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2022 in paleoentomology

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List of years in paleoentomology
In paleontology
2019
2020
2021
2022
2023
2024
2025
In paleobotany
2019
2020
2021
2022
2023
2024
2025
In arthropod paleontology
2019
2020
2021
2022
2023
2024
2025
In paleoichthyology
2019
2020
2021
2022
2023
2024
2025
In paleomalacology
2019
2020
2021
2022
2023
2024
2025
In paleomammalogy
2019
2020
2021
2022
2023
2024
2025
In reptile paleontology
2019
2020
2021
2022
2023
2024
2025
In archosaur paleontology
2019
2020
2021
2022
2023
2024
2025

This paleoentomology list records new fossil insect taxa that are to be described during the year 2022, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

Clade Amphiesmenoptera

Lepidoptera

Name Novelty Status Authors Age Type locality Location Notes Images

Acanthocorona hedida[1]

Sp. nov

Valid

Zhang, Shih & Engel in Zhang et al.

Cretaceous

Burmese amber

 Myanmar

A moth belonging to the family Lophocoronidae.

Acanthocorona venulosa[1]

Sp. nov

Valid

Zhang, Shih & Engel in Zhang et al.

Cretaceous

Burmese amber

 Myanmar

A moth belonging to the family Lophocoronidae.

Dolichoneura jorelisae[2]

Sp. nov

Sarto i Monteys, Hausmann, Baixeras & Peñalver in Sarto i Monteys et al.

Miocene

Dominican amber

 Dominican Republic

A species of Dolichoneura.

Lepidopteran research

  • A description of new specimens of caterpillars from the Cretaceous Burmese amber, expanding the morphological diversity of Cretaceous caterpillars, is published by Gauweiler et al. (2022), who also attempt to determine whether Cretaceous caterpillars might have represented an adequate food source for early birds.[3]
  • Review of the fossil record of caterpillars in Dominican and Mexican amber, including description of new specimens, is published by Haug et al. (2022).[4]

Tarachoptera

Name Novelty Status Authors Age Type locality Location Notes Images

Kinitocelis dashengi[5]

Sp. nov

Valid

Wang, Engel, Shih & Ren in Wang et al.

Cretaceous

Burmese amber

 Myanmar

Trichoptera

Name Novelty Status Authors Age Type locality Location Notes Images

Amberclipsis[6]

Gen. et 3 sp. nov

Valid

Wichard, Müller & Fischer

Cretaceous

Burmese amber

 Myanmar

A member of Annulipalpia related to the genus Pseudoneureclipsis. The type species is A. elegans; genus also includes new species A. oblongus and A. simplex.

Aniburia[7]

Gen. et sp. nov

Valid

Sukatsheva & Aristov

Early Cretaceous (Aptian)

Zaza Formation

 Russia

A member of the family Calamoceratidae. Genus includes new species A. hirta.

Archaeopolycentra longesilentia[8]

Sp. nov

Valid

Melnitsky & Ivanov

Late Cretaceous (Cenomanian)

Dolgan Formation
(Taymyr amber)

 Russia
( Krasnoyarsk Krai)

A member of the family Polycentropodidae.

Baissoferus elegans[7]

Sp. nov

Valid

Sukatsheva & Aristov

Early Cretaceous (Aptian)

Zaza Formation

 Russia

A member of the family Baissoferidae.

Cretapsyche kachini[9]

Sp. nov

Valid

Wichard & Espeland

Cretaceous

Burmese amber

 Myanmar

A member of Integripalpia belonging to the family Cretapsychidae.

Cretapsyche myanmari[9]

Sp. nov

Valid

Wichard & Espeland

Cretaceous

Burmese amber

 Myanmar

A member of Integripalpia belonging to the family Cretapsychidae.

Juraphilopotamus heteroclitus[7]

Sp. nov

Valid

Sukatsheva & Aristov

Early Cretaceous (Aptian)

Zaza Formation

 Russia

A member of the family Philopotamidae.

Khutelia[10]

Gen. et sp. nov

Valid

Sukatsheva & Sinitshenkova

Early Cretaceous

Tsagaantsav Formation

 Mongolia

A member of the family Baissoferidae. The type species is K. jarzembowskii.

Lepidochlamus[5]

Fam. et gen. et sp. nov

Valid

Wang et al.

Cretaceous

Burmese amber

 Myanmar

The type genus of the new family Lepidochlamidae. The type species is L. nodosa.

Neureclipsis acuta[11]

Sp. nov

Valid

Wichard & Xu

Cretaceous

Burmese amber

 Myanmar

A species of Neureclipsis.

Neureclipsis obtusa[11]

Sp. nov

Valid

Wichard & Xu

Cretaceous

Burmese amber

 Myanmar

A species of Neureclipsis.

Neureclipsis triangula[11]

Sp. nov

Valid

Wichard & Xu

Cretaceous

Burmese amber

 Myanmar

A species of Neureclipsis.

Palleptocerus grimaldii[12]

Sp. nov

Valid

Wichard & Müller

Cretaceous

Burmese amber

 Myanmar

A member of Leptoceroidea belonging to the family Palleptoceridae.

Pardoferus[7]

Gen. et sp. et comb. nov

Valid

Sukatsheva & Aristov

Early Cretaceous (Aptian)

Zaza Formation

 Russia

A member of the family Baissoferidae. Genus includes new species P. vitimicus, as well as "Baissoferus" sinitsae Sukatsheva & Vassilenko (2013).

Plectrocnemia bowangi[11]

Sp. nov

Valid

Wichard & Xu

Cretaceous

Burmese amber

 Myanmar

A species of Plectrocnemia.

Plectrocnemia ohlhoffi[11]

Sp. nov

Valid

Wichard & Xu

Cretaceous

Burmese amber

 Myanmar

A species of Plectrocnemia.

Protoclipsis[6]

Gen. et 3 sp. nov

Valid

Wichard, Müller & Fischer

Cretaceous

Burmese amber

 Myanmar

A member of Annulipalpia belonging to the group Psychomyiodea. The type species is P. ulmeri; genus also includes new species P. picteti and P. roeseli.

Other amphiesmenopterans

Name Novelty Status Authors Age Type locality Location Notes Images

Anecrotaulius[13]

Gen. et comb. nov

Valid

Engel

Early Cretaceous

Yixian Formation

 China

A member of the family Necrotauliidae, assigned by Engel (2022) to the new amphiesmenopteran order Necrotrichoptera.[13] The type species is "Necrotaulius" qingshilaense Hong (1984).

Microcarbonella[14]

Gen. et sp. nov

Valid

Prokop et al.

Carboniferous (Moscovian)

Osnabrück Formation

 Germany

A stem group Amphiesmenopteran.
The type species is M. paradoxa.

Necrotaulius americanus[15]

Sp. nov

Valid

Pierwola & Grimaldi

Late Triassic (Norian)

Cow Branch Formation

 United States

A Necrotauliidae Amphiesmenoptera.

Paranecrotaulius[13]

Gen. et comb. nov

Valid

Engel

Triassic

Madygen Formation

 Kyrgyzstan

A member of the family Necrotauliidae. The type species is "Necrotaulius" proximus Sukatsheva (1973).

Other amphiesmenopteran research

  • Khramov, Naugolnykh & Węgierek (2022) report the presence of elongate mouthparts with similar morphology to those of some present-day nectarivorous beetles and hymenopterans in Permian protomeropids from Russia, representing the earliest record of insects with siphonate-like mouthparts, which were possibly used to gather sugary fluids from the semi-closed ovulate organs of the gymnosperms of a peltaspermalean affinity known from the same locality.[16]

Clade Antliophora

Dipterans

Brachycerans

Name Novelty Status Authors Age Type locality Location Notes Images

Araripus[17]

Gen. et sp. nov

Valid

Carmo & Sampronha in Carmo et al.

Early Cretaceous (Aptian)

Crato Formation

 Brazil

A horse-fly. The type species is A. crassitibialis.

Archeoxyla[18]

Gen. et sp. nov

In press

Han et al.

Cretaceous

Burmese amber

 Myanmar

A wood soldier fly.
The type species is A. gigantea.

Brevitrichia messogenes[19]

Sp. nov

Valid

Greenwalt & Winterton in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A member of the family Scenopinidae belonging to the subfamily Scenopininae and the tribe Metatrichini.

Clemoxyla[18]

Gen. et sp. nov

In press

Han et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Xylomyidae. Genus includes new species C. aculeolata.

Cratotabanus cearensis[17]

Sp. nov

Valid

Carmo & Sampronha in Carmo et al.

Early Cretaceous (Aptian)

Crato Formation

 Brazil

A horse-fly.

Gujaratmyia[20]

Gen. et sp. nov

Valid

Bickel in Bickel et al.

Eocene

 India

A member of the family Dolichopodidae. The type species is G. rotunda.

Hoplocyrtoma eocenica[19]

Sp. nov

Valid

Sinclair & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A member of the family Hybotidae belonging to the subfamily Bicellariinae.

