2012 in paleontology

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Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2012.

Plants

Gymnosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Calocedrus huashanensis	Sp. nov	Valid	Shi, Zhou, & Xie	Oligocene	Ningming Formation	China	An extinct species in the modern genus Calocedrus
Ginkgo cranei	Sp. nov	Valid	Zhou, Quan, & Liu	upper Paleocene	Sentinel Butte Formation, North Dakota	USA	An extinct species in the modern genus Ginkgo

Arthropods

Main article: 2012 in arthropod paleontology

Bryozoans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Admirandopora	Nom. nov.	Valid	Ariunchimeg	Lower Carboniferous		Mongolia	A replacement name for the genus Admiranda Ariunchimeg, 1996.
Argentinodictya	Gen. et sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A cryptostome bryozoan, a member of Ptilodictyina. The type species is Argentinodictya lenticulata.
Basyaylella	Gen. et sp. nov	In press	Zágoršek & Gordon	Late Tortonian	Mut Basin	Turkey	An ascophoran bryozoan. Its type species is Basyaylella elsae.
Chasmatopora rossae	Sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A phyloporinine, a species of Chasmatopora.
Lunostoma	Gen. et sp. nov	Valid	Ernst, Taylor, Bohatý & Wyse Jackson	Middle Devonian		Germany	A cryptostome bryozoan. The type species is Lunostoma pulchra.
Ostrovskia	Gen. et sp. nov	In press	Zágoršek & Gordon	Late Tortonian	Mut Basin	Turkey	An ascophoran bryozoan. Its type species is Ostrovskia triforamina.
Parvohallopora parvula	Sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A trepostome bryozoan, a species of Parvohallopora.
Ptilodictya intermedia	Sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A cryptostome bryozoan, a species of Ptilodictya.
Rorypora	Gen. et comb. nov.	Valid	Taylor	Jurassic		Switzerland	A new genus for "Diastopora" retiformis.
Trigonodictya parvula	Sp. nov	Valid	Ernst & Carrera	Late Ordovician (Sandbian)		Argentina	A cryptostome bryozoan, a species of Trigonodictya.

Brachiopods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Amydroptychus markowitzi	sp nov	Valid	Feldman et al.	Callovian	Mughanniyya Formation	Jordan	A tetrarhynchiid rhynchonellide, a species of Amydroptychus.
Askepasma saproconcha	sp nov	In press	Topper et al..	Lower Cambrian	Wirrapowie Limestone	Australia	A member of Paterinata, a species of Askepasma.
Bolilaspirifer riccardii	Sp. nov	Valid	Manceñido in Damborenea & Manceñido	Late Triassic		Argentina	A lepismatinid brachiopod, a species of Bolilaspirifer.
Cincinnetina	Gen, sp. et comb. nov	Valid	Jin	Late Ordovician		United States	A dalmanellid brachiopod, with C. multisecta, C. meeki, C. minnesotensis
Crurithyris tazawai	sp nov	In press	He et al..	Late Permian (Changhsingian)		China	An ambocoeliid brachiopod, a species of Crurithyris.
Daghanirhynchia susanae	Species	Valid	Feldman et al.	Callovian	Mughanniyya Formation	Jordan	A tetrarhynchiid rhynchonellide, a species of Daghanirhynchia.
Eodmitria anauris	sp nov	Valid	Oleneva	Late Devonian (middle Frasnian)		Russia	A cyrtospiriferid spiriferid, a species of Eodmitria.
Gatosella	Gen. et sp. nov	Valid	Benedetto	Middle Ordovician (early Darriwilian)	San Juan Formation	Argentina	A plectambonitoid brachiopod. The type species is Gatosella muricata.
Gracianella paulula	sp nov	In press	Baliński	Early Devonian	Khudykivtsi Beds	Ukraine	An atrypinid brachiopod, a species of Gracianella.
Plectodonta mariae pantherae	Subsp nov	In press	Baliński	Early Devonian	Khudykivtsi Beds	Ukraine	A sowerbyellid brachiopod, a subspecies of Plectodonta mariae.
Rotutaspirifer	Gen. et sp. nov	Valid	Oleneva	Late Devonian (middle Frasnian)		Russia	A cyrtospiriferid spiriferid. The type species is Rotutaspirifer rotutus.
Skenidioides tatyanae	sp nov	In press	Baliński	Early Devonian	Khudykivtsi Beds	Ukraine	A skenidiid brachiopod, a species of Skenidioides.

Molluscs

Newly named ammonites

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Alocolytoceras isztimeri	Sp nov	Valid	Galácz & Kassai	Bajocian		Hungary	A member of Lytoceratina, a species of Alocolytoceras.
Baidites	Gen. et comb. et sp. nov.	Valid	Brühwiler et al.	Early Triassic			A flemingitid ammonite. Contains "Ophiceras" crassicostatum, "Ophiceras" tenue and a new species Baidites hermanni.
Boreotrachyceras	Gen. et comb. nov	Valid	Konstantinov	Early Carnian		Russia	A trachyceratid, a new genus for "Protrachyceras" omkutchanicum Bytschkov.
Craspedites sachsi	Sp nov	Valid	Igolnikov	Berriasian		Russia	A species of Craspedites.
Curacoites	Gen. et sp. nov	Valid	Aguirre-Urreta & Rawson	Valanginian or early Barremian	Agrio Formation	Argentina	The type species is Curacoites rotundus.
Dimorphites noricus	Sp nov	Valid	Balini et al.	Late Triassic	Scillato Formation	Italy	A species of Dimorphites.
Dimorphoceratoides adamsi	Sp. nov	Valid	Work, Mason & Boardman	Pennsylvanian	Four Corners Formation	United States	A species of Dimorphoceratoides.
Dissimilites intermedius	Sp nov	In press	Lukeneder & Lukeneder	Barremian		Italy	An acrioceratid ancyloceratoid, a species of Dissimilites.
Galfettites kyrae	Sp nov	Valid	Brühwiler et al.	Early Triassic		Oman	A galfettitid ammonite, a species of Galfettites.
Galfettites omani	Sp nov	Valid	Brühwiler et al.	Early Triassic		Oman	A galfettitid ammonite, a species of Galfettites.
Gastrioceras magoffinense	Sp. nov	Valid	Work, Mason & Boardman	Pennsylvanian	Four Corners Formation	United States	A species of Gastrioceras.
Goudemandites	Gen. et sp. nov	Valid	Brühwiler et al.	Early Triassic		China Oman	A palaeophyllitid ammonite. The type species is Goudemandites sinensis.
Hildaites bisulcatus	Sp nov	Valid	Sassaroli & Venturi	Early Toarcian	Bugarone Formation	Italy	A hildoceratid, a species of Hildaites.
Hildaites calamantis	Sp nov	Valid	Sassaroli & Venturi	Early Toarcian	Bugarone Formation	Italy	A hildoceratid, a species of Hildaites.
Hildaites evolutus	Sp nov	Valid	Sassaroli & Venturi	Early Toarcian	Bugarone Formation	Italy	A hildoceratid, a species of Hildaites.
Hildaites planiventris	Sp nov	Valid	Sassaroli & Venturi	Early Toarcian	Bugarone Formation	Italy	A hildoceratid, a species of Hildaites.
Indosphinctes tsatiaensis	Sp. nov	Valid	Hall & Poulton	Callovian		Canada	A perisphinctid ammonite, a species of Indosphinctes.
Infraparkinsonia pierae	Sp. nov	Valid	Pavia & Zunino	Late Bajocian		France	A species of Infraparkinsonia, a possible member of Stephanoceratidae.
Inyoites sedini	Sp nov	In press	Zakharov & Abnavi	Olenekian	Zhitkov Formation	Russia	An inyoitid, a species of Inyoites.
Kashmirites baidi	Sp nov	Valid	Brühwiler et al.	Early Triassic		Oman	A kashmiritid ammonite, a species of Kashmirites.
Lepteocaloceras	Gen. et 2 sp. nov	Valid	Dommergues	Hettangian		France	A psiloceratid ammonite. Genus contains the type species Lepteocaloceras evrardii and possibly also the species Lepteocaloceras ? strictum.
Leptosphinctes (Cleistosphinctes) umbilicatus	Sp nov	Valid	Galácz	Bajocian		Hungary	A species of Leptosphinctes.
Lucasites	Gen. et 2 sp. nov	Valid	Brühwiler et al.	Early Triassic		Oman	A prionitid ammonite. Genus contains two species: Lucasites involutus and L. evolutus.
Maximites nassichuki	Sp. nov	Valid	Work, Mason & Boardman	Pennsylvanian	Four Corners Formation	United States	A species of Maximites.
Mesohedenstroemia olgae	Sp nov	In press	Zakharov & Abnavi	Olenekian	Zhitkov Formation	Russia	A hedenstroemiid, a species of Mesohedenstroemia.
Nannolytoceras gibbosum	Sp nov	Valid	Galácz & Kassai	Bajocian		Hungary	A member of Lytoceratina, a species of Nannolytoceras.
Neograhamites primus	Sp. nov	Valid	Olivero	Upper Campanian	Snow Hill Island Formation	Antarctica	A kossmaticeratid ammonite, a species of Neograhamites.
Neokossmaticeras	Gen. et sp. nov	Valid	Olivero	Upper Campanian	Rabot Formation	Antarctica	A kossmaticeratid ammonite. The type species is Neokossmaticeras redondensis.
Notosilesitoides	Gen. et sp. nov.	Valid	Matsukawa et al.	Aptian to early Albian		Philippines	The type species is Notosilesitoides philippinensis.
Obataceras	Gen. et sp. nov	Valid	Shigeta, Futakami & Hoffmann	Late Albian		Japan	A gabbioceratine ammonite. The type species is Obataceras manjiense.
Okhototrachyceras	Gen. et comb. nov	Valid	Konstantinov	Early Carnian		Russia	A trachyceratid, a new genus for "Protrachyceras" seimkanense Bytschkov.
Omanites	Gen. et sp. nov	Valid	Brühwiler et al.	Early Triassic		Oman	A paranannitid ammonite. The type species is Omanites musjahensis.
Oxyarietites	Gen. et sp. nov	Valid	Dommergues & Goolaerts	Early Sinemurian		France	An arietitoid, a member of Phylloceratida. The type species is Oxyarietites boletzkyi.
Paranannites baudi	Gen. et sp. nov	Valid	Brühwiler et al.	Early Triassic		Oman	A paranannitid ammonite, a species of Paranannites.
Phaneroceras chesnuti	Sp. nov	Valid	Work, Mason & Boardman	Pennsylvanian	Four Corners Formation	United States	A species of Phaneroceras.
Phaulostephanus apertus	Sp nov	Valid	Galácz	Bajocian		Hungary	A species of Phaulostephanus.
Phaulostephanus exilis	Sp nov	Valid	Galácz	Bajocian		Hungary	A species of Phaulostephanus.
Procolumbites safraensis	Sp nov	Valid	Brühwiler et al.	Late Early Triassic		Oman	A columbitid ammonite, a species of Procolumbites.
Pseudaspidites planus	Sp nov	Valid	Brühwiler et al.	Early Triassic		Oman	A proptychitid ammonite, a species of Pseudaspidites.
Rohillites omanensis	Sp nov	Valid	Brühwiler et al.	Early Triassic		Oman	A flemingitid ammonite, a species of Rohillites.
Sabaudiella riverorum	Sp nov	Valid	Aguirre-Urreta & Rawson	Valanginian or early Barremian	Agrio Formation	Argentina	A species of Sabaudiella.
Safraites	Gen. et sp. nov	Valid	Brühwiler et al.	Early Triassic		Oman	A galfettitid ammonite. The type species is Safraites simplex.
Stephanosphinctes tuberculatus	Sp nov	Valid	Galácz	Bajocian		Hungary	A species of Stephanosphinctes.
Subbalhaeceras	Gen. et sp. nov	In press	Zakharov & Abnavi	Olenekian	Zhitkov Formation	Russia	A flemingitid. The type species is Subbalhaeceras shigetai.
Tanabeceras	Gen. et sp. nov	Valid	Shigeta, Futakami & Hoffmann	Late Albian		Japan	A gabbioceratine ammonite. The type species is Tanabeceras pombetsense.
Ussurijuvenites	Gen. et 2 sp. nov	Valid	Smyshlyaeva & Zakharov	Early Olenekian		Russia	A melagathiceratid. The type species is U. popovi. U.s artyomensis is a second species.

