancer camilosantanai† Santana, Tavares, Martins, Melo & Pinheiro (Crustacea, Decapoda, Brachyura) from the Romualdo Formation, Araripe Sedimentary Basin, Brazil". Papéis Avulsos de Zoologia. 63. e202363014. doi:10.11606/1807-0205/2023.63.014. S2CID 258596942.

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[Sienalia-43] Garassino, A.; Pasini, G.; Pizzolato, F. (2022). "A review of some leucosiid crabs from the lower Pleistocene beds of Poggi Gialli (Tuscany, central Italy)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 304 (3): 263–273. doi:10.1127/njgpa/2022/1070. S2CID 250010128.

[Eoacmaeopleura-44] De Angeli, A.; Bellin, B. (2022). "Nuovi crostacei brachiuri dell'Eocene di cava "Main" di Arzignano (Vicenza, Italia nordorientale)". Lavori – Società Veneziana di Scienze Naturali. 47: 93–101.

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[Eureotropis-47] Mendes, I. D.; Santana, W.; Carvalho, I. S. (2022). "Fossil hymenosomatid crabs (Crustacea: Decapoda) from the Lower Cretaceous of NE Brazil". Cretaceous Research. 134: Article 105174. Bibcode:2022CrRes.13405174M. doi:10.1016/j.cretres.2022.105174. S2CID 246894368.

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[55] Beschin, C.; De Angeli, A. (2022). "Stimdromia conternoi n. sp., (Decapoda, Brachyura, Dromiidae) dell'Eocene di Nanto (Monti Berici, Vicenza, Italia nordorientale)". Studi e Ricerche - Associazione Amici del Museo - Museo Civico "G. Zannato" Montecchio Maggiore (Vicenza). 29: 5–8.

[56] Audo, D.; Charbonnier, S. (2022). "Teruzzicheles popeyei, an early Sinemurian polychelid lobster from the Osteno Lagerstätte (Italy)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 306 (3): 187–194. doi:10.1127/njgpa/2022/1103. S2CID 254820598.

[57] Garassino, A.; Pasini, G.; Nazarkin, M. V. (2022). "A new caridean shrimp (Crustacea, Decapoda) from the middle–late Miocene of Sakhalin Island, Russia". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 303 (1): 1–9. doi:10.1127/njgpa/2022/1031. S2CID 246449498.

[CRViaia-58] Gašparič, R.; van Bakel, B. W. M.; Fraaije, R. H. B.; Jagt, J. W. M. (2022). "First record of viaiid crabs (Decapoda, Brachyura) from the Lower Cretaceous of Slovenia, with the description of two new species". Cretaceous Research. 138: Article 105293. Bibcode:2022CrRes.13805293G. doi:10.1016/j.cretres.2022.105293. S2CID 250034465.

[59] Pazinato, P. G.; Haug, C.; Leipner, A.; Haug, J. T. (2022). "Pygocephalomorphan crustaceans further emphasise the similarities between the Carboniferous Piesberg quarry in Germany and the Mazon Creek Lagerstätte in North America". Palaeontologia Electronica. 25 (1): Article number 25.1.2A. doi:10.26879/1051.

[60] Tshudy, D. M.; Hyžný, M.; Kočová Veselská, M.; Jagt, J. W. M. (2022). "Taxonomic revision of the extinct clawed lobster genus Oncopareia Bosquet, 1854 (Decapoda, Astacidea, Nephropidae)". Palaeontologia Electronica. 25 (2): Article number 25.2.a20. doi:10.26879/1190.

[61] Fraaije, R. H. B.; Van Bakel, B. W. M.; Jagt, J. W. M.; Charbonnier, S.; Schweigert, G.; Garcia, G.; Valentin, X. (2022). "The evolution of hermit crabs (Crustacea, Decapoda, Anomura, Paguroidea) on the basis of carapace morphology: a state-of-the-art-report". Geodiversitas. 44 (1): 1–16. doi:10.5252/geodiversitas2022v44a1. S2CID 245907939.

[62] Jones, A. R.; Schweitzer, C. E.; Feldmann, R. M. (2022). "Sexual dimorphism and rare intersex individuals in Cretaceous (Maastrichtian) Dakoticancer Rathbun, 1917 (Decapoda: Brachyura: Dakoticancroida)". Journal of Crustacean Biology. 42 (1): ruac010. doi:10.1093/jcbiol/ruac010.