Leptogaster? cerestensis[21]

Sp. nov

In press

Nel & Jouault

Early Oligocene

 France

A member of the family Asilidae belonging to the subfamily Leptogastrinae.

Microphorella fragilis[19]

Sp. nov

Valid

Cumming & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A species of Microphorella.

Opetia americana[19]

Sp. nov

Valid

Amorim & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A member of the family Opetiidae.

Palaeoapsilocephala[19]

Gen. et sp. nov

Valid

Hauser & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A member of the family Apsilocephalidae. The type species is P. kishenehnensis.

Palaeomedeterus cambayensis[20]

Sp. nov

Valid

Bickel in Bickel et al.

Eocene

 India

A member of the family Dolichopodidae, possibly belonging to the subfamily Peloropeodinae.

Rajpardia[20]

Gen. et sp. nov

Valid

Bickel in Bickel et al.

Eocene

 India

A member of the family Dolichopodidae. The type species is R. grimaldii.

Tmemophlebia carolinae[19]

Sp. nov

Valid

Evenhuis & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A species of Tmemophlebia.

Nematocerans

Name Novelty Status Authors Age Type locality Location Notes Images

Afroscatopse[22]

Gen. et sp. nov

Valid

Szadziewski et al.

Miocene

Ethiopian amber

 Ethiopia

A member of the family Scatopsidae. The type species is A. haennii.

Antodicranomyia rubra[23]

Sp. nov

Valid

Kania-Kłosok, Perrichot & Krzemiński

Late Cretaceous (Cenomanian)

Charentese amber

 France

A Limoniid cranefly.

Archizelmira jarzembowskii[24]

Sp. nov

Valid

Lukashevich

Middle Jurassic (Bathonian)

Itat Formation

 Russia
( Krasnoyarsk Krai)

A member of the family Archizelmiridae.

Azana akarenos[19]

Sp. nov

Valid

Kerr & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A member of the family Mycetophilidae belonging to the subfamily Sciophilinae.

Chaoborus kishenehnicus[25]

Sp. nov

Valid

Raranov & Haug in Baranov et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A member of the family Chaoboridae.

Cretomanota[26]

Gen. et sp. nov

Amorim et al.

Early Cretaceous (Aptian)

Crato Formation

 Brazil

A member of the family Mycetophilidae belonging to the subfamily Leiinae. The type species is C. gondwanica.

Dilophus cerestensis[27]

Sp. nov

Valid

Nel, Legal & Coster

Early Oligocene

 France

A species of Dilophus.

Dilophus idanos[19]

Sp. nov

Valid

Fitzgerald & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A species of Dilophus.

Docidiadia grimaldii[28]

Sp. nov

Valid

Amorim & Brown

Cretaceous

Burmese amber

 Myanmar

A member of the family Diadocidiidae.

Eltxo grimaldii[29]

Sp. nov

Valid

Peñalver, Arillo & Nel

Early Cretaceous (Albian)

El Soplao amber

 Spain

A member of the family Cecidomyiidae.

Eloeophila eocenica[30]

Sp. nov

Santos, Teramoto Klein & Ribeiro

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A species of Eloeophila.

Eloeophila scudderi[30]

Nom. nov

Santos, Teramoto Klein & Ribeiro

Eocene

Florissant Formation

 United States
( Colorado)

A species of Eloeophila; a replacement name for Cladura maculata Scudder (1894).

Fushuniola[31]

Gen. et sp. nov

Valid

Fedotova & Perkovsky in Fedotova et al.

Eocene (Ypresian)

Fushun amber

 China

A member of the family Cecidomyiidae. The type species is F. mai.

Gonomyia burmitica[32]

Sp. nov

Valid

Kania-Kłosok et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A limoniid cranefly, a species of Gonomyia.

Gonomyia penalveri[32]

Sp. nov

Valid

Kania-Kłosok et al.

Early Cretaceous (Albian)

 Spain

A limoniid cranefly, a species of Gonomyia.

Hintelmanniella[25]

Gen. et sp. nov

Valid

Baranov & Haug in Baranov et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A member of the family Chironomidae belonging to the tribe Chironomini. The type species is H. noncatafractata.

Hoploscatopse[33]

Gen. et sp. nov

In press

Haenni & Ševčík

Cretaceous

Burmese amber

 Myanmar

A member of the family Scatopsidae. The type species is H. prisca.

Leptotarsus acciolii[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus alemaoi[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus bonifacioi[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus capanemai[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus coutoi[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus feijoi[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus ferreirai[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus lagosi[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus lemesi[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus pereirai[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus petrii[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Leptotarsus velosoi[34]

Sp. nov

Santos, Carvalho & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A species of Leptotarsus.

Liassorhyphus[35]

Gen. et sp. nov

Valid

Nel & Huang in Wang et al.

Early Jurassic (Pliensbachian-Toarcian)

Sangonghe Formation

 China

A member of the family Anisopodidae. The type species is L. liaoi.

Libanonemopalpus[36]

Gen. et sp. nov

Valid

Azar et al.

Early Cretaceous (Barremian)

Lebanese amber

 Lebanon

A member of the family Psychodidae belonging to the subfamily Bruchomyiinae. The type species is L. grimaldii.

Libanophlebotomites[37]

Gen. et sp. nov

Valid

Azar, Maalouf & Maksoud

Early Cretaceous (Barremian)

Lebanese amber

 Lebanon

A member of the family Psychodidae belonging to the subfamily Phlebotominae. Genus includes new species L. ramyi.

Macrocera apithanos[19]

Sp. nov

Valid

Kerr & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A species of Macrocera.

Monogedania[38]

Gen. et comb. nov

Valid

Szadziewski et al.

Eocene

Baltic amber

Baltic Sea region
 Germany  Ukraine

A member of the tribe Ceratopogonini. The type species is "Ceratopogon" clunipes Loew (1850).

Paleopsychoda jarzembowskii[39]

Sp. nov

Valid

Azar & Maksoud

Early Cretaceous (Barremian)

Lebanese amber

 Lebanon

A member of the family Psychodidae.

Plecia jorgecaridadi[40]

Sp. nov

In press

Rosse-Guillevic et al.

Miocene
Burdigalian

Dominican Amber

 Dominican Republic

A species of Plecia.

Plecia surieli[40]

Sp. nov

In press

Rosse-Guillevic et al.

Miocene
Burdigalian

Dominican Amber

 Dominican Republic

A species of Plecia.

Protoanisolarva[41]

Gen. et sp. nov

Valid

Peñalver et al.

Middle Triassic (Anisian)

Estellencs Formation

 Spain

A member of the stem group of the family Anisopodidae. The type species is P. juarezi.

Psectrosciara crassieton[19]

Sp. nov

Valid

Amorim & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A member of the family Scatopsidae belonging to the subfamily Psectrosciarinae.

Psectrosciara makrochaites[19]

Sp. nov

Valid

Amorim & Greenwalt in Greenwalt et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A member of the family Scatopsidae belonging to the subfamily Psectrosciarinae.

Pseudoerioptera[42]

Gen. et sp. nov

Men, Hu & Starkevich

Cretaceous

Burmese amber

 Myanmar

A member of the subfamily Chioneinae. Genus includes new species P. hubenqii.

Rheotanytarsus lacustris[25]

Sp. nov

Valid

Baranov & Haug in Baranov et al.

Eocene

Kishenehn Formation

 United States
( Montana)

A species of Rheotanytarsus.

Sycorax longistyla[43]

Sp. nov

Valid

Wagner, Agnihotri & Singh

Early Eocene

Cambay amber

 India

A species of Sycorax.

Toxorhina (Ceratocheilus) christelius[44]

Sp. nov

Valid

Kania-Kłosok, Krzemiński & Szwedo

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Toxorhina.

Triassomyia[45]

Gen. et sp. nov

Valid

Lukashevich

Triassic (LadinianCarnian)

Madygen Formation

 Kyrgyzstan

A member of the family Chaoboridae. The type species is T. shcherbakovi

Trichomyia (Trichomyia) fudalai[46]

Sp. nov

Valid

Azar, Pielowska-Ceranowska & Szwedo

Miocene

Dominican amber

 Dominican Republic

A species of Trichomyia.

Trichomyia (Septemtrichomyia) grimaldii[46]

Sp. nov

Valid

Azar, Pielowska-Ceranowska & Szwedo

Miocene

Dominican amber

 Dominican Republic

A species of Trichomyia.

Zwickina[47]

Gen. et sp. nov

Lukashevich & Vorontsov

Cretaceous

Burmese amber

 Myanmar

A member of the family Blephariceridae. Genus includes new species Z. minaevi.

Dipteran research
  • A redescription of Minyohelea nexuosa, based on data from a new specimen from the Lower Cretaceous Lebanese amber, is published by Pielowska-Ceranowska, Azar & Szwedo (2022).[48]
  • A review of the fossil record of the subfamily Microphorinae is published by Shamshev & Perkovsky (2022).[49]

Mecoptera

Name Novelty Status Authors Age Type locality Location Notes Images

Agetopanorpa sinica[50]

Sp. nov

Valid

Lian, Cai & Huang

Middle Triassic

 China

A member of the family Permochoristidae.