Other cephalopods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Bathmoceras taichoutense	Sp nov	Valid	Kröger & Lefebvre	Early Ordovician	Fezouata Formation	Morocco	A species of Bathmoceras.
Destombesiceras	Gen. et sp. nov	Valid	Kröger & Lefebvre	Early Ordovician	Fezouata Formation	Morocco	A discosorid cephalopod. The type species is Destombesiceras zagorense.
Germanoteuthis	Gen. et sp. nov	Valid	Schweigert & Fuchs	Ladinian		Germany	A coleoid cephalopod. The type species is Germanoteuthis donai.
Reitneriteuthis	Gen. et comb. nov	Valid	Schweigert & Fuchs	Norian		Germany	A new genus for "Loligosepia" neidernachensis Reitner, 1978.

Newly named gastropods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Ananias riccardii	Sp. nov	Valid	Pinilla	Early Permian		Argentina	An eotomariid gastropod, a species of Ananias.
Angulathilda	Gen. et comb. nov	In press	Gründel & Nützel	Bathonian to Albian		Germany	A mathildid, a new genus for "Carinathilda" calloviensis (Gründel, 1997) and a few other species of Mesozoic mathildids.
Anulifera chubutensis	Sp nov	In press	Ferrari	Late Pliensbachian to early Toarcian	Mulanguiñeu Formation	Argentina	A protorculid, a species of Anulifera.
Bulinus corici	Sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Bulinus.
Campanile zakhoense	Sp. nov	In press	Harzhauser, Hoşgör & Pacaud	Thanetian	Kolosh Formation	Iraq	A campanilid, a species of Campanile.
Cryptaulax redelii	Sp. nov	Valid	Ferrari	Pliensbachian or Toarcian		Argentina	A procerithiid caenogastropod, a species of Cryptaulax.
Emmericia roetzeli	sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Emmericia.
Eunerinea mendozana	Sp. nov	In press	Cataldo	Early Cretaceous	Agrio Formation	Argentina	A nerineoid gastropod, a species of Eunerinea.
Europrotomus	Gen. et comb. nov	Valid	Kronenberg & Harzhauser	Miocene		Europe	A strombid caenogastropod, a new genus for "Strombus" schroeckingeri. Strombus almerai (Crosse, 1885) may be a second species.
"Faunus" dominicii	Sp. nov	In press	Harzhauser, Hoşgör & Pacaud	Thanetian	Kolosh Formation	Iraq	A pachychilid; provisionally assigned to the genus Faunus, but probably actually belongs to a separate genus (to be described by Pacaud and Harzhauser).
Ferrissia crenellata	sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Ferrissia.
Gyraulus sachsenhoferi	sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Gyraulus.
Kaneconcha	Gen. et sp. nov	Valid	Kaim, Tucholke & Warén	Late Pliocene		Mid-Atlantic Ridge, Atlantic Ocean	A provannid gastropod. The type species is Kaneconcha knorri.
Lepsiella ukika	sp nov	In press	Gordillo & Nielsen	Pleistocene		Chile	A haustrine muricid, a species of Lepsiella.
Nematurella zuschini	sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Nematurella.
Odontohydrobia groisenbachensis	sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Odontohydrobia.
Odontohydrobia pompatica	Sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Odontohydrobia.
Odontohydrobia styriaca	Sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Odontohydrobia.
Pachymelania islamogluae	Sp. nov	In press	Harzhauser, Hoşgör & Pacaud	Thanetian	Kolosh Formation	Iraq	A thiarid, a species of Pachymelania.
Persististrombus pannonicus	Sp nov	In press	Harzhauser & Kronenberg	Middle Miocene	Tauchen Formation	Austria	A strombid, a species of Persististrombus.
Planorbis austroalpinus	Sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Planorbis.
Procerithium (Infacerithum) nodosum	Sp. nov	Valid	Ferrari	Pliensbachian or Toarcian		Argentina	A procerithiid caenogastropod, a species of Procerithium.
Procerithium (Rhabdocolpus) patagoniensis	Sp. nov	Valid	Ferrari	Pliensbachian or Toarcian		Argentina	A procerithiid caenogastropod, a species of Procerithium.
Provanna alexi	Sp nov	In press	Amano & Little	Lower Middle Miocene	Chikubetsu Formation	Japan	A species of Provanna.
Provanna hirokoae	Sp nov	In press	Amano & Little	Middle Miocene	Ogaya Formation	Japan	A species of Provanna.
Pseudoaluco mesopotamicus	Sp. nov	In press	Harzhauser, Hoşgör & Pacaud	Thanetian	Kolosh Formation	Iraq	A cerithiid, a species of Pseudoaluco.
Pseudomelania feruglioi	Sp nov	In press	Ferrari	Late Pliensbachian to early Toarcian	Mulanguiñeu Formation	Argentina	A pseudomelaniid, a species of Pseudomelania.
Pyrazopsis hexagonpyramidalis	Sp. nov	In press	Harzhauser, Hoşgör & Pacaud	Thanetian	Kolosh Formation	Iraq	A batillariid, a species of Pyrazopsis.
Romania fastigata	Species	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Romania.
"Serradonta" kimberleyae	Sp. nov	Valid	Saether et al.	Miocene		New Zealand	A pectinodontid gastropod, possibly a species of Serradonta.
Stagnicola reinholdkunzi	Sp nov	Valid	Harzhauser & Neubauer in Harzhauser et al.	Middle Miocene	Feistring Formation	Austria	A species of Stagnicola.
Varicipotamides	Nom. nov	In press	Pacaud & Harzhauser in Harzhauser, Hoşgör & Pacaud	Bartonian	Paris Basin	France	A replacement name for Exechestoma Cossmann (1889; preoccupied).

Other molluscs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Asoella campbellorum	Sp. nov	Valid	Damborenea in Damborenea & Manceñido	Late Triassic		Argentina	An asoellid bivalve, a species of Asoella.
Hannestheronia	Gen. et sp. nov	Valid	Vinther et al.	Devonian		South Africa	A multiplacophoran. The type species is Hannestheronia australis.
Oriocrassatella piauiensis	Sp. nov	Valid	Anelli et al.	Carboniferous (Bashkirian to Moscovian)	Piauí Formation	Brazil	A crassatellid bivalve, a species of Oriocrassatella.
Protobalanus spinicoronatus	Sp. nov	Valid	Vinther et al.	Devonian		United States	A multiplacophoran, a species of Protobalanus.
Pseudocatillus nevesskayae	Sp nov	Valid	Esu & Popov	Late Messinian		Italy	A lymnocardiine cockle, a species of Pseudocatillus.
Pseudolimea riccardii	Sp. nov	Valid	Damborenea in Damborenea & Manceñido	Late Triassic		Argentina	A limid bivalve, a species of Pseudolimea.

Echinoderms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Anatifopsis fillmorensis	Sp. nov	Valid	Sumrall et al.	Early Ordovician		United States	A mitrate, a species of Anatifopsis.
Anatifopsis ninemilensis	Sp. nov	Valid	Sumrall et al.	Early Ordovician	McKelligon Canyon Formation Ninemile Shale	United States	A mitrate, a species of Anatifopsis.
Andymetra	Gen. et sp. nov	Valid	Hess	Early Bathonian		France	A comatulid crinoid. The type species is Andymetra galei.
Aorocrinus meyeri	Sp. nov	Valid	Ausich & Roeser	Late Kinderhookian	Cuyahoga Formation	United States	A camerate crinoid, a species of Aorocrinus.
Aphelecrinus gracilis	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Aphelecrinus.
Aryballocrinus martini	Sp. nov	Valid	Ausich & Roeser	Late Kinderhookian	Cuyahoga Formation	United States	A camerate crinoid, a species of Aryballocrinus.
Atelestocrinus meszarosi	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Atelestocrinus.
Bathyovulaster	Gen. et sp. nov	In press	Smith	Santonian		Italy	A spatangoid. The type species is Bathyovulaster disjunctus.
Brissopsis (Kleinia) riccardii	Sp. nov	Valid	Parma	Eocene to Oligocene or early Miocene		Argentina	A brissopsid echinoid, a species of Brissopsis.
Ctenoimbricata	Gen. et sp. nov	Valid	Zamora, Rahman & Smith	Cambrian		Spain	A stem group echinoderm. The type species is Ctenoimbricata spinosa.
Cuyahogacrinus	Gen. et sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid. The type species is Cuyahogacrinus lodiensis.
Cyathocrinites simplex	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Cyathocrinites.
Cyrtocrinus praenutans	Sp. nov	Valid	Hess	Early Bathonian		France	A cyrtocrinid crinoid, a species of Cyrtocrinus.
Diploblastus fadigai	Sp. nov	Valid	Atwood & Sumrall	Carboniferous (Mississippian)	Glen Dean Formation	United States	A blastoid, a species of Diploblastus.
Drepanocystis	Gen. et sp. nov	Valid	Sumrall et al.	Early Ordovician	Wah Wah Limestone	USA	A possible solute. The type species is Drepanocystis dubius.
Goniocrinus sceletus	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Goniocrinus.
Heckerocrinus	Nom nov.	In press	Doweld	Ordovician			A replacement name for the crinoid genus Bockia Hecker 1940.
Lebetocrinus ohioensis	Sp. nov	Valid	Kammer & Roeser	Kinderhookian	Cuyahoga Formation	United States	A cladid crinoid, a species of Lebetocrinus.
Palaeocomaster messingi	Sp. nov	Valid	Hess	Early Bathonian		France	A comatulid crinoid, a species of Palaeocomaster.
Pentacrinites ausichi	Sp. nov	Valid	Hess	Late Bajocian or early Bathonian		France	A species of Pentacrinites.
Pentremites fredericki	Sp. nov	Valid	Atwood & Sumrall	Carboniferous (Mississippian)	Glen Dean Formation	United States	A blastoid, a species of Pentremites.
Pentremites meganae	Sp. nov	Valid	Atwood & Sumrall	Carboniferous (Mississippian)	Glen Dean Formation	United States	A blastoid, a species of Pentremites.
Phyllocrinus voultensis	Sp. nov	Valid	Hess	Early Bathonian		France	A cyrtocrinid crinoid, a species of Phyllocrinus.
Phymosoma ravni	Sp nov.	Valid	Schlüter et al.	Late Cretaceous		Western Europe and Middle East	A phymosomatoid sea urchin, a species of Phymosoma.
Platycrinites burkei	Sp. nov	Valid	Ausich & Roeser	Late Kinderhookian	Cuyahoga Formation	United States	A camerate crinoid, a species of Platycrinites sensu lato.
Praetetracrinus bathonicus	Sp. nov	Valid	Hess	Early Bathonian		France	A cyrtocrinid crinoid, a species of Praetetracrinus.
Pygolampas	Gen. et sp. nov	Valid	Saucède, Dudicourt & Courville	Early Cretaceous (Early Hauterivian)		France	A sea urchin. The type species is Pygolampas edita.
Rhenopyrgus piojoensis	Sp nov	In press	Sumrall, Heredia, Rodríguez & Mestre	Silurian (lower Ludlow)	Los Espejos Formation	Argentina	A rhenopyrgid edrioasteroid, a species of Rhenopyrgus.
Salvaster	Gen. et sp. nov	Valid	Saucède, Dudicourt & Courville	Early Cretaceous (Early Hauterivian)		France	A sea urchin. The type species is Salvaster roberti.
Scutellacrinus	Gen. et sp. nov	Valid	Hess	Late Bajocian or early Bathonian		France	A cyrtocrinid crinoid. The type species is Scutellacrinus tenuis.
Singillatimetra	Gen. et sp. nov	Valid	Hess	Early Bathonian		France	A comatulid crinoid. The type species is Singillatimetra inordinata.
Solanocrinites voultensis	Sp. nov	Valid	Hess	Early Bathonian		France	A comatulid crinoid, a species of Solanocrinites.
Thylechinus (Thylechinus) sinaiensis	Sp. nov	Valid	Abdelhamid & Azab	Turonian		Egypt	A phymosomatid echinoid, a species of Thylechinus.
Toxaster dakhlensis	Sp. nov	Valid	Abdelhamid & Azab	Turonian		Egypt	A toxasterid echinoid, a species of Thylechinus.
Yanjiahella	Gen. et 3 sp. nov	Valid	Guo et al.	Early Cambrian	Yanjiahe Formation	China	An animal of uncertain phylogenetic position, a possible relative of echinoderms. Genus contains Yanjiahella ancarpa (the type species), Y. monocarpa and Y. biscarpa.