[63] Ossó, À.; Hyžný, M.; Amoudji, Y. Z.; Kassegne, K. E.; Johnson, A. K. C.; Da Costa, P. Y. D.; Bondant, C.; Perrier, V.; Hautier, L.; Guinot, G. (2022). "Pleolobites (Decapoda: Brachyura) from the Paleogene of Africa revisited, with implications on taxonomy of fossil portunoid crabs" (PDF). Annales de Paléontologie. 108 (1): Article 102541. Bibcode:2022AnPal.10802541O. doi:10.1016/j.annpal.2022.102541. S2CID 249972329.

[64] Feldmann, R. M.; Schweitzer, C. E. (2022). "Unique occurrence of the Miocene Trichopeltarion greggi Dell, 1969 (Decapoda: Brachyura: Trichopeltariidae): potential documentation of maternal care". Journal of Crustacean Biology. 42 (1): ruab080. doi:10.1093/jcbiol/ruab080.

[PJ562-65] Tesakova, E. M. (2022). "Late Bajocian and early Bathonian ostracods of the Russian Plate. Part I: genera Plumhoffia Brand and Aaleniella Plumhoff". Paleontological Journal. 56 (2): 173–186. doi:10.1134/S0031030122020125. S2CID 248302847.

[PJ563-66] Tesakova, E. M. (2022). "Late Bajocian and early Bathonian ostracods of the Russian Plate. Part II. Genera: Procytherura Whatley, Pseudohutsonia Wienholz, Acrocythere Neale, Nanacythere Herrig and Trachycythere Triebel et Klingler". Paleontological Journal. 56 (3): 250–262. doi:10.1134/S0031030122030145. S2CID 249627638.

[RIPS1282Foreletal-67] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Forel, M.-B.; Del Piero, N.; Rigaud, S.; Martini, R. (2022). "Ostracods from the Late Triassic (Norian) of Yukon, canada: new taxonomic and palaeobiogeographic insights". Rivista Italiana di Paleontologia e Stratigrafia. 128 (2): 325–368. doi:10.54103/2039-4942/15439. S2CID 248900869.

[Asculdoracythereis-68] ^ ^a ^b ^c ^d ^e ^f Puckett, T. M.; Hunt, G. (2022). "New Taxa of Marine Ostracods (Anticytherideinae, n. subfam.) from the Upper Cretaceous (Campanian and Maastrichtian) of Mississippi, Alabama and Tennessee, U. S. Gulf Coastal Plain". Micropaleontology. 68 (5): 433–504. Bibcode:2022MiPal..68..433P. doi:10.47894/mpal.68.5.01. S2CID 252058741.

[RM77Sciutoetal-69] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k Sciuto, F.; Baldanza, A.; Reitano, A. (2022). "First report of Early Pleistocene ostracods from the Agira sedimentary succession (Central Eastern Sicily, Italy)". Revue de Micropaléontologie. 77. 100693. Bibcode:2022RvMic..7700693S. doi:10.1016/j.revmic.2022.100693. S2CID 252688669.

[70] Song, J.; Guo, W.; Sun, Y.; Ma, J.; Huang, J.; Zhang, Y.; Qie, W. (2022). "First record of the latest Devonian ostracods from the Xainza region, Tibet, China: implications on palaeoenvironment and palaeobiogeography". Palaeoworld. in press. doi:10.1016/j.palwor.2022.01.002. S2CID 245994353.

[71] Mojon, P.-O.; De Kaenel, E. (2022). "New paleontological and biostratigraphical data (calcareous nannofossils, ostracods, brachiopods), correlations and lithostratigraphic units in the Urgonian facies (latest Hauterivian-Barremian) of the Swiss and French Jura Mountains: the Falaises Member and the Saars Formation (former "Gorges de l'Orbe Formation")". Swiss Journal of Geosciences. 115: Article 17. doi:10.1186/s00015-022-00416-x. S2CID 248574040.

[Fortistriginglymus-72] ^ ^a ^b ^c ^d ^e ^f McDonald, A. P.; Warne, M. T. (2022). "Latest Miocene ostracods from the Bookpurnong Formation in the Murray Basin of southeastern Australia: shallow marine migrants into an epicontinental sea". Alcheringa: An Australasian Journal of Palaeontology. 46 (3–4): 301–339. Bibcode:2022Alch...46..301M. doi:10.1080/03115518.2022.2133169. S2CID 253934707.

[PJ564Tesakova-73] Tesakova, E. M. (2022). "Late Bajocian and early Bathonian ostracods of the Russian Plate. Part III. Genera Camptocythere Triebel and Procytheridea Peterson". Paleontological Journal. 56 (4): 400–411. doi:10.1134/S003103012204013X. S2CID 251519387.