Baltipanorpa oppressiva[51]

Sp. nov

Valid

Soszyńska-Maj & Krzemiński in Soszyńska-Maj et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Panorpidae.

Burmorthophlebia[51]

Gen. et 2 sp. nov

Valid

Soszyńska-Maj, Krzemiński & Wang in Soszyńska-Maj et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Orthophlebiidae. The type species is B. multiprocessa Soszyńska-Maj, Krzemiński & Wang in Soszyńska-Maj et al.; genus also includes B. macularis Zhang et al..[52]

Burmothauma[53]

Gen. et sp. nov

In press

Zhang et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Eomeropidae. Genus includes new species B. eureka.

Cantabra[51]

Fam. et gen. et sp. nov

Valid

Soszyńska-Maj, Pérez-de la Fuente, Krzemiński & Wang in Soszyńska-Maj et al.

Early Cretaceous (Albian)

El Soplao amber

 Spain

A member of Panorpoidea belonging to the new family Cantabridae. The type species is C. soplao.

Chaohuchorista[54]

Gen. et sp. nov

Valid

Lian, Cai & Huang

Permian (Guadalupian)

Yinping Formation

 China

A member of the family Permochoristidae. Genus includes new species C. liaoi.

Chimaeropteron[55]

Gen. et 2 sp. nov

Valid

Bashkuev

 Russia

A member of the family Permotanyderidae. Genus includes new species C. antiquum and C. agmatos.

Eremobittacus opeth[56]

Sp. nov

Valid

Poschmann & Nel

Late Oligocene

Enspel Formation

 Germany

A hangingfly.

Itaphlebia elegana[57]

Sp. nov

Valid

Cao, Shih & Ren in Cao et al.

Middle Jurassic (BathonianCallovian)

Jiulongshan Formation

 China

A member of the family Nannochoristidae.

Itaphlebia procera[57]

Sp. nov

Valid

Cao, Shih & Ren in Cao et al.

Middle Jurassic (Bathonian–Callovian)

Jiulongshan Formation

 China

A member of the family Nannochoristidae.

Mesochorista tongchuanensis[50]

Sp. nov

Valid

Lian, Cai & Huang

Middle Triassic

 China

A member of the family Permochoristidae.

Permeca chaohuensis[58]

Sp. nov

Valid

Lian, Cai & Huang

Permian (Capitanian)

Yinping Formation

 China

A member of the family Permochoristidae.

Permotanyderites[55]

Gen. et sp. nov

Valid

Bashkuev

 Russia

A member of the family Permotanyderidae. Genus includes new species P. pseudopolycentropoides.

Permotanyderus sensibilis[55]

Sp. nov

Valid

Bashkuev

 Russia

A member of the family Permotanyderidae.

Permotanyderus sibiricus[55]

Sp. nov

Valid

Bashkuev

 Russia

A member of the family Permotanyderidae.

Rudimentanus[55]

Gen. et 2 sp. nov

Valid

Bashkuev

 Russia

A member of the family Permotanyderidae. Genus includes new species R. rieki and R. cisuralicus.

Clade Archaeorthoptera

†Cnemidolestida

Name Novelty Status Authors Age Type locality Location Notes Images

Roqueia[59]

Fam. et gen. et sp. nov

In press

Nel, Vallois & Duquesne

Carboniferous (Kasimovian)

 France

An archaeorthopteran belonging to the new family Roqueiidae, possibly belonging to the group Cnemidolestodea. Genus includes new species R. alpine.

Orthopterans

Name Novelty Status Authors Age Type locality Location Notes Images

Aenigmaraphidophora[60]

Gen. et sp. nov

Valid

Azar, Maalouf & Nel

Early Cretaceous

Lebanese amber

 Lebanon

A member of Tettigoniidea of uncertain affinities. The type species is A. mouniri.

Apiculatus[61]

Gen. et sp. nov

Valid

Yuan, Ma & Gu in Yuan et al.

Cretaceous (Albian-Cenomanian)

Burmese amber

 Myanmar

A member of the family Gryllidae belonging to the subfamily Oecanthinae. The type species is A. cretaceus.

Archelcana numbergerae[62]

Sp. nov

Valid

Heads, Thuy & Tamarri

Early Jurassic (Toarcian)

 Luxembourg

A member of the family Elcanidae.

Archelcana tina[62]

Sp. nov

Valid

Heads, Thuy & Tamarri

Early Jurassic (Toarcian)

 Luxembourg

A member of the family Elcanidae.

Bacharaboilus curvus[63]

Sp. nov

Valid

Gu et al.

Middle Jurassic

 China

A member of the family Prophalangopsidae.

Birmanioecanthus[61]

Gen. et sp. nov

Valid

Yuan, Ma & Gu in Yuan et al.

Cretaceous (Albian-Cenomanian)

Burmese amber

 Myanmar

A member of the family Gryllidae belonging to the subfamily Oecanthinae. The type species is B. haplostichus.

Burmaripipteryx[64]

Gen. et sp. nov

Zhao et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Ripipterygidae. Genus includes new species B. oblongus.

Chunxiania[65]

Gen. et sp. nov

Disputed

Xu et al.

Cretaceous

Burmese amber

 Myanmar

A mole cricket or a member of the family Gryllidae.[66] Genus includes new species C. fania. Cadena-Castañeda et al. (2023) considered it to be a junior synonym of Pherodactylus rectanguli (Xu, Fang & Wang, 2020).[66]

Curvospurus[67]

Gen. et sp. nov

In press

Liu, Yu & He

Cretaceous

Burmese amber

 Myanmar

A member of the family Trigonidiidae. The type species is C. huzhengkun.

Gryllobencain[68]

Gen. et sp. nov

Valid

Haug et al.

Cretaceous

Burmese amber

 Myanmar

An ensiferan of uncertain affinities. The type species is G. patrickmuelleri.

Lusitadischia[69]

Gen. et sp. nov

In press

Correia & Nel

Carboniferous (Gzhelian)

 Portugal

A member of the family Oedischiidae. The type species is L. sai.

Magnidactylus gracilis[70]

Sp. nov

Valid

Gu, Zheng, Cao & Yue in Gu et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Ripipterygidae.

Magnidactylus mirus[70]

Sp. nov

Valid

Gu, Zheng, Cao & Yue in Gu et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Ripipterygidae.

Monitelcana[71]

Gen. et sp. nov

Xu et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Elcanidae. The type species is M. penalveri.

Novaboilus ovatus[63]

Sp. nov

Valid

Gu et al.

Middle Jurassic

 China

A member of the family Prophalangopsidae.

Panorpidium acusicaudum[72]

Sp. nov

Valid

Tian, Ren & Gu

Early Cretaceous

Yixian Formation

 China

A member of the family Elcanidae.

Panorpidium maculosum[73]

Sp. nov

In press

Zhou et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Elcanidae.

Panorpidium validum[72]

Sp. nov

Valid

Tian, Ren & Gu

Early Cretaceous

Yixian Formation

 China

A member of the family Elcanidae.

Parabintoniella[74]

Gen. et sp. nov

Valid

Béthoux & Grauvogel-Stamm

Middle Triassic

Grès à Voltzia Formation

 France

A member of the family Bintoniellidae. The type species is P. papierae.

Proararipegryllus[75]

Gen. et sp. et comb. nov

In press

Gorochov & Coram

Early Cretaceous

Wealden Group

 United Kingdom

A member of Grylloidea. Genus includes new species P. andrewi, as well as "Araripegryllus" orientalis Gorochov, Jarzembowski & Coram (2006).

Probaisselcana minuta[76]

Sp. nov

In press

Gu et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Elcanidae.

Probaisselcana zhengi[76]

Sp. nov

In press

Gu et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Elcanidae.

Republicopteron[77]

Gen. et sp. nov

Valid

Archibald, Gu & Mathewes

Early Eocene

Klondike Mountain Formation

 United States
( Washington)

A Palaeorehniidae ensiferan.
The type species is R. douseae.

?Sibelcana websteri[75]

Sp. nov

In press

Gorochov & Coram

Early Cretaceous

Purbeck Group

 United Kingdom

A member of Elcanoidea.

Transigryllus[75]

Gen. et sp. nov

In press

Gorochov & Coram

Early Cretaceous

Wealden Group

 United Kingdom

A member of Grylloidea. Genus includes new species T. edmundi.

Tresdigitus gracilis[78]

Sp. nov

Disputed

Jiang et al.

Cretaceous

Burmese amber

 Myanmar

A mole cricket or a member of the family Gryllidae.[66] Cadena-Castañeda et al. (2023) considered it to be a junior synonym of Pherodactylus rectanguli (Xu, Fang & Wang, 2020).[66]

Ypopteron[77]

Gen. et sp. nov

Valid

Archibald, Gu & Mathewes

Early Eocene

Quilchena site

 Canada
( British Columbia)

A Palaeorehniidae ensiferan.
The type species is Y. nicola.