Ascidians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Burykhia	Gen. et sp.	Valid	Fedonkin et al.	Vendian	Verkhovka Formation or Ust-Pinega Formation	Russia	An ausiid, a possible ascidian, Its type species is Burykhia hunti.

Conodonts

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acanthodus humachensis	Sp nov	In press	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Acanthodus.
Acanthodus raqueli	Sp nov	In press	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Acanthodus.
Acodus primitivus	Sp nov	In press	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Acodus.
Carnepigondolella angulata	Sp nov	Valid	Mazza, Cau & Rigo	Late Triassic		Italy	A species of Carnepigondolella.
Carnepigondolella gulloae	Sp nov	Valid	Mazza, Rigo & Gullo	Late Triassic		Italy	A species of Carnepigondolella.
Carnepigondolella tuvalica	Sp nov	Valid	Mazza, Rigo & Gullo	Late Triassic		Italy	A species of Carnepigondolella.
Caudicriodus schoenlaubi	Sp nov	In press	Drygant & Szaniawski	Early Devonian (middle to late Lochkovian)	Chortkiv Formation	Ukraine	An icriodontid, a species of Caudicriodus.
Epigondolella heinzi	Sp nov	Valid	Mazza, Cau & Rigo	Late Triassic		Italy	A species of Epigondolella.
Epigondolella miettoi	Sp nov	Valid	Mazza, Cau & Rigo	Late Triassic		Italy	A species of Epigondolella.
Kallidontus gondwanicus	Sp nov	In press	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Kallidontus.
Nealeodus	Gen. et comb. nov	Valid	Stouge	Middle Ordovician		Canada	A new genus for "Lenodus" martinpointensis (Johnston and Barnes, 2000).
Norigondolella trinacriae	Sp nov	Valid	Mazza, Cau & Rigo	Late Triassic		Italy	A species of Norigondolella.
Pandorinellina? parva	Sp nov	In press	Drygant & Szaniawski	Early Devonian (late Lochkovian or Pragian)	Ivanye Formation	Ukraine	A spathognathodontid, possibly a species of Pandorinellina.
Periodon hankensis	Sp nov	Valid	Stouge	Middle Ordovician		Canada	A species of Periodon.
Spinodus wardi	Sp nov	Valid	Stouge	Middle Ordovician		Canada	A species of Spinodus.
Tilcarodus	Gen. et comb. nov	In press	Zeballo & Albanesi	Early Ordovician	Rupasca Formation	Argentina	A new genus for "Utahconus" humahuacensis Albanesi and Aceñolaza (2005).
Utahconus purmamarcensis	Sp nov	In press	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Utahconus.
Utahconus scandodiformis	Sp nov	In press	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Utahconus.
Utahconus tortibasis	Sp nov	In press	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Utahconus.
Variabiloconus crassus	Sp nov	In press	Zeballo & Albanesi	Late Cambrian (late Furongian) or early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Variabiloconus.

Fishes

Newly named jawless vertebrates

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Dunyu	Gen. et sp. nov	Valid	Zhu, Liu, Jia & Gai	Silurian	Kuanti Formation	China	A eugaleaspidiform galeaspid. Its type species is Dunyu longiforus.
Kerreralepis	Gen. et sp. nov	Valid	Blom	Lower Devonian		United Kingdom	A birkeniid anaspid. The type species is Kerreralepis carinata.

Newly named acanthodians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Halimacanthodes	Gen. et sp. nov	Valid	Burrow, Trinajstic & Long	Late Devonian (Frasnian)	Gogo Formation	Australia	An acanthodiform acanthodian. The type species is Halimacanthodes ahlbergi.
Podoliacanthus	Gen. et sp. nov	In press	Voichyshyn & Szaniawski	Early Devonian, possibly also late Silurian.		Ukraine Greenland	An ischnacanthiform, possibly an ischnacanthid. The type species is Podoliacanthus zychi; also contains three yet unnamed species.

Newly named cartilaginous fishes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Abdounia lata	Sp. nov	Valid	Malyshkina	Late Eocene	Tavda Formation	Russia	A requiem shark, a species of Abdounia.
Abdounia vassilyevae	Sp. nov	Valid	Malyshkina	Late Eocene	Tavda Formation	Russia	A requiem shark, a species of Abdounia.
Agaleorhynchus	Gen. et sp. nov	Valid	Guinot et al.	Late Cretaceous		United Kingdom	A batoid. The type species is Agaleorhynchus britannicus.
Argoubia	Gen. et sp. nov	Valid	Adnet et al.	Late Eocene		Morocco	A mobulid. The type species is Argoubia barbei.
Cretaplatyrhinoidis	Gen. et sp. nov	Valid	Guinot et al.	Late Cretaceous			A batoid. The type species is Cretaplatyrhinoidis ornatus.
Duffinselache	Gen. et comb. nov	Valid	Andreev & Cuny	Rhaetian		United Kingdom	A selachimorph, a new genus for "Polyacrodus" holwellensis (Duffin, 1998).
Eoplinthicus underwoodi	Sp. nov	Valid	Adnet et al.	Late Eocene		Morocco	A mobulid, a species of Eoplinthicus.
Eosqualiolus skrovinai	Sp nov	In press	Underwood & Schlogl	Miocene	Lakšárska Nová Ves Formation	Slovakia	A dalatiid, a species of Eosqualiolus.
Isacrodus	Gen. et sp. nov	Valid	Ivanov, Nestell & Nestell	Middle Permian (Roadian)	Cutoff Formation	USA	A jalodontid chondrichthyan. The type species is Isacrodus marthae.
Kaibabvenator	Gen. et sp. nov	Valid	Hodnett et al.	Permian (Late Leonardian)	Kaibab Formation	USA	A ctenacanthiform. The type species is Kaibabvenator swiftae.
Nanocetorhinus	Gen. et sp. nov	In press	Underwood & Schlogl	Miocene	Lakšárska Nová Ves Formation	Slovakia	A neoselachian of uncertain affinities. The type species is Nanocetorhinus tuberculatus.
Nanoskalme	Gen. et sp. nov	Valid	Hodnett et al.	Permian (Late Leonardian)	Kaibab Formation	USA	A ctenacanthiform. The type species is Nanoskalme natans.
Neosaivodus	Gen. et sp. nov	Valid	Hodnett et al.	Permian (Late Leonardian)	Kaibab Formation	USA	A ctenacanthiform. The type species is Neosaivodus flagstaffensis.
Oromobula	Gen. et sp. nov	Valid	Adnet et al.	Late Eocene		Morocco	A mobulid. The type species is Oromobula dakhlaensis.
Paraetmopterus horvathi	Sp nov	In press	Underwood & Schlogl	Miocene	Lakšárska Nová Ves Formation	Slovakia	An etmopterid, a species of Paraetmopterus.
Parasquatina jarvisi	Sp nov	Valid	Guinot et al.	Late Cretaceous			A squatiniform, a species of Parasquatina.
Parasquatina justinensis	Sp nov	Valid	Guinot et al.	Late Cretaceous			A squatiniform, a species of Parasquatina.
Pristiophorus striatus	Species	In press	Underwood & Schlogl	Miocene	Lakšárska Nová Ves Formation	Slovakia	A sawshark, a species of Pristiophorus
Pseudoplatyrhina	Gen. et sp. nov	Valid	Guinot et al.	Late Cretaceous			A batoid. The type species is Pseudoplatyrhina crispa.
Ptychotrygonoides sabatieri	Sp. nov	Valid	Guinot et al.	Late Cretaceous			A batoid, a species of Ptychotrygonoides.
"Rhinobatos" seruensis	Sp. nov	Valid	Guinot et al.	Late Cretaceous			A batoid, a possible species of Rhinobatos.
Rhomaleodus	Gen. et sp. nov	Valid	Andreev & Cuny	Anisian		Bulgaria	A selachimorph. The type species is Rhomaleodus budurovi.
Scyliorhinotheca	Gen. et sp. nov	In press	Kiel, Peckmann & Simon	Late Eocene		USA	Catshark egg capsules. The type species is Scyliorhinotheca goederti.
Squatirhina draytoni	Sp. nov	Valid	Guinot et al.	Late Cretaceous			A batoid, a species of Squatirhina.
Tantalepis	Gen. et sp. nov	Valid	Sansom et al.	Middle Ordovician (Darriwilian)	Stairway Sandstone	Australia	A gnathostome of uncertain phylogenetic placement, a possible cartilaginous fish. The type species is Tantalepis gatehousei.
Texasodus	Gen. et sp. nov	Valid	Ivanov, Nestell & Nestell	Middle Permian (late Wordian or early Capitanian)	Bell Canyon Formation	USA	A jalodontid chondrichthyan. The type species is Texasodus varidentatus.