[PJ565-74] Tesakova, E. M.; Seltser, V. B. (2022). "New Callovian–lower Oxfordian ostracods from the Russian Plate (genera Camptocythere Triebel and Procytherura Whatley) and their stratigraphic significance". Paleontological Journal. 56 (5): 526–547. doi:10.1134/S0031030122050124. S2CID 252717464.

[BG973ostracods-75] Poursalehi, F.; Voldman, G. G.; Bahrami, A.; Salas, M. J.; Ameri, H. (2022). "New data on conodonts and ostracods of the Katkoyeh Formation (Lower-Upper Ordovician) at the Banestan Section of East-Central Iran: biostratigraphical and palaeobiogeographical significance". Bulletin of Geosciences. 97 (3): 363–384. doi:10.3140/bull.geosci.1847. S2CID 251238719.

[EJT804-76] Guillam, E.; Forel, M.-B.; Song, J.; Crasquin, S. (2022). "Late Devonian–early Carboniferous ostracods (Crustacea) from South China: taxonomy, diversity and implications". European Journal of Taxonomy (804): 1–62. doi:10.5852/ejt.2022.804.1689. S2CID 247488088.

[RBP251-77] Linhares, A. P.; Ramos, M. I. F. (2022). "Neogene ostracods from the Solimões Formation (Atalaia do Norte, Amazonas State, Brazil), with the description of two new species". Revista Brasileira de Paleontologia. 25 (1): 61–75. doi:10.4072/rbp.2022.1.05. S2CID 248172465.

[JSAESCyprideis-78] Sousa, F. S. D. E.; Ramos, M. I. F. (2022). "Taxonomic study of the genus Cyprideis JONES, 1857 from the Pebas Formation (Miocene), Iquitos (Peru), with description of three new species". Journal of South American Earth Sciences. 123. 104126. doi:10.1016/j.jsames.2022.104126. S2CID 254090098.

[RM77Guzmanetal-79] Guzmán, J.; Piovesan, E. K.; Almeida-Lima, D. S.; Sousa, A. J.; Neumann, V. H. M. L. (2022). "Aptian ostracods from the Santana Group, Araripe Basin, Brazil". Revue de Micropaléontologie. 77. 100694. Bibcode:2022RvMic..7700694G. doi:10.1016/j.revmic.2022.100694. S2CID 252732442.

[Klyasinella-80] Melnikova, L. M.; Mirantsev, G. V.; Terentiev, S. S.; Anekeeva, G. A. (2022). "Ostracods of the Gryazno Formation (Upper Ordovician, Sandbian Stage and Idavere Regional Stage) in the west of Leningrad Region". Paleontological Journal. 56 (5): 510–525. doi:10.1134/S0031030122050082. S2CID 252717674.

[81] Wang, H.; Matzke-Karasz, R.; Horne, D. J. (2022). "Mid-Cretaceous coastal amber forest palaeoenvironment revealed by exceptionally preserved ostracods from an extant lineage". Fossil Record. 25 (1): 147–172. doi:10.3897/fr.25.e84604.

[82] Tesakova, E. M. (2022). "Late Bajocian and early Bathonian ostracods of the Russian Plate. Part IV. The lineage Glyptocythere aspera (Khabarova) → G. bathonica sp. nov". Paleontological Journal. 56 (6): 646–662. doi:10.1134/S0031030122060120. S2CID 254248807.

[Karlingrella-83] Groos-Uffenorde, H.; Schindler, E.; Becker, R. T.; Dojen, C.; Brocke, R.; Jansen, U. (2022). "Late Early Devonian ostracodes from the Torkoz area (SW Morocco) and the Emsian/Eifelian boundary". PalZ. 96 (4): 689–747. doi:10.1007/s12542-022-00603-z. S2CID 248969995.

[84] McMenamin, M. A. S. (2022). "Early Carboniferous Ostracods (Crustacea) from Death Valley, California, USA". Geosciences. 12 (8). 300. Bibcode:2022Geosc..12..300M. doi:10.3390/geosciences12080300.

[85] Forel, M.-B. (2022). "Thoughts on the Late Paleozoic-Early Mesozoic records of deep-sea ostracods". Micropaleontology. 68 (3): 217–231. Bibcode:2022MiPal..68..217F. doi:10.47894/mpal.68.3.02. S2CID 248565579.