Orthopteran research

  • Putative liomopterid Alekhosara reticulata is reinterpreted as a possible locustavid caeliferan, and thus possibly the oldest known caeliferan, by Aristov & Gorochov (2022).[79]
  • Evidence of the presence of the earliest tympanal ears and sound-producing system in exceptionally preserved Mesozoic katydids, interpreted as indicating that katydids evolved a high diversity of singing frequencies at least by the Late Triassic, and complex acoustic communication at least by the Middle Jurassic, is presented by Xu et al. (2022).[80]

†Titanoptera

Name Novelty Status Authors Age Type locality Location Notes Images

Magnatitan[81]

Gen. et sp. nov

Valid

Park et al.

Late Triassic

Amisan Formation

 South Korea

A titanopteran belonging to the family Paratitanidae. The type species is M. jongheoni.

Other panorthopterans

Name Novelty Status Authors Age Type locality Country Notes Images

Hispanopteron[82]

Gen. et sp. nov

Valid

Santos et al.

Late Carboniferous

 Spain

Genus includes new species H. romerali. Announced in 2022; the final article version was published in 2023.

Clade Coleopterida

Coleopterans

Adephaga

Name Novelty Status Authors Age Type locality Country Notes Images

Antephilorhizus zerovae[83]

Sp. nov

Valid

Kirichenko-Babko & Perkovsky in Kirichenko-Babko, Perkovsky & Vasilenko

Eocene

Rovno amber

 Ukraine

A lebiine ground beetle.

Apermunda[84]

Gen. et comb. nov

In press

Kirejtshuk & Ansorge

Permian-Triassic transition

 Russia
( Kemerovo Oblast)

A member of the family Trachypachidae; a new genus for "Petrodromeus" minor Ponomarenko & Volkov (2013).

Cretomigadops[85]

Gen. et sp. nov

Liu et al.

Cretaceous

Burmese amber

 Myanmar

A migadopine ground beetle. Genus includes new species C. bidentatus.

Holcoptera rasnitsyni[86]

Sp. nov

Valid

Ponomarenko & Fedorenko in Ponomarenko, Fedorenko & Bashkuev

Late Triassic (CarnianNorian)

Protopivka Formation

 Ukraine

A member of the family Coptoclavidae.

Hujia[87]

Gen. et sp. nov

In press

Song, Jarzembowski & Xiao

Cretaceous

Burmese amber

 Myanmar

A tiger beetle belonging to the tribe Manticorini. The type species is H. rolfi.

Liassodites[84]

Gen. et sp. nov

In press

Kirejtshuk & Ansorge

Early Jurassic (Toarcian)

Ciechocinek Formation

 Germany

A member of the family Trachypachidae. The type species is L. obsti.

Novunda[84]

Nom. nov

In press

Kirejtshuk & Ansorge

Jurassic

 China
 Russia
( Zabaykalsky Krai)

A member of the family Trachypachidae; a replacement name for Unda Ponomarenko (1977).

Quasicalathus[88]

Gen. et 2 sp. et comb. nov

Valid

Schmidt & Will in Schmidt, Scholz & Will

Eocene

Rovno amber

Baltic Sea region
 Ukraine

A ground beetle belonging to the tribe Sphodrini. The type species is Q. conservans; genus also includes new species Q. agonicollis, as well as "Calathus" elpis Ortuño & Arillo (2009).

Archostemata

Name Novelty Status Authors Age Type locality Country Notes Images

Lepidomma beuteli[89]

Sp. nov

In press

Song, Jarzembowski & Xiao

Cretaceous

Burmese amber

 Myanmar

An ommatine.

Mallecupes zhangi[90]

Sp. nov

In press

Song, Jarzembowski & Xiao

Cretaceous

Burmese amber

 Myanmar

A member of the family Cupedidae.

Notocupes premeris[91]

Sp. nov

In press

Lee, Nam & Li

Early Cretaceous (Albian)

Jinju Formation

 South Korea

Myxophaga

Name Novelty Status Authors Age Type locality Country Notes Images

Bezesporum burmiticum[92]

Sp. nov

Fikáček et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Sphaeriusidae, related to extant taxa Bezesporum minutum (formerly Sphaerius minutus, the type species of the genus Bezesporum) and Bezesporum papulosum (formerly Sphaerius papulosus).

†Protocoleoptera

Protocoleopteran research
  • A study on the anatomy and affinities of Coleopsis archaica is published by Schädel, Yavorskaya & Beutel (2022).[93]
  • A study on the anatomy and phylogenetic affinities of members of the family Tshekardocoleidae is published by Boudinot et al. (2022), who name new clades Mesocoleoptera (containing all coleopterans except tshekardocoleids) and Metacoleoptera (containing all mesocoleopterans except permocupedids).[94]

Polyphaga

Bostrichiformia
Name Novelty Status Authors Age Type locality Country Notes Images

Alitrepanum[95]

Gen. et sp. nov

Disputed

Peng et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Bostrichidae. Genus includes new species A. aladelicatum. Legalov & Háva (2022) considered the genus Alitrepanum to be a junior synonym of the genus Poinarinius, though the authors maintained A. aladelicatum as a distinct species within the latter genus.[96]

Anthrenus larvalis[97]

Comb nov

In press

(Cockerell) Háva

Cenomanian

Burmese amber

 Myanmar

Moved from Dermestes larvalis (1917)

Lasioderma michalskii[98]

Sp. nov

Valid

Háva & Zahradník

Eocene

Baltic amber

 Poland

A species of Lasioderma.

Nothattagenus[99]

Gen. et comb. nov

Valid

Li & Cai in Li, Huang & Cai

Cretaceous

Burmese amber

 Myanmar

A member of the family Dermestidae belonging to the subfamily Orphilinae; a new genus for "Attagenus" burmiticus Cai, Háva & Huang (2017).

Poinarinius antonkozlovi[96]

Sp. nov

Valid

Legalov & Háva

Cretaceous

Burmese amber

 Myanmar

A member of the family Bostrichidae.

Poinarinius aristovi[96]

Sp. nov

Valid

Legalov & Háva

Cretaceous

Burmese amber

 Myanmar

A member of the family Bostrichidae.

Poinarinius borowskii[96]

Sp. nov

Valid

Legalov & Háva

Cretaceous

Burmese amber

 Myanmar

A member of the family Bostrichidae.

Poinarinius cretaceus[96]

Sp. nov

Valid

Legalov & Háva

Cretaceous

Burmese amber

 Myanmar

A member of the family Bostrichidae.

Poinarinius lesnei[96]

Sp. nov

Valid

Legalov & Háva

Cretaceous

Burmese amber

 Myanmar

A member of the family Bostrichidae.

Poinarinius perkovskyi[96]

Sp. nov

Valid

Legalov & Háva

Cretaceous

Burmese amber

 Myanmar

A member of the family Bostrichidae.

Poinarinius zahradniki[96]

Sp. nov

Valid

Legalov & Háva

Cretaceous

Burmese amber

 Myanmar

A member of the family Bostrichidae.

Stagetus michalskii[98]

Sp. nov

Valid

Háva & Zahradník

Eocene

Baltic amber

 Poland

A species of Stagetus.

Stagetus szydlowskae[98]

Sp. nov

Valid

Háva & Zahradník

Eocene

Baltic amber

 Poland

A species of Stagetus.

Stagetus zahradniki[100]

Sp. nov

Valid

Háva

Eocene

Baltic amber

 Poland

A species of Stagetus.

Sucinoptinus zahradniki[101]

Sp. nov

Valid

Háva

Eocene

Baltic amber

 Poland

A species of Sucinoptinus.

Tuberphradonoma secunda[102]

Sp. nov

Valid

Háva

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Dermestidae belonging to the subfamily Megatominae and the tribe Megatomini.

Cucujiformia
Name Novelty Status Authors Age Type locality Country Notes Images

Alloterocucus[103]

Gen. et sp. nov

Valid

Li, Leschen, Liu & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of Cucujoidea that may be allied to the family Lamingtoniidae. The type species is A. atratus.

Anancites[104]

Nom. nov

Valid

Perkovsky

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Lymexylidae; a replacement name for Adamas Chen & Zhang (2020).

Archaeoheilus gallicus[105]

Sp. nov

Valid

Legalov, Kirejtshuk & Nel

Paleocene

 France

A member of the family Curculionidae belonging to the subfamily Molytinae.

Atomaria (Anchicera) alekseevi[106]

Sp. nov

Valid

Lyubarsky & Bukejs

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A species of Atomaria.

Atomaria (Anchicera) perkovskyi[106]

Sp. nov

Valid

Lyubarsky & Bukejs

Eocene

Rovno amber

 Ukraine

A species of Atomaria.

Austronausibius aenigmatista[107]

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Rovno amber

 Ukraine

A species of Austronausibius.

Baltoconapium vilhelmseni[108]

Sp. nov

Valid

Legalov

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Brentidae.

Chramesus boninoi[109]

Sp. nov

Legalov

Miocene

Dominican amber

 Dominican Republic

A species of Chramesus.