Newly named bony fishes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aglyptorhynchus hakataramea	sp nov	Valid	Gottfried, Fordyce & Rust	Late Oligocene		New Zealand	A billfish
Apsopelix miyazakii	Sp. nov	Valid	Yabumoto, Hikida & Nishino	Late Cretaceous (Turonian)		Japan	A species of Apsopelix.
Aspistor verumquadriscutis	sp nov	Valid	Aguilera & Marceniuk	Late Miocene	Urumaco Formation	Venezuela	An ariid, a species of Aspistor.
Bawitius	Gen. et comb. nov	Valid	Grandstaff et al..	Cenomanian	Bahariya Formation	Egypt	A polypterid actinopterygian, a new genus for "Polypterus" bartheli (Schaal, 1984).
Bruehnopteron	Gen. et sp. nov	Valid	Schultze & Reed	Middle Devonian		United States	A tristichopterid sarcopterygian. The type species is Bruehnopteron murphyi.
Caruso	Gen. et comb. nov	Valid	Carnevale & Pietsch	Eocene		Italy	A lophiid anglerfish, a new genus for "Lophius" brachysomus (Agassiz).
Echinelops	Gen. et sp. nov.	Valid	Murray & Hoşgör	Early Oligocene		Turkey	An elopiform fish. The type species is Echinelops ozcani.
Galaxias angustiventris	Sp nov	Valid	Schwarzhans in Schwarzhans et al..	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias brevicauda	Sp nov	Valid	Schwarzhans in Schwarzhans et al..	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias bobmcdowalli	Sp nov	Valid	Schwarzhans in Schwarzhans et al..	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias papilionis	Sp nov	Valid	Schwarzhans in Schwarzhans et al..	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias parvirostris	Sp nov	Valid	Schwarzhans in Schwarzhans et al..	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias tabidus	Sp nov	Valid	Schwarzhans in Schwarzhans et al..	Early Miocene		New Zealand	A galaxiid, a species of Galaxias.
Gerpegezhus	Gen. et sp. nov	Valid	Bannikov & Carnevale	Early Eocene		Russia	A close relative of centriscids. The type species is Gerpegezhus paviai.
Hippotropiscis	Gen. et sp. nov	Valid	Žalohar & Hitij	Middle Miocene		Slovenia	A hippocampine syngnathid. The type species is Hippotropiscis frenki.
Illusionella	Gen. et 2 sp. nov	Valid	Baykina	Miocene		Russia	An clupeid fish. Genus contains two species: Illusionella tsurevica and Illusionella pshekhensis.
Isanichthys lertboosi	Sp. nov	In press	Deesri et al.	Probably Late Jurassic	Phu Kradung Formation	Thailand	A lepisosteiform, a species of Isanichthys.
Isurichthys breviusculus	Sp nov	Valid	Bannikov	Lower Oligocene		Russia	An ariommatid, a species of Isurichthys.
Kyphosichthys	Gen. et sp. nov	Valid	Xu & Wu	Middle Anisian	Guanling Formation	China	A ginglymodian actinopterygian.
Luopingcoelacanthus	Gen. et sp. nov	In press	Wen et al..	Anisian	Guanling Formation	China	A coelacanth. The type species is Luopingcoelacanthus eurylacrimalis.
Luoxiongichthys	Gen. et sp. nov	Valid	Wen et al.	Middle Anisian	Guanling Formation	China	A basal actinopterygian.
Macrosemimimus	Gen. et sp. et comb. nov	Valid	Schröder, López-Arbarello & Ebert	Late Jurassic		France Germany United Kingdom	A semionotiform. The type species is Macrosemimimus fegerti; "Lepidotes" lennieri (Sauvage, 1893) is a second species of Macrosemimimus.
Mataichthys	Gen. et sp. nov	Valid	Schwarzhans, Scofield, Tennyson & T. Worthy in Schwarzhans et al..	Early Miocene	Bannockburn Formation	New Zealand	An eleotrid. Contains M. bictenatus, M. procerus, M. rhinoceros, and M. taurinus
Nanaichthys	Gen. et sp. nov	Valid	Amaral & Brito	Early Cretaceous, possibly Aptian	Marizal Formation	Brazil	A member of Chanidae. The type species is Nanaichthys longipinnus.
Palatinichthys	Gen. et sp. nov	Valid	Witzmann & Schoch	Lower Permian	Saar–Nahe Basin	Germany	A megalichthyid sarcopterygian. The type species is Palatinichthys laticeps.
Paleotetra	Gen. et 2 sp.	Valid	Weiss, Malabarba & Malabarba	Eocene to Oligocene		Brazil	A characiform, probably a characid. The genus contains two species: Paleotetra entrecorregos and Paleotetra aiuruoca.
Paranaichthys	Gen. et sp. nov	Valid	Dias	Permian, Guadalupian (Wordian or Capitanian)	Rio do Rasto Formation	Brazil	A probable relative of Platysomus and other platysomids. The type species is Paranaichthys longianalis.
Parawenzichthys	Gen. et sp. nov	Valid	de Figueiredo, Gallo & Delarmelina	Turonian	Atlântida Formation	Brazil	A protacanthopterygian.
Potanichthys	Gen. et sp. nov	In press	Xu et al.	Middle Triassic (Ladinian)	Falang Formation	China	A relative of Thoracopterus capable of over-water gliding. The type species is Potanichthys xingyiensis.
Prototroctes modestus	Sp nov	Valid	Schwarzhans in Schwarzhans et al..	Early Miocene	Bannockburn Formation	New Zealand	A retropinnid, a species of Prototroctes.
Prototroctes vertex	Sp nov	Valid	Schwarzhans in Schwarzhans et al..	Early Miocene	Bannockburn Formation	New Zealand	A retropinnid, a species of Prototroctes.
Rebellatrix	Gen. et sp. nov	Valid	Wendruff & Wilson	Lower Triassic	Sulphur Mountain Formation	Canada	A coelacanth. The type species is Rebellatrix divaricerca.
Reidus	Gen. et sp. nov	Valid	Graf	Albian		United States	A coelacanth. The type species is Reidus hilli.
Rhinodipterus kimberleyensis	Sp nov	Valid	Clement	Late Devonian	Gogo Formation	Australia	A lungfish, a species of Rhinodipterus.
Tinirau	Gen. et sp. nov	Valid	Swartz	Middle Devonian	Red Hill I beds	United States	An eotetrapodiform tetrapodomorph. The type species is Tinirau clackae.
Tomognathus gigeri	sp nov	Valid	Cavin & Giner	Valanginian		France	A halecomorph fish, a species of Tomognathus.
Tungsenia	Gen. et sp. nov	Valid	Lu et al.	Early Devonian		China	A basal member of Tetrapodomorpha. The type species is Tungsenia paradoxa.
Voelklichthys	Gen. et sp. nov	Valid	Arratia & Schultze	Kimmeridgian		Germany	A macrosemiiform. The type species is Voelklichthys comitatus.
Yunnancoelacanthus	Gen. et sp. nov	In press	Wen et al..	Anisian	Guanling Formation	China	A coelacanth. The type species is Yunnancoelacanthus acrotuberculatus.

Amphibians

Newly named basal tetrapods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Ymeria	Gen. et sp. nov	Valid	Clack, Ahlberg, Blom & Finney	Famennian	Celsius Bjerg Group	Greenland	A stem tetrapod closely related to Ichthyostega.

Newly named temnospondyls

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Arachana	Gen. et sp. nov	Valid	Piñeiro, Ramos & Marsicano	? Late Permian	Buena Vista Formation	Uruguay	A rhinesuchid-like stereospondyl.
Benthosuchus gusevae	Sp. nov	Valid	Novikov	Early Triassic	Kamennyi Yar Formation	Russia	A trematosauroid.
Cacops woehri	Sp. nov	Valid	Fröbisch & Reisz	Leonardian		USA	A dissorophid.
Microposaurus averyi	Sp nov	Valid	Warren	Anisian	Rouse Hill Siltstone	Australia	A trematosaurine stereospondyl, a species of Microposaurus.	Microposaurus averyi.
Parotosuchus ptaszynskii	Sp nov	In press	Sulej & Niedźwiedzki	Late Olenekian		Poland	A species of Parotosuchus.
Qantas	Gen. et sp. nov	Valid	Novikov	Early Triassic	Kamennyi Yar Formation	Russia	A trematosauroid temnospondyl. The type species is Qantas samarensis.
Sclerocephalus stambergi	Sp nov	Valid	Klembara & Steyer	Early Permian	Boskovice Basin	Czech Republic	A stereospondylomorph, a species of Sclerocephalus.
Trematosuchoides	Gen. et sp. nov	Valid	Novikov	Triassic	Cynognathus Assemblage Zone	South Africa	A trematosaurid temnospondyl. The type species is Trematosuchoides africanus.

Newly named lepospondyls

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Altenglanerpeton	Gen. et sp. nov	Valid	Glienke	Gzhelian or Asselian	Altenglan Formation	Germany	A microsaur with an elongated body.

Newly named lissamphibians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Bakonybatrachus	Gen. et sp. nov	Valid	Szentesi & Venczel	Santonian	Csehbánya Formation	Hungary	A discoglossine discoglossid.
Beiyanerpeton	Gen. et sp. nov	Valid	Gao & Shubin	Oxfordian	Tiaojishan Formation	China	A salamandroid. The type species is Beiyanerpeton jianpingensis.
Eobarbourula	Gen. et sp. nov	In press	Folie et al..	Early Eocene	Cambay Shale Formation	India	A bombinatorid. Its type species is Eobarbourula delfinoi.
Indorana	Gen. et sp. nov	In press	Folie et al..	Early Eocene	Cambay Shale Formation	India	A possible rhacophorid. Its type species is Indorana prasadi.
Seminobatrachus	Gen. et sp. nov	Valid	Skutschas & Gubin	late Paleocene - early Eocene		Ukraine	A urodelan salamander
Uberabatrachus	Gen. et sp. nov	Valid	Báez et al.	Maastrichtian	Marília Formation	Brazil	A neobatrachian, a possible member of Nobleobatrachia. The type species is Uberabatrachus carvalhoi.
Wesserpeton	Gen. et sp. nov	In press	Sweetman & Gardner	Barremian	Wessex Formation	England	An albanerpetontid.

Anapsids

Newly named anapsids

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Feeserpeton	Gen. et sp. nov.	Valid	Macdougall & Reisz	Early Permian		United States	A lanthanosuchoid. The type species is Feeserpeton oklahomensis.
Lasasaurus	Gen. et sp. nov.	Valid	Falconnet et al.	Lower Triassic	Middle Sakamena Formation	Madagascar	A procolophonid. The type species is Lasasaurus beltanae.