[RM76-86] Forel, M.-B.; Kolar-Jurkovšek, T.; Jurkovšek, B. (2022). "Marine ostracods from the Ladinian of Slovenia". Revue de Micropaléontologie. 76. 100691. Bibcode:2022RvMic..7600691F. doi:10.1016/j.revmic.2022.100691. S2CID 252152269.

[Zootaxa51594-87] ^ ^a ^b ^c ^d ^e ^f ^g Tomé, M. E.; Araripe, R.; Oliveira, D.; Barreto, A.; Prado, L.; Pedrosa, F.; Pereira, P.; Nascimento, L. R.; Ng, C. (2022). "Early Cretaceous Ostracoda (Crustacea) from south-central Araripe Basin, Brazil, with descriptions of seven new species". Zootaxa. 5159 (4): 535–557. doi:10.11646/zootaxa.5159.4.4. PMID 36095533. S2CID 250241849.

[PalaeoworldZhang-88] Zhang, Y.-C. (2022). "Late Darriwilian–early Katian (Ordovician) ostracods of northeastern Yunnan, Southwest China". Palaeoworld. 32 (2): 303–332. doi:10.1016/j.palwor.2022.04.002. S2CID 248332032.

[89] Antonietto, L. S.; Eaton, J. G.; Park Boush, L. E. (2022). "A newly found ostracod fauna from the Paleocene? - Eocene Claron Formation at Sweetwater Creek, Utah, USA with description of a new species". Micropaleontology. 68 (4): 375–388. Bibcode:2022MiPal..68..375A. doi:10.47894/mpal.68.4.02.

[Paranacythere-90] Bergue, C. T.; Maranhão, M. S. A. S.; Ng, C.; Naumcheva, M. A. (2022). "A Permian mixohaline ostracod association of the Teresina Formation, Paraná Basin, Brazil". Revista Brasileira de Paleontologia. 25 (1): 24–37. doi:10.4072/rbp.2022.1.02. hdl:10183/240424. S2CID 248173978.

[Zootaxa51416-91] Piovesan, E. K.; Melo, G. D.; Cabral, M. C.; Guzmán, J. (2022). "New Early Cretaceous species of Zonocypris G. W. Müller, 1898 (Crustacea: Ostracoda) from the Crato Formation, Brazil". Zootaxa. 5141 (6): 581–591. doi:10.11646/zootaxa.5141.6.4. PMID 36095772. S2CID 249221037.

[92] Karasawa, H.; Amano, K. (2022). "New records for pedunculate barnacles (Cirripedia: Thoracica) from the Miocene Higashibessho Formation in Toyama Prefecture, central Japan". Bulletin of the Mizunami Fossil Museum. 49: 123–127. doi:10.50897/bmfm.49.0_123.

[93] Collareta, A.; Bosselaers, M.; Holroyd, P. A.; Dineen, A. (2022). "A forgotten cirripedological gem: a new species of whale barnacle of the genus Cetopirus from the Pleistocene of the United States West Coast". Earth and Environmental Science Transactions of the Royal Society of Edinburgh: 1–8. doi:10.1017/S1755691022000214. S2CID 253404871.

[PalaeoworldDabashanella-94] Zhang, H.-Q. (2022). "A new assemblage of dabashanellids (Crustacea: Phosphatocopida) from Cambrian Stage 3 of western Hubei Province, South China". Palaeoworld. 32: 14–26. doi:10.1016/j.palwor.2022.06.004. S2CID 250080273.

[95] Peel, J. S. (2022). "The oldest tongue worm: a stem-group pentastomid arthropod from the early middle Cambrian (Wuliuan Stage) of North Greenland (Laurentia)". GFF. 144 (2): 97–105. Bibcode:2022GFF...144...97P. doi:10.1080/11035897.2022.2064543. S2CID 249028918.

[Thuringiops-96] Werneburg, R.; Schneider, J. W. (2022). "New branchiopod crustaceans from the late Carboniferous and early Permian of the Thuringian Forest Basin, Germany, with a review of Permian notostracans from the Lodève basin, France". Semana. Naturwissenschaftliche Veröffentlichungen des Naturhistorischen Museums Schloss Bertholdsburg Schleusingen. 37: 57–103.

[97] Van Houte, E.; Hegna, T. A.; Butler, A. D. (2022). "A new genus and species of ?parthenogenic anostracan (Pancrustacea, Branchiopoda, ?Thamnocephalidae) from the Lower Cretaceous Koonwarra Fossil Bed in Australia". Alcheringa: An Australasian Journal of Palaeontology. 46 (2): 180–187. Bibcode:2022Alch...46..180V. doi:10.1080/03115518.2022.2048267. S2CID 247807263.