Cretakarenni shaoi[110]

Sp. nov

Valid

Li & Cai in Li, Huang & Cai

Cretaceous

Burmese amber

 Myanmar

A member of the family Monotomidae.

Cretostenotarsus[111]

Gen. et sp. nov

Valid

Tomaszewska, Szawaryn & Arriaga-Varela

Cretaceous (Albian-Cenomanian)

Burmese amber

 Myanmar

A member of the family Endomychidae. The type species is C. striatus.

Cyrtanaspis kerneggeri[112]

Sp. nov

Valid

Perkovsky et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Cyrtanaspis.

Delteredolaemus[113]

Gen. et sp. nov

Valid

Li & Cai in Li, Huang & Cai

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of the family Teredidae. The type species is D. hei.

Demimaea gratshevi[114]

Sp. nov

Valid

Legalov, Vasilenko & Perkovsky

Eocene

Rovno amber

 Ukraine

A member of the family Curculionidae.

Demimaea zherichini[114]

Sp. nov

Valid

Legalov, Vasilenko & Perkovsky

Eocene

Rovno amber

 Ukraine

A member of the family Curculionidae.

Eduardoxenus rasnitsyni[115]

Sp. nov

Legalov

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A member of the family Anthribidae belonging to the subfamily Choraginae.

Eocenocolaspis[116]

Gen. et sp. nov

Valid

Bukejs, Moseyko & Alekseev

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A leaf beetle belonging to the subfamily Eumolpinae. Genus includes new species E. aurichalceus.

Eustrophus praecursor[117]

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A species of Eustrophus.

Gongyloceria[118]

Gen. et sp. nov

Valid

Legalov & Poinar

Miocene

Dominican amber

 Dominican Republic

An ambrosia beetle belonging to the tribe Mecopelmini. The type species is G. dominicana.

Grzymalia[119]

Gen. et sp. nov

In press

Bao et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Aderidae. Genus includes new species G. wukong.

Homalenchodes[120]

Gen. et sp. nov

Valid

Li, Hsiao, Yoshitomi & Cai in Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Melandryidae. The type species is H. jarzembowskii.

Hylastes perkovskyi[109]

Sp. nov

Legalov

Eocene

Rovno amber

 Ukraine

A species of Hylastes.

Leiestes tomaszewskae[121]

Sp. nov

Valid

Alekseev & Bukejs in Alekseev, Kairišs & Bukejs

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A species of Leiestes.

Paonaupactus zosimovichi[122]

Sp. nov

Valid

Legalov, Vasilenko & Perkovsky

Eocene

Rovno amber

 Ukraine

A member of the family Curculionidae belonging to the subfamily Entiminae and the tribe Anypotactini.

Pelretes bicolor[123]

Sp. nov

In press

Zhao et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Kateretidae.

Placonotus sepulvedae[124]

Sp. nov

Valid

Gomes & Souza

Miocene

Mexican amber

 Mexico

A species of Placonotus.

Pogonocherus minimus[125]

Sp. nov

Valid

Vitali

Eocene

Baltic amber

Europe (Baltic Sea region)

A longhorn beetle, a species of Pogonocherus.

Pogonocherus scutellaris[125]

Sp. nov

Junior homonym

Vitali

Eocene

Baltic amber

Europe (Baltic Sea region)

A longhorn beetle, a species of Pogonocherus.
The specific name is shared with Pogonocherus scutellaris Mulsant (1846);[126] Vitali (2023) coined a replacement name Pogonocherus scutellatus.[127]

Poliaenus europaeus[128]

Sp. nov

Valid

Vitali & Perkovsky

Eocene

Rovno amber

 Ukraine

A species of Poliaenus.

Promethis undulatus[129]

Sp. nov

Valid

Nabozhenko & Tanaka

Miocene

Isarizawa Formation

 Japan

A darkling beetle belonging to the subfamily Stenochiinae and the tribe Cnodalonini.

Protonitidula[130]

Gen. et sp. nov

Valid

Zhao, Huang & Cai

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A sap beetle. The type species is P. neli.

Pseudanthonomus antonkozlovi[131]

Sp. nov

Valid

Legalov

Miocene

Mexican amber

 Mexico

A species of Pseudanthonomus.

Pseudauletes (Eopseudauletes) balticus[132]

Sp. nov

Valid

Legalov

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Rhynchitidae.

Raractocetus sverlilo[133]

Sp. nov

Valid

Nazarenko, Perkovsky & Yamamoto in Yamamoto et al.

Late Eocene

Rovno amber

 Ukraine

A member of the family Lymexylidae. Originally described as a species of Raractocetus, but subsequently transferred by Li et al. (2022) to the genus Cretoquadratus.[134]

Rhomeocalpsua[135]

Gen. et sp. nov

Valid

Li, Tomaszewska & Cai in Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Endomychidae. The type species is R. torosa.

Rovnoeurops[136]

Gen. et sp. nov

In press

Alekseev & Bukejs

Eocene

Rovno amber

 Ukraine

A member of the family Monotomidae. Genus includes new species R. mckellari.

Scaporetes[137]

Gen. et sp. nov

Valid

Zhao, Huang & Cai

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of the family Kateretidae. The type species is S. rectus.

Synchita andrisbukejsi[138]

Sp. nov

Valid

Alekseev

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A species of Synchita.

Thallisellites[139]

Gen. et sp. nov

Valid

Kupryjanowicz, Lyubarsky & Perkovsky

Late Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Erotylidae belonging to the subfamily Languriinae. The type species is T. olgae.

Thanerosus[140]

Gen. et sp. nov

Valid

Peris & Kolibáč in Peris, Mähler & Kolibáč

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Thanerocleridae. The type species is T. antiquus.

Vetuprostomis angularis[141]

Sp. nov

Valid

Li & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of the family Prostomidae.

Vetuprostomis gaoi[141]

Sp. nov

Valid

Li & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of the family Prostomidae.

Wanachia pinisucciniferae[142]

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Baltic amber

Europe
(Baltic Sea region)

A member of the family Melandryidae.

Xenostanus[143]

Gen. et sp. nov

Valid

Li, Szawaryn & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of the family Corylophidae. The type species is X. jiangkuni.

Yakutia[144]

Gen. et sp. nov

In press

Telnov et al.

Late Cretaceous (Cenomanian)

Yakutian amber

 Russia
( Sakha)

A member of the family Mordellidae. Genus includes new species Y. sukachevae.

Zavaljus lyubarskyi[145]

Sp. nov

In press

Alekseev & Bukejs

Eocene

Rovno amber

 Ukraine

A species of Zavaljus.

Cucujiform research
  • Description of new specimens of scraptiid larvae with large, elongated terminal ends from the Eocene Baltic amber and Cretaceous Burmese amber (the latter representing the oldest record of these larvae reported to date), and a study on the morphological variation of these larvae is published by Zippel et al. (2022).[146]
  • Revision of the fossil lymexylids originally assigned to the genus Raractocetus is published by Li et al. (2022), who transfer the species Raractocetus balticus, R. extinctus, R. fossilis and R. sverlilo to the genus Cretoquadratus, and consider the species Cretoquadratus engeli to be a junior synonym of Cretoquadratus fossilis.[134]
Elateriformia
Name Novelty Status Authors Age Type locality Country Notes Images

Ajkaelater[147]

Gen. et sp. nov

Valid

Szabó et al.

Late Cretaceous (Santonian)

Ajka Coal Formation

 Hungary

An elaterid click beetle.
The type species is A. merkli.

Anomocephalobus[148]

Gen. et sp. nov

Valid

Li, Jäch & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of the family Limnichidae belonging to the subfamily Cephalobyrrhinae. The type species is A. liuhaoi.

Asiocnemis colombicus[149]

Sp. nov

Valid

Muona

Quaternary, exact age uncertain

Colombian resin

 Colombia

A member of the family Eucnemidae belonging to the subfamily Macraulacinae.

Bipogonia[150]

Gen. et 2 sp. nov

Valid

Li, Kundrata & Cai in Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Artematopodidae. Genus includes new species B. trivialis and B. fortis.

Brevipterus[151]

Gen. et comb. et 3 sp. nov

Valid

Zhao et al.

Cretaceous

Burmese amber

 Myanmar

A soldier beetle. Genus includes "Sanaungulus" strungei Fanti & Damggard (2019), as well as new species B. acutiapicis, B. abtusiapicis and B. megacephalus.

Burmochares[152]

Gen. et sp. nov

Valid

Kirejtshuk & Prokin

Cretaceous (Albian-Cenomanian)

Burmese amber

 Myanmar

A member of the family Limnichidae belonging to the subfamily Limnichinae. The type species is B. groehni.

Burmogonus[153]

Gen. et sp. nov

Valid

Triskova et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A click beetle belonging to the tribe Elaterini. The type species is B. cretaceus.

Burmomiles parisii[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle belonging to the subfamily Cantharinae.

Burmomiles volpei[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle belonging to the subfamily Cantharinae.