Newly named turtles

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Atoposemys	Gen. et sp. nov	Valid	Hutchison	Early Paleocene (Puercan)	Fort Union Formation	United States	A trionychid. The type species is Atoposemys entopteros.
Axestemys cerevisia	sp nov	Valid	Vitek	Eocene	Bridger Formation	United States	A trionychid, a species of Axestemys.
Axestemys montinsana	sp nov	Valid	Vitek	Paleocene	Denver Formation Fort Union Formation Melville Formation	United States	A trionychid, a species of Axestemys.
Bairdemys healeyorum	Sp. nov	Valid	Weems & Knight	Late Oligocene	Chandler Bridge Formation	United States	A podocnemidid pleurodiran, a species of Bairdemys.
Bashuchelys	Gen. et comb. et sp. nov	Valid	Tong, Danilov, Ye, Ouyang & Peng	Middle Jurassic	Xiashaximiao Formation	China	A bashuchelyid cryptodiran, a new genus for "Chengyuchelys" zigongensis (Ye, 1982). Genus also contains a new species Bashuchelys youngi.
Basilemys gaffneyi	Sp. nov	Valid	Sullivan, Jasinski & Lucas	Late Campanian		United States	A nanhsiungchelyid turtle, a species of Basilemys.
Berruchelus	Gen. et sp. nov	Valid	Pérez-García	Upper Paleocene		Western Europe	A paracryptodiran closely related to Compsemys. The type species is Berruchelus russelli.
Brachyopsemys	Gen. et sp. nov	Valid	Tong & Meylan	Early Paleocene	Ouled Abdoun Basin	Morocco	A cryptodiran closely related to Sandownia. The type species is Brachyopsemys tingitana.
Brodiechelys royoi	Sp. nov	In press	Pérez-García, Gasulla & Ortega	Early Aptian	Arcillas de Morella Formation	Spain	A xinjiangchelyid, a species of Brodiechelys.
Carbonemys	Gen. et sp. nov	Valid	Cadena et al.	Late Palaeocene	Cerrejón Formation	Colombia	A podocnemidid turtle. The type species is Carbonemys cofrinii.
Cardichelyon	Gen. et sp. nov	Valid	Hutchison	Early Eocene (Wasatchian)		United States	A platysternid. The type species is Cardichelyon rogerwoodi.
Chuannanchelys	Gen. et comb. nov	Valid	Tong, Danilov, Ye, Ouyang & Peng	Middle Jurassic	Xiashaximiao Formation	China	A bashuchelyid cryptodiran, a new genus for "Chengyuchelys" dashanpuensis (Fang, 1987).
Foxemys trabanti	sp nov	Valid	Rabi, Tong & Botfalvai	Santonian	Csehbánya Formation	Hungary	A bothremydid, a species of Foxemys.
Galvechelone	Gen. et sp. nov	Valid	Pérez-García & Murelaga	Early Cretaceous		Spain	A cryptodiran turtle. The type species is Galvechelone lopezmartinezae.
Hoyasemys	Gen. et sp. nov	Valid	Pérez-García, Fuente & Ortega	Late Barremian	Calizas de La Huérguina Formation	Spain	A basal eucryptodiran.
Iberoccitanemys	Gen. et comb. nov	Valid	Pérez-García, Ortega & Murelaga	Late Campanian to Maastrichtian		France Spain	A bothremydid, a new genus for "Elochelys" convenarum (Laurent et al. 2002).
Kappachelys	Gen. et sp. nov	Valid	Hirayama, Isaji & Hibino	Early Cretaceous	Akaiwa Formation	Japan	A relative of trionychids. The type species is Kappachelys okurai.
Kinosternon pojoaque	Species	Valid	Bourque	Middle Miocene		United States	A species of Kinosternon.
Kinosternon skullridgescens	Species	Valid	Bourque	Early middle Miocene	Tesuque Formation	United States	A species of Kinosternon.
Laganemys	Gen. et sp. nov	Valid	Sereno & ElShafie	Aptian or Albian	Elrhaz Formation	Niger	An araripemydid pleurodiran. The type species is Laganemys tenerensis.
Neurankylus lithographicus	Sp. nov	Valid	Larson et al.	Santonian	Milk River Formation	Canada	A neurankyline baenid, a species of Neurankylus.
Nostimochelone	Gen. et sp. nov	Valid	Georgalis et al.	Early Miocene	Zeugostasion Formation	Greece	A pleurodiran, a member of Podocnemidoidea. The type species is Nostimochelone lampra.
Planetochelys	Gen. et sp. nov	Valid	Hutchison	Early Eocene (Wasatchian)		United States	A relative of trionychids. The type species is Planetochelys dithyros.
Polysternon isonae	Species	Valid	Marmi et al..	Late Maastrichtian		Spain	A bothremydid, a species of Polysternon.
Protoxinjiangchelys	Gen. et sp. nov	Valid	Tong, Danilov, Ye, Ouyang & Peng	Middle Jurassic	Xiashaximiao Formation	China	A xinjiangchelyid cryptodiran.
Psilosemys	Gen. et sp. nov	Valid	Hutchison	Early Eocene (Wasatchian)		United States	The earliest North American emydid. The type species is Psilosemys wyomingensis.
Puentemys	Gen. et sp. nov.	Valid	Cadena, Bloch & Jaramillo	Paleocene	Cerrejón Formation	Colombia	A bothremydid turtle. The type species is Puentemys mushaisaensis.
Rhinoclemmys panamaensis	Sp nov	Valid	Cadena et al.	Miocene	Cucaracha Formation	Panama	A geoemydid turtle, a species of Rhinoclemmys.
Scabremys	Gen. et comb. nov	Valid	Sullivan, Jasinski & Lucas	Late Campanian		United States	A baenid cryptodiran, a new genus for "Baena" ornata Gilmore (1935).
Spoochelys	Gen. et sp. nov	Valid	Smith & Kear	Early or middle Albian	Griman Creek Formation	Australia	A meiolaniid-like turtle. The type species is Spoochelys ormondea.
Staurotypus moschus	Sp nov	Valid	Cadena et al.	Miocene	Cucaracha Formation	Panama	A kinosternid turtle, a species of Staurotypus.
Terrapene parornata	Sp nov	Valid	Joyce et al..	Miocene/Pliocene boundary		United States	A box turtle.
Tullochelys	Gen. et sp. nov	Valid	Hutchison	Maastrichtian to early Paleocene (Puercan)	Fort Union Formation Hell Creek Formation	United States	A chelydrid. The type species is Tullochelys montanus.
Xinjiangchelys radiplicatoides	Sp. nov	Valid	Brinkman et al.	Late Middle or early Late Jurassic	Shishugou Formation	China	A species of Xinjiangchelys.
Yuchelys	Gen. et sp. nov	In press	Tong et al.	Late Cretaceous	Gaogou Formation	China	A nanhsiungchelyid cryptodiran. The type species is Yuchelys nanyangensis.

Diapsids

Newly named ichthyosaurs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acamptonectes	Gen. et sp. nov	Valid	Fischer et al..	Hauterivian–Cenomanian	Cambridge Greensand Formation	United Kingdom	An ophthalmosaurid. Type species is Acamptonectes densus.	Skull roof of Acamptonectes densus.
Cryopterygius	Gen. et sp. nov	Valid	Druckenmiller et al.	Late Jurassic	Agardhfjellet Formation	Norway	An ophthalmosaurid. The type species is Cryopterygius kristiansenae.
Palvennia	Gen. et sp. nov	Valid	Druckenmiller et al.	Late Jurassic	Agardhfjellet Formation	Norway	An ophthalmosaurid. The type species is Palvennia hoybergeti.
Stenopterygius aaleniensis	Sp. nov	Valid	Maxwell, Fernández & Schoch	Aalenian		Germany	A species of Stenopterygius.	The holotype of Stenopterygius aaleniensis.
Temnodontosaurus azerguensis	Sp. nov	Valid	Martin et al.	Middle Toarcian		France	A species of Temnodontosaurus.

Lepidosauromorphs

Newly named sauropterygians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Albertonectes	Gen. et sp. nov	Valid	Kubo, Mitchell & Henderson	Upper Campanian	Bearpaw Formation	Canada	An elasmosaurid. The type species is Albertonectes vanderveldei.	Albertonectes vanderveldei.
Anningasaura	Gen. et sp. nov.	Valid	Vincent & Benson	Early Jurassic		United Kingdom	A basal member of Plesiosauria. The type species is Anningasaura lymense.
Avalonnectes	Gen. et sp. nov	Valid	Benson, Evans & Druckenmiller	Early Jurassic, most likely earliest Hettangian.	Blue Lias Formation	United Kingdom	A rhomaleosaurid. The type species is Avalonnectes arturi.	Avalonnectes.
Cryonectes	Gen. et sp. nov	In press	Vincent, Bardet & Mattioli	Pliensbachian		France	A pliosaurid. The type species is Cryonectes neustriacus
Djupedalia	Gen. et sp. nov.	Valid	Knutsen, Druckenmiller & Hurum	Late Jurassic	Agardhfjellet Formation	Norway	A long-necked plesiosaurian. The type species is Djupedalia engeri.
Dolichorhynchops tropicensis	sp nov	Valid	McKean	Early Turonian	Tropic Shale	United States	Third species of Dolichorhynchops.
Eoplesiosaurus	Gen. et sp. nov	Valid	Benson, Evans & Druckenmiller	Early Jurassic, most likely earliest Hettangian.		United Kingdom	A basal plesiosauroid. The type species is Eoplesiosaurus antiquior.	Eoplesiosaurus.
Largocephalosaurus	Gen. et sp. nov	Valid	Cheng, Chen, Zeng & Cai	Middle Triassic	Guanling Formation	China	A relative of Wumengosaurus, pachypleurosaurs and nothosauroids.
Lusonectes	Gen. et sp. nov	Valid	Smith, Araújo & Mateus	Toarcian	São Gião Formation	Portugal	A plesiosaurid plesiosaur.
Pliosaurus funkei	Sp. nov	Valid	Knutsen, Druckenmiller & Hurum	Late Jurassic	Agardhfjellet Formation	Norway	A species of Pliosaurus.
Spitrasaurus	Gen. et 2 sp. nov.	Valid	Knutsen, Druckenmiller & Hurum	Late Jurassic	Agardhfjellet Formation	Norway	A long-necked plesiosaurian. Genus contains two species: Spitrasaurus wensaasi and S. larseni.
Stratesaurus	Gen. et sp.	Valid	Benson, Evans & Druckenmiller	Early Jurassic, most likely earliest Hettangian.	Blue Lias Formation	United Kingdom	A rhomaleosaurid. The type species is Stratesaurus taylori	Stratesaurus.

Newly named rhynchocephalians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Oenosaurus	Gen. et sp. nov	Valid	Rauhut et al.	Late Jurassic (early Tithonian)	Mörnsheim Formation	Germany	A sphenodontid rhynchocephalian. The type species is Oenosaurus muehlheimensis.
Sphenocondor	Gen. et sp. nov	Valid	Apesteguía, Gómez & Rougier	Middle Jurassic		Argentina	A rhynchocephalian lepidosaur. The type species is Sphenocondor gracilis.

Newly named lizards

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Chianghsia	Gen. et sp. nov	Valid	Mo, Xu & Evans	Late Cretaceous	Nanxiong Formation	China	A platynotan lizard, probably a monstersaurian. The type species is Chianghsia nankangensis.
Dornosaurus	Gen. et sp. nov	Valid	Alifanov	Eocene		Mongolia	An arretosaurid iguanian. The type species is Dornosaurus gobiensis.
Eremiasaurus	Gen. et sp. nov	Valid	Leblanc, Caldwell & Bardet	Maastrichtian		Morocco	A mosasaur.
Ergiliinsaurus	Gen. et sp. nov	Valid	Alifanov	Oligocene	Ergiliin Zoo Formation	Mongolia	An arretosaurid iguanian. The type species is Ergiliinsaurus postumus.
Jucaraseps	Gen. et sp. nov	Valid	Bolet & Evans	Early Cretaceous		Spain	A lizard related to scleroglossan lizards. The type species is Jucaraseps grandipes.
Khaichinguana	Gen. et sp. nov	Valid	Alifanov	Eocene	Khaychin Formation	Mongolia	An arretosaurid iguanian. The type species is Khaichinguana eocaenica.
Pseudopus ahnikoviensis	Sp. nov	Valid	Klembara	Early Miocene		Czech Republic	An anguid, a species of Pseudopus.
Uquiasaurus	Gen. et sp. nov	Valid	Daza et al.	Late Pliocene	Uquía Formation	Argentina	An iguanian lizard. The type species is Uquiasaurus heptanodonta.
Varanus (Varaneades) amnhophilis	Subgen. et sp. nov	Valid	Conrad, Balcarcel & Mehling	Miocene (Turolian)	Mytilini Formation	Greece	A monitor lizard.

Archosauromorphs

Newly named basal archosauromorphs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Chanaresuchus ischigualastensis	sp nov	Valid	Trotteyn, Martínez & Alcober	Late Triassic	Ischigualasto Formation	Argentina	A proterochampsian archosauriform, a species of Chanaresuchus.
Protome	Gen. et sp. nov	Valid	Stocker	Late Triassic	Chinle Formation	USA	A phytosaur. The type species is Protome batalaria.
Smok	Gen. et sp. nov	Valid	Niedźwiedzki, Sulej & Dzik	Latest Norian/Early Rhaetian	Lipie Śląskie clay-pit	Poland	A large predatory archosaur of uncertain phylogenetic placement, the largest known predatory archosaur from the Late Triassic and Early Jurassic of central Europe.

Newly named pseudosuchians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aegisuchus	Gen. et sp. nov	Valid	Holliday & Gardner	Cenomanian	Kem Kem Formation	Morocco	An aegyptosuchid eusuchian.	Aegisuchus witmeri.
Aetobarbakinoides	Gen. et sp. nov	Valid	Desojo, Ezcurra & Kischlat	Late Carnian or early Norian	Santa Maria Formation	Brazil	An aetosaur.
Barreirosuchus	Gen. et sp. nov	Valid	Iori & Garcia	Turonian to Santonian	Adamantina Formation	Brazil	A trematochampsid. The type species is Barreirosuchus franciscoi.
Borealosuchus threeensis	Sp. nov	Valid	Brochu, Parris, Grandstaff, Denton & Gallagher	Maastrichtian or Danian		USA	A crocodilian from New Jersey.
Bystrowisuchus	Gen. et sp. nov	Valid	Sennikov	Early Triassic		Russia	A ctenosauriscid. The type species is Bystrowisuchus flerovi.
Crocodylus thorbjarnarsoni	sp nov	Valid	Brochu & Storrs	Pliocene to Pleistocene		Kenya	A species of Crocodylus.	Crocodylus thorbjarnarsoni.
Diandongosuchus	Gen. et sp. nov.	Valid	Li et al.	Ladinian	Falang Formation	China	A poposauroid. The type species is Diandongosuchus fuyuanensis.	Diandongosuchus fuyuanensis.
Gasparinisuchus	Gen. et sp. nov	Valid	Martinelli et al.	Late Cretaceous		Argentina	A peirosaurid. The type species is Gasparinisuchus peirosauroides.