[98] Teng, X.; Li, G.; Zhang, Y.-Z. (2022). "Linglongtaestheria (Spinicaudata) from the Upper Jurassic of Qinglong, Hebei Province, northeastern China". Palaeoworld. doi:10.1016/j.palwor.2022.11.008. S2CID 254449513.

[99] Mychko, E. V.; Alekseev, A. S.; Zaytseva, E. L.; Schweitzer, C. E.; Feldmann, R. M. (2022). "New finding of Cyclida (Crustacea) from Mississippian and not‑cyclidan from Permian of Russia". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 305 (1): 75–85. doi:10.1127/njgpa/2022/1077. S2CID 250706723.

[100] Gale, Andy (2022-05-12). "A giant barnacle (Crustacea, Cirripedia, Thoracica) from the Upper Cretaceous (Maastrichtian) of Canada revisited". Proceedings of the Geologists' Association. 133 (3): 212–217. Bibcode:2022PrGA..133..212G. doi:10.1016/j.pgeola.2022.04.002. ISSN 0016-7878. S2CID 248758434.

[101] Perreault, R. T.; Collareta, A.; Buckeridge, J. S. (2022). "A new species of the archaic "turtle barnacle" genus Protochelonibia (Coronuloidea, Chelonibiidae) from the upper Rupelian Chickasawhay Formation of Mississippi (U.S.A.)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 305 (3): 225–235. doi:10.1127/njgpa/2022/1087. S2CID 252744143.

[102] Shen, Y.; Wu, X. (2022). "Revision of the Devonian clam shrimp (Diplostraca: Spinicaudata) from the Canadian Arctic and discussion on the early morphotypes of the leaiins". Acta Palaeontologica Sinica. 61 (2): 217–225. doi:10.19800/j.cnki.aps.2021027.

[103] Li, G. (2022). "New spinicaudatan species (Crustacea) of the Lower Cretaceous (Valanginian–Hauterivian) Jehol Biota from northern Hebei, China". Cretaceous Research. 143. 105425. doi:10.1016/j.cretres.2022.105425. S2CID 253998860.

[104] Collareta, A.; Varas-Malca, R.; Bosio, G.; Urbina, M.; Coletti, G. (2022). "Ghosts of the Holobiont: Borings on a Miocene Turtle Carapace from the Pisco Formation (Peru) as Witnesses of Ancient Symbiosis". Journal of Marine Science and Engineering. 11 (1). 45. doi:10.3390/jmse11010045. hdl:10281/433198.

[:0-105] Zeng, Han; Zhao, Fangchen; Zhu, Maoyan (2022-09-07). "Innovatiocaris, a complete radiodont from the early Cambrian Chengjiang Lagerstätte and its implications for the phylogeny of Radiodonta". Journal of the Geological Society. 180. doi:10.1144/jgs2021-164. ISSN 0016-7649. S2CID 252147346.

[106] Moysiuk, J.; Caron, J.-B. (2022). "A three-eyed radiodont with fossilized neuroanatomy informs the origin of the arthropod head and segmentation". Current Biology. 32 (15): 3302–3316.e2. doi:10.1016/j.cub.2022.06.027. PMID 35809569. S2CID 250361698.

[Forteyaspis-107] Karim, T. S.; Adrain, J. M. (2022). "The phylogenetic affinity of the Ordovician trilobites Agerina, Forteyaspis gen. nov., and related genera, with new and revised species from Canada and the United States". Canadian Journal of Earth Sciences. 59 (3): 156–179. doi:10.1139/cjes-2021-0105. S2CID 246603380.

[Aguaditaspis-108] Randolfe, E. A.; Rustán, J. J.; Bignon, R. (2022). "New dalmanitid trilobite taxa from the Lower Devonian of Argentina: comments on spinosity in endemic taxa from southern high paleolatitudes". Comptes Rendus Palevol. 21 (2): 17–38. doi:10.5852/cr-palevol2022v21a2.

[Aitkenaspis-109] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Handkamer, N. M.; Pratt, B. R.; MacNaughton, R. B. (2022). "Biostratigraphy and paleoecology of the trilobite faunas from the Mount Clark and Mount Cap formations (early and middle Cambrian), eastern Mackenzie Mountains, northwestern Canada". Journal of Paleontology. 96 (Supplement S89): 1–47. Bibcode:2022JPal...96S...1H. doi:10.1017/jpa.2022.13. S2CID 249959858.