Cacomorphocerus thomasiwentzeli[155]

Sp. nov

Valid

Wentzel, Bonino & Fanti

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A soldier beetle belonging to the subfamily Cantharinae.

Carinibipogonia[150]

Gen. et sp. nov

Valid

Li, Kundrata & Cai in Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Artematopodidae. The type species is C. xiai.

Cessator anachronicus[156]

Sp. nov

Valid

Ferrera & Ivie in Ferreira et al.

Oligocene-Miocene

Dominican amber

 Dominican Republic

A member of the family Lycidae belonging to the subfamily Lycinae and the tribe Leptolycini.

Chauliognathus koonwarra[157]

Sp. nov

Valid

Fanti

Early Cretaceous (Aptian)

Koonwarra Fossil Bed

 Australia

A soldier beetle, a species of Chauliognathus.

Cretaceocoleus[158]

Gen. et sp. nov

Valid

Tihelka & Cai in Tihelka et al.

Cretaceous (Albian–Cenomanian)

Burmese amber

 Myanmar

A member of Dryopoidea belonging to the family Mastigocoleidae. The type species is C. saetosus Tihelka, Kundrata & Cai.

Damzenium[159]

Gen. et sp. nov

Valid

Kazantsev & Bocak

Late Eocene

Rovno amber

 Ukraine

A member of the family Lycidae belonging to the subfamily Erotinae and the tribe Erotini. The type species is D. rivnense.

Dariuszelater[160]

Gen. et sp. nov

In press

Zhao, Shih & Ren

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A click beetle. The type species is D. tarnawskii.

Elektrokleinia thiloi[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle belonging to the subfamily Cantharinae.

Elektrokleinia zahradniki[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle belonging to the subfamily Cantharinae.

Falsoceratoprion fumagalliae[161]

Sp. nov

Valid

Ferreira in Ferreira & Tettamanzi

Oligocene-Miocene

Dominican amber

 Dominican Republic

A member of the family Lycidae belonging to the subfamily Lycinae and the tribe Calopterini. The type species of the genus Falsoceratoprion, which also includes an extant species Falsoceratoprion pecki.[162]

Helcophorus ekaterinae[163]

Sp. nov

Valid

Kazantsev & Perkovsky

Eocene

Rovno amber

 Ukraine

A member of the family Lycidae. Originally described as a species of Helcophorus, but subsequently transferred to the genus Hiekeolycus.[164]

Hernandochares[152]

Gen. et comb. nov

Valid

Kirejtshuk & Prokin

Eocene

Baltic amber

Europe (Gdańsk Bay area)

A member of the family Limnichidae. The type species is "Platypelochares" electricus Hernando, Szawaryn & Ribera (2018).

Hukawngichthyurus barsevskisi[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle belonging to the subfamily Cantharinae.

Hukawngichthyurus ditaddeoi[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle belonging to the subfamily Cantharinae.

Isorhipis muonai[165]

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Baltic amber

Europe
(Baltic Sea region)

A species of Isorhipis.

Lasiosyne integera[166]

Sp. nov

In press

Guo et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Lasiosynidae.

Lasiosyne parva[166]

Sp. nov

In press

Guo et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Lasiosynidae.

Malthodes markpankowskii[167]

Sp. nov

Valid

Pankowski & Fanti

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A species of Malthinus.

Malthodes masoni[167]

Sp. nov

Valid

Pankowski & Fanti

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A species of Malthinus.

Mantimalthinus arturi[168]

Sp. nov

Valid

Fanti

Eocene

Baltic amber

 Poland

A soldier beetle belonging to the subfamily Malthininae and the tribe Malchinini.

Mastigocoleus[158]

Fam. et gen. et 2 sp. nov

Valid

Tihelka & Cai in Tihelka et al.

Cretaceous (Aptian–Cenomanian)

Burmese amber

 Brazil
 Myanmar

A member of Dryopoidea, the type genus of the new family Mastigocoleidae Tihelka, Jäch, Kundrata & Cai. The type species is M. resinicola Tihelka & Cai; genus also includes M. rhinoceros Tihelka & Cai.

Neusiokia[149]

Gen. et sp. nov

Valid

Muona

Late Cretaceous

Black Creek Formation

 United States
( North Carolina)

A member of the family Eucnemidae belonging to the subfamily Macraulacinae. The type species is N. appalachiensis.

Nothotytthonyx[169]

Gen. et sp. nov

Valid

Li, Biffi, Kundrata & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A soldier beetle. The type species is N. serratus.

Parelateriformius grimaldii[170]

Sp. nov

Valid

Li & Cai in Li et al.

Jurassic

Daohugou Beds

 China

A member of the family Dascillidae.

Poinarelektronmiles cuaroni[171]

Sp. nov

Valid

Bramanti & Fanti

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Ptilodactyla odnosum[172]

Sp. nov

Telnov, Perkovsky, Kundrata & Bukejs in Telnov et al.

Eocene

Rovno amber

 Ukraine

A species of Ptilodactyla.

Sanaungulus dominikiweissbachi[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus dunlopi[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus electrum[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus emarginaticollis[173]

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus franziskaeweissbachae[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus havai[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus imparitibius[173]

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus kachinensis[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus kirstenaeweissbachae[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus laticoxa[173]

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus leniae[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus multiramus[173]

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus myanmaricus[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus myitkyinaensis[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus nalae[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus nilsi[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus perkovskyi[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus peteriruedeli[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus ruficollis[173]

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus temporiscapsula[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Sanaungulus ypogaeum[154]

Sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle.

Silis (Silis) boninoi[174]

Sp. nov

Valid

Fanti & Pankowski

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A species of Silis.

Thambus woodruffi[149]

Sp. nov

Valid

Muona

Cenozoic (20 to 40 million years old)

Dominican amber

 Dominican Republic

A member of the family Eucnemidae belonging to the subfamily Macraulacinae.

Tytthonyx milleri[175]

Sp. nov

Valid

Ivie, Fanti & Ferreira

Miocene

Dominican amber

 Dominican Republic

A species of Tytthonyx.

Varcalium[176]

Gen. et sp. nov

Valid

Li, Ruta, Tihelka & Cai in Li et al.

Cretaceous (Albian-Cenomanian)

Burmese amber

 Myanmar

A member of the family Scirtidae. The type species is V. lawrencei.

Vetubrachypsectra huchengi[177]

Sp. nov

Valid

Li, Kundrata & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of the family Brachypsectridae.

Vitalfranzius[154]

Gen. et 2 sp. nov

Valid

Fanti & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A soldier beetle. The type species is V. burmiticus; genus also includes V. cretaceus.

Elateriform research
  • The first fossil larva of a representative of Elmidae reported to date is described from the Eocene Baltic amber by Zippel et al. (2022).[178]
Scarabaeiformia
Name Novelty Status Authors Age Type locality Country Notes Images

Anisoodontus[179]

Gen. et 2 sp. nov

In press

Wu et al.

Cretaceous

Burmese amber

 Myanmar

A stag beetle. Genus includes new species A. qizhihaoi and A. xiafangyuani.

Ceratocanthus huarongii[180]

Sp. nov

Valid

Lu, Ballerio & Bai in Lu et al.

Miocene

Dominican amber

 Dominican Republic

A species of Ceratocanthus.

Litholamprima qizhihaoi[181]

Sp. nov

In press

Jiang et al.

Early Cretaceous

Yixian Formation

 China

A stag beetle.

Nesopalla succini[180]

Sp. nov

Valid

Lu, Ballerio & Bai in Lu et al.

Miocene

Dominican amber

 Dominican Republic

A member of the family Hybosoridae belonging to the subfamily Ceratocanthinae.

Palaeopycnus[180]

Gen. et 2 sp. nov

Valid

Lu, Ballerio & Bai in Lu et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Hybosoridae belonging to the subfamily Ceratocanthinae. The type species is P. circus; genus also includes P. fushengii.

Prolucanus[182]

Gen. et sp. nov

Qi et al.

Early Cretaceous

Yixian Formation

 China

A stag beetle. The type species is P. beipiaoensis.

Scarabaeiform research
Staphyliniformia
Name Novelty Status Authors Age Type locality Country Notes Images

Afristenus[184]

Gen. et sp. nov

Mnguni, McKay & Badenhorst

Late Cretaceous (probably Turonian)

 Botswana

A rove beetle belonging to the family Steninae. The type species is A. orapensis.

Amplectister terapoides[185]

Sp. nov

Yamamoto & Caterino

Cretaceous (Albian–Cenomanian)

Burmese amber

 Myanmar

A member of the family Histeridae.

Anapleus kachinensis[186]

Sp. nov

Valid

Jiang, Caterino & Chen

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Anapleus.

Angucharcotes[187]

Gen. et sp. nov

Li et al.

Cretaceous

Burmese amber

 Myanmar

A rove beetle belonging to the subfamily Phloeocharinae. Genus includes new species A. thayerae.

Coomania enkarsios[188]

Sp. nov

Valid

Jenkins Shaw et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A rove beetle belonging to the subfamily Coomaniinae.