Newly named basal dinosauriforms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Diodorus	Gen. et sp. nov	Valid	Kammerer, Nesbitt, & Shubin	?Carnian-Norian	Timezgadiouine Formation	Morocco	A silesaurid.

Newly named non-avian dinosaurs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Alnashetri	Gen. et sp. nov	Valid	Makovicky, Apesteguía & Gianechini	Late Cretaceous (Cenomanian or Turonian)	Candeleros Formation	Argentina	A coelurosaurian, a probable relative of alvarezsaurids. The type species is Alnashetri cerropoliciensis.
Batyrosaurus	Gen. et sp. nov	Valid	Godefroit et al.	Late Cretaceous		Kazakhstan	A basal hadrosauroid. The type species is Batyrosaurus rozhdestvenskyi.
Bicentenaria	Gen. et sp. nov	Valid	Novas et al.	Cenomanian	Candeleros Formation	Argentina	A coelurosaur. The type species is Bicentenaria argentina.
Bonapartenykus	Gen. et sp. nov	Valid	Agnolin, Powell, Novas, & Kundrát	Campanian/Maastrichtian	Allen Formation	Argentina	An alvarezsaurid.
Camarillasaurus	Gen. et sp. nov	In press	Sánchez-Hernández & Benton	Early Barremian	Camarillas Formation	Spain	A basal ceratosaurian. The type species is Camarillasaurus cirugedae.
Elaltitan	Gen. et sp. nov	Valid	Mannion & Otero	Middle Cenomanian to Turonian	Bajo Barreal Formation	Argentina	A lithostrotian titanosaur. The type species is Elaltitan lilloi.
Eoabelisaurus	Gen. et sp. nov	Valid	Pol & Rauhut	Aalenian or Bajocian	Cañadon Asfalto Formation	Argentina	An abelisaurid. The type species is Eoabelisaurus mefi.
Gideonmantellia	Gen. et sp. nov	Valid	Ruiz-Omeñaca et al.	Barremian	Camarillas Formation	Spain	A basal ornithopod. The type species is Gideonmantellia amosanjuanae.
Gryphoceratops	Gen. et sp. nov	Valid	Ryan, Evans, Currie, Brown, & Brinkman	Late Santonian	Milk River Formation	Canada	A leptoceratopsid ceratopsian.
Hexing	Gen. et sp. nov	Valid	Jin, Chen & Godefroit	Early Valanginian to early Barremian	Yixian Formation	China	A basal ornithomimosaur. The type species is Hexing qingyi.
Huehuecanauhtlus	Gen. et sp.	Valid	Ramírez-Velasco et al..	Santonian		Mexico	A basal hadrosauroid. The type species is Huehuecanauhtlus tiquichensis.	Huehuecanauhtlus.
Ichthyovenator	Gen. et sp. nov	Valid	Allain et al.	Late Early Cretaceous	Savannakhet Basin	Laos	A spinosaurid. The type species is Ichthyovenator laosensis.
Juratyrant	Gen. et comb. nov	In press	Brusatte & Benson	Tithonian	Kimmeridge Clay	United Kingdom	A basal tyrannosauroid, a new genus for "Stokesosaurus" langhami (Benson, 2008).	Juratyrant.
Kundurosaurus	Gen. et sp. nov	Valid	Godefroit, Bolotsky & Lauters	Late Cretaceous, possibly late Maastrichtian	Udurchukan Formation	Russia	A saurolophine hadrosaurid. The type species is Kundurosaurus nagornyi.	Kundurosaurus.
Magnapaulia	Gen. et comb. nov	Valid	Prieto-Márquez, Chiappe & Joshi	Late Campanian	El Gallo Formation	Mexico	A lambeosaurine hadrosaurid, a new genus for "Lambeosaurus" laticaudus (Morris, 1981).	Magnapaulia.
Martharaptor	Gen. et sp. nov	Valid	Senter, Kirkland & DeBlieux	Early Aptian	Cedar Mountain Formation	United States	A possible therizinosauroid. The type species is Martharaptor greenriverensis.	Martharaptor.
Microraptor hanqingi	sp nov	Valid	Gong et al.	Early Cretaceous		China	A species of Microraptor.
Mochlodon vorosi	Sp. nov	Valid	Ősi et al.	Santonian	Csehbánya Formation	Hungary	A rhabdodontid ornithopod, a species of Mochlodon.
Ostafrikasaurus	Gen. et sp. nov	Valid	Buffetaut	Late Jurassic	Tendaguru Formation	Tanzania	A spinosaurid. The type species is Ostafrikasaurus crassiserratus.
Pachyrhinosaurus perotorum	sp nov	Valid	Fiorillo & Tykoski	Maastrichtian	Prince Creek Formation	USA	Third known species of Pachyrhinosaurus.
Pegomastax	Gen. et sp. nov	Valid	Sereno	Hettangian to Sinemurian	Elliot Formation	South Africa	A heterodontosaurid. The type species is Pegomastax africana.
Philovenator	Gen. et sp. nov	Valid	Xu et al.	Campanian	Wulansuhai Formation	China	A troodontid. Its type species is Philovenator curriei.
Rugocaudia	Gen. et sp. nov	Valid	Woodruff	Aptian or Albian	Cloverly Formation	USA	A titanosauriform sauropod. The type species is Rugocaudia cooneyi.
Saurolophus morrisi	sp nov	In press	Prieto-Márquez & Wagner	Maastrichtian	Moreno Formation	USA	A species of Saurolophus.
Sauroniops	Gen. et sp. nov	Valid	Cau, Dalla Vecchia & Fabbri	Cenomanian	Kem Kem Beds	Morocco	A carcharodontosaurid. The type species is Sauroniops pachytholus.
Sciurumimus	Gen. et sp. nov	Valid	Rauhut et al.	Upper Kimmeridgian		Germany	A megalosauroid. The type species is Sciurumimus albersdoerferi.	Sciurumimus.
Unescoceratops	Gen. et sp. nov	Valid	Ryan, Evans, Currie, Brown, & Brinkman	Late Campanian	Dinosaur Park Formation	Canada	A leptoceratopsid ceratopsian.
Yueosaurus	Gen. et sp. nov	Valid	Zheng, Jin, Shibata, Azuma, & Yu	Aptian-Cenomanian	Liangtoutang Formation	China	A basal ornithopod.
Yurgovuchia	Gen. et sp. nov	Valid	Senter et al..	Early Cretaceous, possibly Barremian	Cedar Mountain Formation	USA	A dromaeosaurid. The type species is Yurgovuchia doellingi.	Yurgovuchia.
Yutyrannus	Gen. et sp. nov	Valid	Xu et al..	Early Cretaceous	Yixian Formation	China	A basal tyrannosauroid. The type species is Yutyrannus huali.	Yutyrannus.

Newly named birds

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aegypius jinniushanensis	Sp nov	Valid	Zhang, Huang, James, & Hou	Middle Pleistocene		China	An Old World vulture.
Aix praeclara	Sp. nov	Valid	Zelenkov & Kurochkin	Middle Miocene		Mongolia	A duck, a species of Aix.
Anhinga walterbolesi	Sp nov	Valid	Worthy	Late Oligocene or early Miocene		Australia	A darter.
Asphaltoglaux	Gen. et sp. nov	In press	Campbell & Bocheński	Late Pleistocene	La Brea Tar Pits	USA	A miniature true owl. The type species is Asphaltoglaux cecileae.
Australopicus	Gen. et sp. nov	Valid	Manegold & Louchart	Early Pliocene	Varswater Formation	South Africa	A woodpecker. The type species is Australopicus nelsonmandelai.
Avolatavis	Gen. et sp. nov	Valid	Ksepka & Clarke	Eocene	Green River Formation	USA	A stem parrot, a member of Pan-Psittaciformes. The type species is Avolatavis tenens.
Brodavis	Gen. et sp. nov	Valid	Martin, Kurochkin, & Tokaryk	Maastrichtian, possibly also Campanian	Frenchman Formation Hell Creek Formation Nemegt Formation ?Sharon Springs Formation	Canada Mongolia USA	A hesperornithiform. The type species is B. americanus; the genus also contains B. baileyi, B. mongoliensis and possibly also B. varneri.
Chenoanas	Gen. et sp. nov	Valid	Zelenkov	Middle Miocene		Mongolia	A duck. The type species is Chenoanas deserta.
Glaucidium kurochkini	Sp nov	In press	Campbell & Bocheński	Late Pleistocene	La Brea Tar Pits	USA	A pygmy owl.
Hirundo selengenica	Sp nov	Valid	Zelenkov & Kurochkin	Late Pliocene	Chikoi Formation	Mongolia	A swallow, a species of Hirundo.
Jeholornis palmapenis	Sp nov	Valid	O'Connor, Sun, Xu, Wang & Zhou	Early Aptian	Jiufotang Formation	China	A species of Jeholornis.
Kairuku	Gen. et sp. nov	Valid	Ksepka, Fordyce, Ando, & Jones	Oligocene		New Zealand	A penguin. The genus contains two species: Kairuku waitaki and Kairuku grebneffi.
Lutetodontopteryx	Gen. et sp. nov	Valid	Mayr & Zvonok	Middle Eocene		Ukraine	A pelagornithid. The type species is Lutetodontopteryx tethyensis.
Mioquerquedula	Gen. et sp. et comb. nov.	Valid	Zelenkov & Kurochkin	Middle Miocene		France Mongolia	A duck. The type species is Mioquerquedula minutissima; genus also contains "Anas" velox (Milne-Edwards, 1868).
Otus mauli	Sp nov	Valid	Rando et al.	Quaternary		Portugal (Madeira)	A scops owl.
Picavus	Gen. et sp. nov	Valid	Mayr & Gregorová	Early Oligocene	Menilite Formation	Czech Republic	A stem group representative of Pici. The type species is Picavus litencicensis.
Pliocalcarius	Gen. et sp. nov	Valid	Zelenkov & Kurochkin	Late Pliocene		Mongolia	A probable close relative of longspurs. The type species is Pliocalcarius orkhonensis.
Pterocnemia mesopotamica	Sp. nov	Valid	Agnolín & Noriega	Late Miocene	Ituzaingó Formation	Argentina	A rhea.
Rhodospiza shaamarica	Sp nov	Valid	Zelenkov & Kurochkin	Late Pliocene		Mongolia	A finch, a species of Rhodospiza.
?Rupelrallus belgicus	Sp. nov	In press	Mayr	Early Oligocene		Belgium	A parvigruid gruiform, a possible species of Rupelrallus.
Samrukia	Gen. et sp. nov	Valid	Naish, Dyke, Cau, Escuillié, Godefroit	Late Cretaceous	Kyzylorda District	Kazakhstan	A possible basal ornithuromorph; alternatively, it could be a large pterosaur.
Sanshuiornis	Gen. et sp. nov	Valid	Wang, Mayr, Zhang, & Zhou	Middle Eocene	Huayong Formation	China
Schizooura	Gen. et sp. nov	Valid	Zhou, Zhou, & O’Connor	Aptian	Jiufotang Formation	China	A basal ornithuromorph. The type species is Schizooura lii.
Shengjingornis	Gen. et sp. nov	Valid	Li et al.	Early Cretaceous	Jiufotang Formation	China	A longipterygid enantiornithine. The type species is Shengjingornis yangi.
Tydea	Gen. et sp. nov	Valid	Mayr & Smith	Early Oligocene		Belgium	An albatross. The type species is Tydea septentrionalis.
Xiangornis	Gen. et sp. nov	Valid	Hu et al.	Early Cretaceous	Jiufotang Formation	China	An enantiornithine bird. The type species is Xiangornis shenmi.