[PalaeoworldWeietal-110] Wei, X.; Wang, K.; Zhou, Z.-Q.; Cui, Y.-N.; Zhang, Z.-T.; Liu, J.-B. (2023). "A new late Katian (Late Ordovician) trilobite association from Zhenxiong, northeastern Yunnan, southwest China and its palaeoecological implications". Palaeoworld. 32 (2): 333–353. Bibcode:2023Palae..32..333W. doi:10.1016/j.palwor.2022.01.008. S2CID 246513780.

[Anebocephalus-111] Sundberg, F. A.; Webster, M. (2022). ""Ptychoparioid" trilobites of the Harkless Formation and Mule Spring Limestone (Cambrian Series 2, Stage 4), Clayton Ridge, Nevada". Journal of Paleontology. 96 (4): 886–920. Bibcode:2022JPal...96..886S. doi:10.1017/jpa.2021.124. S2CID 246656292.

[Auritolithus-112] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Holloway, D. J.; Banks, M. R.; Banks, D. M. (2022). "Late Ordovician and early Silurian trilobites from Tasmania". Bulletin of Geosciences. 97 (4): 397–444. doi:10.3140/bull.geosci.1868. S2CID 255365827.

[:2-113] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Viersen, Allart P. VAN; J. Kloc, Gerald (2022-12-09). "Functional morphology, coaptation and palaeoecology of Hollardops (Trilobita, Acastidae), with descriptions of new species and two new genera from the Devonian of Morocco". Geologica Belgica. 25 (3–4): 99–144. doi:10.20341/gb.2022.005. ISSN 1374-8505. S2CID 254532294.

[FS67-114] ^ ^a ^b ^c ^d ^e ^f ^g Cederström, P.; Geyer, G.; Ahlberg, P.; Nilsson, C. H.; Ahlgren, J. (2022). Ellipsocephalid trilobites from Cambrian Series 2 and Stage 4, with emphasis on the taxonomy, morphological plasticity and biostratigraphic significance of ellipsocephalids from Scania, Sweden. Fossils and Strata Series. Vol. 67. pp. 1–131. doi:10.18261/9788215065779-2022-01. ISBN 978-8-215-06576-2.

[115] Westrop, S. R.; Dengler, A. A. (2022). "A new Cambrian catillicephalid trilobite from the Shallow Bay Formation of western Newfoundland, Canada". Acta Palaeontologica Polonica. 67 (1): 27–33. doi:10.4202/app.00922.2021. S2CID 247904399.

[116] Collantes, L.; Pereira, S.; Mayoral, E.; Gozalo, R. (2022). "First report of Chelediscus Rushton, 1966 (Trilobita) from Western Gondwana, with description of a new species from the Cambrian Series 2 of Spain". Historical Biology: An International Journal of Paleobiology. 35 (9): 1701–1708. doi:10.1080/08912963.2022.2109966. S2CID 251576690.

[117] Crônier, C.; Waters, J. A. (2022). "Late Devonian (Famennian) phacopid trilobites from western Xinjiang, Northwest China". Palaeobiodiversity and Palaeoenvironments. 103 (2): 327–340. doi:10.1007/s12549-022-00547-x. S2CID 253075986.

[118] Zong, R.-W. (2023). "Variation in eye lenses of two new Late Devonian phacopid trilobites from western Junggar, NW China". Journal of Paleontology: 1–15. doi:10.1017/jpa.2023.31. S2CID 258981236.

[119] Zong, R.-W. (2023). "Variation in eye lenses of two new Late Devonian phacopid trilobites from western Junggar, NW China – CORRIGENDUM". Journal of Paleontology: 1. doi:10.1017/jpa.2023.42.

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[121] Fortey, R. A.; Gutiérrez-Marco, J. C. (2022). "Extraordinary Ordovician trilobite Fantasticolithus gen. nov. from Peru and its bearing on the trinucleimorph hypothesis". Papers in Palaeontology. 8 (2): e1423. doi:10.1002/spp2.1423. hdl:10261/266631. S2CID 247460358.

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[165] Pari, G.; Briggs, D. E. G.; Gaines, R. R. (2022). "The soft-bodied biota of the Cambrian Series 2 Parker Quarry Lagerstätte of northwestern Vermont, USA". Journal of Paleontology. 96 (4): 770–790. Bibcode:2022JPal...96..770P. doi:10.1017/jpa.2021.125. S2CID 246933116.

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