Coomania megistos[188]

Sp. nov

Valid

Jenkins Shaw et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A rove beetle belonging to the subfamily Coomaniinae.

Coomania yini[188]

Sp. nov

Valid

Jenkins Shaw et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A rove beetle belonging to the subfamily Coomaniinae.

Crenossidium[189]

Gen. et sp. nov

Valid

Li, Newton & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of the family Ptiliidae. The type species is C. slipinskii.

Cretaceonanobius[190]

Gen. et sp. nov

Valid

Liu et al.

Early Cretaceous

Yixian Formation

 China

A rove beetle belonging to the subfamily Pseudopsinae. The type species is C. fossilis.

Derolathrus capdoliensis[191]

Sp. nov

Valid

Tihelka et al.

Cretaceous (Albian-Cenomanian)

Charentese amber

 France

A species of Derolathrus.

Europharinodes[192]

Gen. et sp. nov

Valid

Yin & Cai in Yin et al.

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A rove beetle belonging to the subfamily Pselaphinae and the tribe Hybocephalini. The type species is E. schaufussi.

Pareburniola[193]

Gen. et sp. nov

Valid

Zilberman, Yin & Cai

Miocene

Dominican amber

 Dominican Republic

A rove beetle belonging to the subfamily Aleocharinae and the tribe Corotocini. The type species is P. dominicana.

Priochirus trisclerite[194]

Sp. nov

Valid

Peng et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Priochirus.

Yprezethinus[195]

Gen. et sp. nov

Valid

Parker

Eocene (Ypresian)

Cambay amber

 India

A rove beetle belonging to the subfamily Pselaphinae and the tribe Bythinoplectini. The type species is Y. grimaldii.

Staphyliniform research

Clade Dictyoptera

Name Novelty Status Authors Age Type locality Country Notes Images

Alienopterix santonicus[197]

Sp. nov

Valid

Szabó et al.

Late Cretaceous (Santonian)

Ajka Coal Formation

 Hungary

A member of the family Alienopteridae.

Anisotermes bourguignoni[198]

Sp. nov

Jouault et al.

Cretaceous

 Myanmar

A termite.

Bimini[199]

Gen. et sp. nov

Vršanský in Vršanský et al.

Late Jurassic

 Russia

A cockroach belonging to the family Corydiidae. Genus includes new species B. gorevoesis

Corydoblatta[199]

Gen. et sp. nov

Hinkelman & Vršanský in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A cockroach belonging to the family Corydiidae. Genus includes new species C. subrotunda.

Cryptotermes pouilloni[200]

Sp. nov

Valid

Jouault & Nel

Miocene

Dominican amber

 Dominican Republic

A termite, a species of Cryptotermes.

Facioblatta[201]

Gen. et sp. nov

Valid

Hinkelman

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A cockroach belonging to the family Liberiblattinidae. Genus includes new species F. perfidia.

Hypercercoula[202]

Gen. et sp. nov

In press

Qiu

Cretaceous

Burmese amber

 Myanmar

A member of Blattoidea. Genus includes new species H. jiewenae.

Litaneutria pilosuspedes[203]

Sp. nov

Valid

Terríquez-Beltrán, Riquelme & Varela-Hernández

Miocene

Mexican amber

 Mexico

An amelid mantis, a species of Litaneutria.

Longitermes[198]

Gen. et sp. nov

Jouault et al.

Cretaceous

 Myanmar

A termite. Genus includes new species L. pulcher.

Magnifitermes[198]

Gen. et sp. nov

Jouault et al.

Cretaceous

 Myanmar

A termite. Genus includes new species M. krishnai.

Magniocula paradoxona[199]

Sp. nov

Vršanská & Vršanský in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A cockroach belonging to the family Corydiidae.

Manipulatoides[204]

Gen. et sp. nov

Valid

Li & Huang

Cretaceous (Albian-Cenomanian)

Burmese amber

 Myanmar

A member of the family Manipulatoridae. The type species is M. obscura.

Mastotermes myanmarensis[198]

Sp. nov

Jouault et al.

Cretaceous

 Myanmar

A termite, a species of Mastotermes.

Mesoblattina libanensis[205]

Sp. nov

Valid

Káčerová & Azar

Early Cretaceous

 Lebanon

A cockroach belonging to the family Mesoblattinidae.

Nuurcala coloris[206]

Sp. nov

Valid

Kováčová

Early Cretaceous

Dzun-Bain Formation

 Mongolia

A cockroach belonging to the family Caloblattinidae.

Obscuroblatta[199]

Gen. et sp. nov

Hinkelman & Vršanský in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A cockroach belonging to the family Corydiidae. Genus includes new species O. rigida.

?Ocelloblattula striata[205]

Sp. nov

Valid

Káčerová & Azar

Early Cretaceous

 Lebanon

A cockroach belonging to the group Corydioidea and the family Blattulidae.

Okruhliak[199]

Gen. et sp. nov

Vršanský & Hinkelman in Vršanský et al.

Mesozoic

A cockroach belonging to the family Corydiidae. Genus includes new species O. vedec.

Piloscutumus[199]

Gen. et sp. nov

Sendi in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A cockroach belonging to the family Corydiidae. Genus includes new species P. discretus.

Pokemon[199]

Gen. et sp. nov

Vršanská, Hinkelman & Vršanský in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A cockroach belonging to the family Corydiidae. Genus includes new species P. oculiapertion.

Poroblattina anadiensis[207]

Sp. nov

In press

Correia et al.

Carboniferous (Gzhelian)

Buçaco Carboniferous Basin

 Portugal

A roachoid belonging to the family Poroblattinidae.

Pozabudnutie[208]

Gen. et 2 sp. nov

Valid

Vršanský, Hinkelman & Aristov in Vršanský et al.

Permian to Cretaceous

 Myanmar
 Russia

A phyloblattid cockroach. The type species is P. antiquorum Hinkelman & Vršanský from the Cretaceous Burmese amber; genus also includes P. anakitensis Vršanský & Aristov from the Permian of Anakit, Russia.

Sclerotermes[209]

Gen. et sp. nov

Valid

Jouault & Nam

Early Cretaceous (Albian)

Jinju Formation

 South Korea

A termite. Genus includes new species S. samsiki.

Sosnowiecia[210]

Gen. et sp. nov

Valid

Dvořák, Krzemiński & Prokop

Carboniferous (Westphalian A)

 Poland

A representative of the stem group of Dictyoptera with resemblance to "blattoid" lines Dictyoptera (Blattodea + Mantodea) and Paoliida, possibly a member of the family Mesorthopteridae. The type species is S. dareki.

Squamicaputa[199]

Gen. et sp. nov

Hinkelman & Koubová in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A cockroach belonging to the family Corydiidae. Genus includes new species S. tubulosa.

Supella dominicana[211]

Sp. nov

Valid

Poinar

Miocene

Dominican amber

 Dominican Republic

An ectobiid cockroach, a species of Supella.

Valkyritermes[212]

Gen. et sp. nov

Valid

Jouault, Engel & Nel in Jouault et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A termite belonging to the family Kalotermitidae. The type species is V. inopinatus.

Vitisma okata[206]

Sp. nov

Valid

Kováčová

Early Cretaceous

Dzun-Bain Formation

 Mongolia

A member of the family Umenocoleidae.

Dictyopteran research

  • An assemblage of specimens of Subioblatta madygenica, with several specimens preserving the coloration on wings, is described from the Triassic Madygen Formation (Kyrgyzstan) by Hinkelman (2022).[213]
  • Probable termite coprolites with different proportions are described from the Lower Cretaceous Huolinhe Formation (Huolinhe Basin, China) by Dong et al. (2022), who interpret this finding as indicating that the Early Cretaceous termites from the Huolinhe Basin had wood-feeding habits, and suggesting that the division of labor among termites already existed in the Early Cretaceous.[214]
  • Borings filled with coprolites similar to those produced by extant termites belonging to the family Kalotermitidae are described from silicified conifer woods from the Albian Kachaike Formation (Argentina) by Greppi et al. (2022), who interpret the presence of kalotermitids as indicative of arid climatic conditions.[215]

†Glosselytrodea

Name Novelty Status Authors Age Type locality Location Notes Images

Moscheloptera[216]

Gen. et sp. nov

Valid

Pérez-de la Fuente, Nel & Poschmann

Permian (Asselian–?Sakmarian)

Meisenheim Formation

 Germany

A member of Glosselytrodea belonging to the family Permoberothidae. The type species is M. phantasma.

Hymenopterans

"Symphyta"

Name Novelty Status Authors Age Type locality Location Notes Images

Brachyoxyela conjunctiva[217]

Sp. nov

In press

Dai et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Xyelidae belonging to the subfamily Macroxyelinae and the tribe Macroxyelini.

Brachyoxyela elliptica[217]

Sp. nov

In press

Dai et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Xyelidae belonging to the subfamily Macroxyelinae and the tribe Macroxyelini.

Brachyoxyela leei[217]

Sp. nov

In press

Dai et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Xyelidae belonging to the subfamily Macroxyelinae and the tribe Macroxyelini.