Newly named pterosaurs

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Bellubrunnus	Gen. et sp. nov	Valid	Hone et al.	Late Kimmeridgian	Pfraundorf-Heitzenhofen Basin	Germany	A rhamphorhynchid. The type species is Bellubrunnus rothgaengeri.	Bellubrunnus.
Cuspicephalus	Gen. et sp. nov	In press	Martill & Etches	Kimmeridgian	Kimmeridge Clay Formation	England	A basal monofenestratan.	Cuspicephalus.
Europejara	Gen. et sp. nov	Valid	Vullo et al.	Upper Barremian	La Huérguina Formation	Spain	A tapejarid. The type species is Europejara olcadesorum.	Europejara.
Gladocephaloideus	Gen. et sp. nov	Valid	Lü, Ji, Wei, & Liu	Barremian	Yixian Formation	China	A gallodactylid ctenochasmatoid.
Guidraco	Gen. et sp. nov	Valid	Wang, Kellner, Jiang, & Cheng	Early Cretaceous	Jiufotang Formation	China	A close relative of Ludodactylus. The type species is Guidraco venator.	Guidraco.
Jianchangnathus	Gen. et sp. nov	Valid	Cheng, Wang, Jiang, & Kellner	Middle Jurassic	Tiaojishan Formation	China	A scaphognathid.
Moganopterus	Gen. et sp. nov	Valid	Lü et al.	Early Cretaceous	Yixian Formation	China	A boreopterid. The type species is Moganopterus zhuiana.
Qinglongopterus	Gen. et sp.	Valid	Lü, Unwin, Zhao, Gao & Shen.	Middle Jurassic	Tiaojishan Formation	China	A rhamphorhynchid.

Synapsids

Newly named non-mammalian synapsids

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Fortunodon	Gen. et comb. nov	Valid	Kurkin	Upper Permian		Russia	A dicynodont, a new genus for "Dicynodon" trautscholdi (Amalitzky, 1922) (considered to be a senior synonym of Vivaxosaurus permirus and assigned to the genus Vivaxosaurus by Kammerer et al., 2011). "Dicynodon" amalitzkii (Sushkin, 1926) is considered by Kurkin (2012) to be a second species of Fortunodon (Kammerer et al., 2011 classify it as the type species of the genus Peramodon.)	File:Pampaphoneus biccai cranium2.png Skull of Pampaphoneus biccai.
Novocynodon	Gen. et sp. nov	Valid	Ivakhnenko	Middle Permian		Russia	A thrinaxodontid cynodont. The type species is Novocynodon kutorgai.
Pampaphoneus	Gen. et sp. nov	Valid	Cisneros, Abdala, et al.	Guadalupian (Middle Permian)	Rio do Rasto Formation	Brazil	An anteosaurid dinocephalian.
Panchetocynodon	Gen. et sp. nov	Valid	Das & Gupta	Early Triassic	Panchet Formation	India	A cynodont. The type species is Panchetocynodon damodarensis.
Purbeckodon	Gen. et sp. nov	Valid	Butler, Sigogneau-Russell, & Ensom	Early Cretaceous	Purbeck Limestone Group	United Kingdom	A possible morganucodontan.
Sludica	Gen. et sp. nov	Valid	Ivakhnenko	Upper Permian		Russia	A procynosuchid cynodont. The type species is Sludica bulanovi.

Newly named non-eutherian mammals

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Abderites aisenense	Sp. nov	Valid	Abello & Rougier	Middle Miocene		Argentina Chile	A marsupial, a member of Paucituberculata, a species of Abderites.
Bolodon hydei	Sp. nov	In press	Cifelli, Davis & Sames	Late Berriasian or Valanginian	Lakota Formation	United States	A plagiaulacid multituberculate, a species of Bolodon.
Condorodon	Gen. et sp. nov	Valid	Gaetano & Rougier	Middle Jurassic	Cañadón Asfalto Formation	Argentina	An amphilestid. The type species is Condorodon spanios.
Infernolestes	Gen. et sp. nov	In press	Cifelli, Davis & Sames	Late Berriasian or Valanginian	Lakota Formation	United States	A spalacotheriid symmetrodontan. The type species is Infernolestes rougieri.
Kouriogenys	Gen. et comb. nov	Valid	Davis	Berriasian		United Kingdom	A peramuran mammal, a new genus for the species originally named Spalacotherium minus Owen 1871.
Lakotalestes	Gen. et sp. nov	In press	Cifelli, Davis & Sames	Late Berriasian or Valanginian	Lakota Formation	United States	A dryolestid trechnotherian. The type species is Lakotalestes luoi.
Neohelos davidridei	Species	In press	Black, Archer, Hand, & Godthelp	Middle Miocene	Riversleigh World Heritage Area fossil deposit	Australia	A diprotodontid, a species of Neohelos.
Neohelos solus	Species	In press	Black, Archer, Hand, & Godthelp	Middle Miocene	Riversleigh World Heritage Area fossil deposit	Australia	A diprotodontid, a species of Neohelos.
Passumys	Gen. et sp. nov	In press	Cifelli, Davis & Sames	Late Berriasian or Valanginian	Lakota Formation	United States	An allodontoid multituberculate. The type species is Passumys angelli.
Peramuroides	Gen. et sp. nov	Valid	Davis	Berriasian		United Kingdom	A peramuran mammal. The type species is Peramuroides tenuiscus.
Priscakoala	Gen. et sp. nov	Valid	Black, Archer, & Hand	Miocene	Riversleigh World Heritage Area fossil deposit	Australia	A phascolarctid.
Tjukuru	Gen. et sp. nov	Valid	Prideaux & Tedford	Pliocene	Tirari formation	Australia	A lagostrophine kangaroo. The type species is Tjukuru wellsi.

Newly named eutherians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acritophiomys	sp nov	Valid	Sallam, Seiffert & Simons	Late Eocene		Egypt	A phiomorph. The type species is Acritophiomys bowni.
Aepyceros datoadeni	sp nov	Valid	Geraads, Bobe & Reed	Pliocene	Hadar Formation	Ethiopia	A species of impala.
Afrasia	Gen. et sp. nov	Valid	Chaimanee et al.	Eocene	Pondaung Formation	Myanmar	A primitive anthropoid. The type species is Afrasia djijidae.
Afrodon gheerbranti	sp nov	Valid	De Bast, Sigé & Smith	Early Palaeocene	Hainin Formation	Belgium	An adapisoriculid, a species of Afrodon.
Aguascalientia minuta	sp nov	Valid	Rincon, Bloch, Suarez, MacFadden & Jaramillo	Early Miocene	Las Cascadas Formation	Panama	A floridatraguline camelid, a species of Aguascalientia.
Aguascalientia panamaensis	sp nov	Valid	Rincon, Bloch, Suarez, MacFadden & Jaramillo	Early Miocene	Las Cascadas Formation	Panama	A floridatraguline camelid, a species of Aguascalientia.
Alloptox (Mizuhoptox) japonicus	Subgen. and sp. nov	Valid	Tomida	Early Miocene		Japan	A pika, a species of Alloptox.
Alloptox katinkae	sp nov	Valid	Angelone & Hír	Early Middle Miocene		Hungary	A pika, a species of Alloptox.
Ancylotherium hellenicum	Sp. nov	Valid	Koufos	Late Miocene		Greece	A chalicothere, a species of Ancylotherium.
Andemys	Gen. et sp. nov	Valid	Bertrand et al.	Tinguirirican		Chile	A caviomorph rodent related to dasyproctids. The type species is Andemys termasi.
Archaeophocaena	Gen. et sp. nov.	Valid	Murakami et al.	Late Miocene	Koetoi Formation	Japan	A porpoise. The type species is Archaeophocaena teshioensis.
Babameryx	Gen. et sp. nov	Valid	Mennecart et al.	Late Oligocene		Switzerland	A ruminant. The type species is Babameryx engesseri.
Barberapithecus	Gen. et sp. nov	Valid	Alba & Moyà-Solà	Middle to Late Miocene		Spain	A crouzeliine pliopithecid. The type species is Barberapithecus huerzeleri.
Berrulestes	Gen. et 3 sp. nov	Valid	Hooker & Russell	Paleocene		France	A macroscelidean. The type species is Berrulestes phelizoni; the other species are B. pellouini and B. poirieri.
Bohaskaia	Gen. et sp. nov	Valid	Vélez-Juarbe & Pyenson	Early Pliocene	Yorktown Formation	United States	A monodontid. The type species is Bohaskaia monodontoides.
Bomburodon	Nom. nov.	Valid	Williamson & Carr	Paleocene		United States	A replacement name for the condylarth genus Bomburia Van Valen, 1978.
Borisodon	Gen. et comb. nov	Valid	Archibald & Averianov	Turonian		Kazakhstan	A zhelestid, a new genus for "Sorlestes" kara (Nessov, 1993).
Bustylus folieae	sp nov	Valid	De Bast, Sigé, & Smith	Early Palaeocene	Hainin Formation	Belgium	An adapisoriculid, a species of Bustylus.
Canis lupus maximus	Subsp. nov.	Valid	Boudadi-Maligne	Late Pleistocene		France	A subspecies of gray wolf.
Carposorex burkarti	sp nov	Valid	Hugueney, Mein & Maridet	Miocene		France	A shrew, a species of Carposorex.
Chubutomys navaensis	sp nov	In press	Pérez, Krause & Vucetich	Late Oligocene		Argentina	A member of Hystricognathi, a cavioid rodent; a species of Chubutomys.
Colpodon antucoensis	sp nov	Valid	Shockey et al.	Miocene	Cura-Mallín Formation	Chile	A leontiniid notoungulate, a species of Colpodon.
Cristadjidaumo skinneri	sp nov	Valid	Emry & Korth	Late Eocene	White River Formation	United States	An eomyid, a species of Cristadjidaumo.
Dryomomys dulcifer	sp nov	Valid	Chester & Beard	Late Paleocene	Big Multi Quarry	United States	A micromomyid plesiadapiform, a species of Dryomomys.
Elmerriggsia	Gen. et sp. nov	Valid	Shockey et al.	Late Oligocene		Argentina	A leontiniid notoungulate. The type species is Elmerriggsia fieldia.
Eotmantsoius	Gen. et sp. nov	Valid	Tabuce et al.	Middle or Late Eocene		Libya	A member of Macroscelidea. The type species is Eotmantsoius perseverans.
Eoviscaccia frassinettii	Sp. nov	Valid	Bertrand et al.	Tinguirirican		Chile	A caviomorph rodent related to chinchillids, a species of Eoviscaccia.
Eptenonnus	Gen. et sp. nov	Valid	Rosina & Semenov	Late Miocene		Ukraine	A vesper bat. The type species is Eptenonnus gritsevensis.
Gandheralophus	Gen. et sp. nov	Valid	Missiaen & Gingerich	Early Eocene	Ghazij Formation	Pakistan	An isectolophid tapiromorph. Its species are G. minor and G. robustus.
Garridomys	Gen. et sp. nov	In press	Kramarz, Vucetich & Arnal	Early Miocene	Cerro Bandera Formation	Argentina	A rodent closely related to chinchillids. The type species is Garridomys curunuquem.
Gazella harmonae	sp nov	Valid	Geraads, Bobe & Reed	Pliocene	Hadar Formation	Ethiopia	A species of gazelle.
Geotrypus minor	sp nov	In press	Ziegler	Early Oligocene		Germany	A talpid, a species of Geotrypus.
Gigarton	Gen. et 3 sp. nov	Valid	Hooker & Russell	Paleocene		France	A macroscelidean. The type species is Gigarton meyeri; the other species are G. sigogneauae and G. louisi.
Gobiolagus aliwusuensis	Sp. nov	Valid	Fostowicz-Frelik et al.	Middle Eocene		China	A palaeolagid lagomorph, a species of Gobiolagus.
Hanakia agadjaniani	sp nov	In press	Rosina & Rummel	Early Miocene		Germany	A vesper bat, a species of Hanakia.
Hesperogaulus shotwelli	sp nov	Valid	Calede & Hopkins	Miocene		United States	A mylagaulid, a species of Hesperogaulus.
Indusomys	Nom. nov	Valid	Gunnell et al.	Eocene		Pakistan	A replacement name for the primate genus Indusius Gunnell, Gingerich, Ul-Haw, Bloch, Kahn and Clyde, 2008.
Kolpochoerus millensis	Sp. nov	In press	Haile-Selassie & Simpson	Pliocene	Woranso-Mille and Gona sites	Ethiopia	A suid, a species of Kolpochoerus.
Kurdalagonus	Gen. et 2 sp. et comb. nov	Valid	Tarasenko & Lopatin	Miocene	Blinovo Formation	Russia	A baleen whale. The genus contains two new species: Kurdalagonus mchedlidzei and K. adygeicus, as well as the species "Cetotherium" maicopicum Spasskii, 1951.
Leptophoca amphiatlantica	sp nov	Valid	Koretsky, Ray & Peters	Miocene	Calvert Formation Breda Formation St Mary’s Formation	United States Netherlands	An earless seal.
Litolophus ghazijensis	Species	Valid	Missiaen & Gingerich	Early Eocene	Ghazij Formation	Pakistan	An eomoropid chalicotherioid, a species of Litolophus.
Louisina marci	sp nov	Valid	Hooker & Russell	Paleocene		France	A macroscelidean, a species of Louisina.
Megaleptictis	Gen. et sp. nov	Valid	Meehan & Martin	Chadronian/?Orellan	White River Group	United States	A large leptictid.
Meherrinia	Gen. et sp. nov	Valid	Geisler, Godfrey 7 Lambert	Late Miocene	Eastover Formation	United States	A river dolphin.
Melaneremia schrevei	Sp. nov	Valid	Hooker	Eocene (earliest Ypresian)	Woolwich Formation	United Kingdom	A microchoerine omomyid primate, a species of Melaneremia.
Mesonyx nuhetingensis	Sp. nov	Valid	Jin	Early Eocene	Arshanto Formation	China	A mesonychid, a species of Mesonyx.
Metanoiamys paradoxus	Species	Valid	Emry & Korth	Late Eocene	White River Formation	United States	An eomyid, a species of Metanoiamys.
Miophocaena	Gen. et sp. nov.	Valid	Murakami et al.	Late Miocene	Koetoi Formation	Japan	A porpoise. The type species is Miophocaena nishinoi.
Miostrellus petersbuchensis	sp nov	In press	Rosina & Rummel	Early Miocene		Germany	A vesper bat, a species of Miostrellus.
Mylagaulus cornusaulax	sp nov	Valid	Czaplewski	Late Miocene		United States	A mylagaulid rodent.
Myotis korotkevichae	Sp. nov	Valid	Rosina & Semenov	Late Miocene		Ukraine	A mouse-eared bat.
Namatomys erythrus	sp nov	Valid	Emry & Korth	Late Eocene	White River Formation	United States	An eomyid, a species of Namatomys.
Nementchatherium rathbuni	Sp. nov	Valid	Tabuce et al.	Middle or Late Eocene		Libya	A member of Macroscelidea, a species of Nementchatherium.
Neophiomys	Gen. et comb. nov	Valid	Coster et al.	Early Oligocene		Libya	A rodent, a member of Hystricognathi; a new genus for "Phiomys" paraphiomyoides Wood, 1968.
Nesiotites rafelinensis	sp nov	Valid	Rofes et al.	Early Pliocene		Spain	A shrew, a species of Nesiotites.
Nestoritherium linxiaense	sp nov	Valid	Chen, Deng, He & Chen	Early Late Miocene	Liushu Formation	China	A chalicothere, a species of Nestoritherium.
Ninamys	Gen. et comb. et sp. nov	In press	Vianey-Liaud, Rodrigues & Marivaux	Oligocene to early Miocene		China Kazakhstan Mongolia	An aplodontiid rodent, a new genus for "Prosciurus" arboraptus Shevyreva 1971; genus also contains "Prosciurus" daxnerae Lopatin 2000 and a new species Ninamys kazimierzi.
Pappocricetodon neimongolensis	Sp. nov	Valid	Li	Middle Eocene	Irdin Manha Formation	China	A cricetid rodent, a species of Pappocricetodon.
Paransomys	Gen. et comb. nov	In press	Vianey-Liaud, Rodrigues & Marivaux	Late Oligocene to early Miocene		Western Europe	An aplodontiid rodent, a new genus for "Sciurodon" descendens Dehm 1950 and "Plesispermophilus" argoviensis Stehlin and Schaub 1951.
Paschatherium levei	sp nov	Valid	Hooker & Russell	Early Eocene		France	A macroscelidean, a species of Paschatherium.
Phiocricetomys atavus	Sp. nov	Valid	Coster et al.	Early Oligocene		Libya	A rodent, a member of Hystricognathi; a species of Phiocricetomys.
Platychoerops antiquus	Sp. nov	Valid	Boyer, Costeur & Lipman	Paleocene		France	A plesiadapid, a species of Platychoerops.
Plecotus schoepfelii	sp nov	In press	Rosina & Rummel	Early Miocene		Germany	A vesper bat, a species of Plecotus.
Plesiotypotherium casirense	Sp. nov	Valid	Cerdeño et al.	Late Miocene		Bolivia	A mesotheriid notoungulatan, a species of Plesiotypotherium.
Primoprismus	Gen. et sp. nov	In press	Maridet et al.	Early Miocene		China	A microtoid cricetid. The type species is Primoprismus fejfari.
Pristifelis	Gen. et comb. nov	Valid	Salesa et al.	Miocene		Western Eurasia	A feline felid, a new genus for Felis attica.
Prodendrogale engesseri	sp nov	Valid	Ni & Qiu	Late Miocene		China	A treeshrew, a species of Prodendrogale.
Proeggysodon	Gen. et sp. nov	Valid	Bai & Wang	Late Eocene	Possibly Ulan Gochu Formation	China	An odd-toed ungulate, an eggysodontine rhinocerotoid. The type species is Proeggysodon qiui.
Prolouisina	Gen. et comb. nov	Valid	Hooker & Russell	Paleocene		Germany	A macroscelidean, a new genus for "Louisina" atavella (Russell, 1964).
Proremiculus	Gen. et sp. nov	Valid	De Bast, Sigé & Smith	Early Palaeocene	Hainin Formation	Belgium	An adapisoriculid. Its type species is Proremiculus lagnauxi.
Protictitherium thessalonikensis	sp nov	Valid	Koufos	Late Miocene		Greece	An ictitheriine hyena, a species of Protictitherium.
Pseudoloris cuestai	sp nov	Valid	Minwer-Barakat, Marigó & Moyà-Solà	Middle Eocene		Spain	A microchoerine omomyid, a species of Pseudoloris.
Pterophocaena	Gen. et sp. nov.	Valid	Murakami et al.	Late Miocene	Wakkanai Formation	Japan	A porpoise. The type species is Pterophocaena nishinoi.
Styriofelis vallesiensis	sp nov	Valid	Salesa et al.	Late Miocene		Spain	A feline felid, a species of Styriofelis.
Sulaimanius	Nom. nov	Valid	Gunnell et al.	Eocene		Pakistan	A replacement name for the primate genus Sulaimania Gunnell, Gingerich, Ul-Haw, Bloch, Kahn and Clyde, 2008.
Sylvochoerus	Gen. et sp. nov	Valid	Frailey & Campbell	Late Miocene		Western Amazon Basin	A peccary. The type species is Sylvochoerus woodburnei.
Tachyoryctoides engesseri	sp nov	Valid	Wang & Qiu	Early Miocene	Lanzhou Basin	China	A muroid rodent, a species of Tachyoryctoides.
Tachyoryctoides minor	sp nov	Valid	Wang & Qiu	Early Miocene	Lanzhou Basin	China	A muroid rodent, a species of Tachyoryctoides.
Teilhardimys brisswalteri	sp nov	Valid	Hooker & Russell	Paleocene		France	A macroscelidean, a species of Teilhardimys.
Thryptodon	Gen. et sp. nov	Valid	Hooker & Russell	Paleocene		France	A macroscelidean. The type species is Thryptodon brailloni.
Tinimomys tribos	sp nov	Valid	Chester & Beard	Late Paleocene	Big Multi Quarry	United States	A micromomyid plesiadapiform, a species of Tinimomys.
Tragomys	Gen. et sp. nov	Valid	Agusti, Bover & Alcover	Pliocene		Spain	A cricetid rodent. The type species is Tragomys macpheei.
Tupaia storchi	sp nov	Valid	Ni & Qiu	Late Miocene		China	A treeshrew, a species of Tupaia.
Turkanamys	Gen. et sp. nov	Valid	Marivaux et al.	Late Oligocene		Kenya	A phiomorph rodent. The type species is Turkanamys hexalophus.
Tuscahomys walshi	Sp. nov	Valid	Anemone, Dawson & Beard	Early Eocene (late early Wasatchian)		United States	A cylindrodontid rodent, a species of Tuscahomys.
Vasseuromys cristinae	sp nov	Valid	Ruiz-Sánchez et al.	Middle Miocene		Spain	A dormouse, a species of Vasseuromys.
Vasseuromys rambliensis	sp nov	Valid	Ruiz-Sánchez et al.	Lower Miocene		Spain	A dormouse, a species of Vasseuromys.
Victoriaceros	Gen. et sp. nov	Valid	Geraads, McCrossin & Benefit	Middle Miocene	Makobo beds	Kenya	An elasmotheriine rhinoceros. Type species is Victoriaceros kenyensis.
Victoriamys	Gen. et comb. nov	In press	Martin	Pleistocene		Italy Spain	A vole, a new genus for the species "Allophaiomys" chalinei.
Viretia	Gen. et comb. nov	Valid	Hugueney, Mein and Maridet	Miocene		France Slovakia	A shrew, a new genus for "Sorex" gracilidens.
Walbeckodon	Gen. et 2 sp. nov	Valid	Hooker & Russell	Paleocene		France Germany	A macroscelidean. The type species is Walbeckodon krumbiegeli; the second species is Walbeckodon girardi.
Waldochoerus	Gen. et sp. nov	Valid	Frailey & Campbell	Late Miocene		Western Amazon Basin	A peccary. The type species is Waldochoerus bassleri.
Witwatia sigei	Sp. nov	Valid	Ravel et al.	Early Eocene		Tunisia	A philisid bat, a species of Witwatia.
Yamatocetus	Gen. et sp. nov	Valid	Okazaki	Late Oligocene		Japan	An eomysticetid baleen whale. The type species is Yamatocetus canaliculatus.
Zhalmouzia	Gen. et sp. nov	In press	Averianov & Archibald in Averianov, Archibald & Dyke	Santonian or Campanian	Bostobe Formation	Kazakhstan	A zhelestid. The type species is Zhalmouzia bazhanovi.