Curvitexis[218]

Gen. et sp. nov

Jouault, Nam & Rasnitsyn

Cretaceous

Burmese amber

 Myanmar

A member of the family Anaxyelidae. Genus includes new species C. kopylovi.

Dolicholyda[219]

Gen. et 3 sp. nov

Valid

Zhuang, Rasnitsyn, Shih & Wang in Zhuang et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Pamphiliidae belonging to the subfamily Juralydinae. The type species is D. obtusata; genus also includes D. angusta and D. confluens.

Emphytus miocenicus[220]

Sp. nov

Valid

Nel et al.

Miocene

 France

An Allantinae Tenthredinid sawfly.

Empria sammuti[220]

Sp. nov

Valid

Nel et al.

Miocene

 France

An Empria species Tenthredinid sawfly.

Eourocerus[221]

Gen. et sp. nov

Valid

Archibald & Rasnitsyn

Early Eocene

Klondike Mountain Formation

 United States
( Washington)

A horntail. The type species is E. anguliterreus.

Eourocerus anguliterreus

Eutomostethus karimae[222]

Sp. nov

Valid

Nel, Niu & Wei

Eocene

Green River Formation

 United States
( Colorado)

A species of Eutomostethus.

Evacuxyela[223]

Gen. et 2 sp. nov

Valid

Dai et al.

Middle Jurassic

Jiulongshan Formation

 China

A member of the family Xyelidae belonging to the subfamily Macroxyelinae and the tribe Angaridyellini. Genus includes new species E. conflata and E. hsiaoae.

Leptoxyela[224]

Gen. et sp. nov

Valid

Dai, Rasnitsyn & Wang in Dai et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Xyelidae belonging to the subfamily Macroxyelinae and the tribe Angaridyelini. The type species is L. eximia.

Paleonematus[225]

Gen. et sp. nov

Valid

Nel

Paleocene

 France

A member of the family Tenthredinidae belonging to the subfamily Nematinae. The type species is P. jarzembowskii.

Paraxiphydria[226]

Gen. et sp. nov

Valid

Gao, Engel & Gao in Gao et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Xiphydriidae[226] or Anaxyelidae.[218] The type species is P. resinata.

Scabolyda latusa[227]

Sp. nov

Valid

Zhuang et al.

Middle Jurassic (Callovian)

Jiulongshan Formation

 China

A member of the family Pamphiliidae.

Scabolyda tenuis[227]

Sp. nov

Valid

Zhuang et al.

Early Cretaceous

Yixian Formation

 China

A member of the family Pamphiliidae.

Scleroxyela cephalota[224]

Sp. nov

Valid

Dai, Rasnitsyn Shih & Wang in Dai et al.

Middle Jurassic (Callovian)

Jiulongshan Formation

 China

A member of the family Xyelidae belonging to the subfamily Macroxyelinae and the tribe Xyeleciini.

"Symphytan" research

  • A study on the phylogenetic affinities of the Oligocene pamphiliid Tapholyda caplani is published by Jouault et al. (2022), who transfer this species to the subfamily Juralydinae.[228]

Apocrita

Apoidea

Name Novelty Status Authors Age Type locality Location Notes Images

Angarosphex ryonsangiensis[229]

Sp. nov

Ri et al.

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

 North Korea

A member of the family Angarosphecidae.

Dactylurina aethiopica[230]

Sp. nov

Valid

Lepeco & Melo

Miocene

Ethiopian amber

 Ethiopia

A stingless bee. Originally described as a species of Dactylurina, but subsequently made the type species of the separate genus Adactylurina.[231]

Exallopterus[232]

Gen. et sp. nov

In press

Rosa, Melo & Ribeiro

Early Cretaceous (Aptian)

Crato Formation

 Brazil

A member of the family Crabronidae. Genus includes new species E. spectabilis.

Heterosphex[233]

Gen. et sp. nov

In press

Li, Rosa, Melo & Shih in Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Cirrosphecidae. Genus includes species is H. wuni.

Liotrigona aethiopica[234]

Sp. nov

Valid

Engel in Engel & Aber

Miocene

Ethiopian amber

 Ethiopia

A stingless bee.

Pompilopterus sinuijuensis[229]

Sp. nov

Ri et al.

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

 North Korea

A member of the family Angarosphecidae.

Chalcidoidea

Name Novelty Status Authors Age Type locality Location Notes Images

Archaeocercoides[235]

Gen. et sp. nov

Valid

Simutnik in Simutnik et al.

Eocene

Rovno amber

 Ukraine

A member of the family Encyrtidae. The type species is A. puchkovi.

Balticalcarus[236]

Gen. et sp. nov

Valid

Simutnik in Simutnik, Perkovsky & Vasilenko

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A member of the family Encyrtidae belonging to the subfamily Tetracneminae. The type species is B. archibaldi.

Electronoyesella[237]

Gen. et sp. nov

Valid

Simutnik in Simutnik, Perkovsky & Vasilenko

Eocene

Rovno amber

 Ukraine

A member of the family Encyrtidae belonging to the subfamily Tetracneminae. The type species is E. antiqua.

Protaphycus[238]

Gen. et sp. nov

Valid

Simutnik in Simutnik, Perkovsky & Vasilenko

Eocene

Rovno amber

 Ukraine

A member of the family Encyrtidae belonging to the subfamily Encyrtinae. The type species is P. shuvalikovi.

Rovnopositor[235]

Gen. et sp. nov

Valid

Simutnik in Simutnik et al.

Eocene

Rovno amber

 Ukraine

A member of the family Encyrtidae. The type species is R. voblenkoi.

Chrysidoidea

Name Novelty Status Authors Age Type locality Location Notes Images

Acanthabythus[239]

Gen. et sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

 Myanmar

A member of the family Scolebythidae. Genus includes new species A. exilispineus.

Ajkanesia[240]

Gen. et sp. nov

In press

Szabó & Brazidec in Szabó et al.

Late Cretaceous (Santonian)

Ajka Coal Formation

 Hungary

A member of the family Bethylidae belonging to the subfamily Pristocerinae. The type species is A. harmincipsziloni.

Alongatepyris pedrocai[241]

Sp. nov

Valid

Colombo & Azevedo in Colombo, Perkovsky & Azevedo

Miocene

Dominican amber

 Dominican Republic

A member of the family Bethylidae.

Amissidigitus[240]

Gen. et sp. nov

In press

Szabó & Brazidec in Szabó et al.

Late Cretaceous (Santonian)

Ajka Coal Formation

 Hungary

A member of the family Bethylidae belonging to the subfamily Scleroderminae. The type species is A. belai.

Arcapenesia[239]

Gen. et sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

 Myanmar

A member of the family Scolebythidae. Genus includes new species A. compacta.

Bakeriella nanyhelae[242]

Sp. nov

In press

Brazidec & Perrichot

Miocene

Mexican amber

 Mexico

A member of the family Bethylidae belonging to the subfamily Epyrinae.

Bocchus rex[243]

Sp. nov

Valid

Olmi et al.

Eocene

Rovno amber

 Ukraine

A member of the family Dryinidae.

Burmapenesia[239]

Gen. et 2 sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

 Myanmar

A member of the family Scolebythidae. Genus includes new species B. iridescens and B. longicephala.

Cephalobythus[239]

Gen. et 4 sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

 Myanmar

A member of the family Scolebythidae. Genus includes new species C. clypeatus, C. deformis, C. gladiator and C. simplex.

Cretopsenella[239]

Gen. et sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

 Myanmar

A member of the family Scolebythidae. Genus includes new species C. spinifera.

Dryinus carsteni[244]

Sp. nov

Valid

Olmi et al.

Cretaceous

Burmese amber

 Myanmar

A species of Dryinus.

Gnathapenesia[239]

Gen. et sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

 Myanmar

A member of the family Scolebythidae. Genus includes new species G. burmensis.

Goniozus cotyi[242]

Sp. nov

In press

Brazidec & Perrichot

Miocene

Mexican amber

 Mexico

A species of Goniozus.

Hedychridium rosai[245]

Sp. nov

Brazidec & Perrichot

Miocene

Zhangpu amber

 China

A species of Hedychridium.

Parascleroderma palaeosinica[246]

Sp. nov

In press

Brazidec & Perrichot

Miocene

Zhangpu amber

 China

A member of the family Bethylidae.

Rhynchopsenella[239]

Gen. et 3 sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

 Myanmar

A member of the family Scolebythidae. Genus includes new species R. argentea, R. bicarinata and R. diaphana.

Rovnodryinus[247]

Gen. et sp. nov

Valid

Olmi et al.

Eocene

Rovno amber

 Ukraine

A member of the family Dryinidae belonging to the subfamily Apodryininae. Genus includes new species R. khomychi.

Sierola colomboi[245]

Sp. nov

Brazidec & Perrichot

Miocene

Zhangpu amber

 China

A member of the family Bethylidae.

Solepyris electromexicanus[242]

Sp. nov

In press

Brazidec & Perrichot

Miocene

Mexican amber

 Mexico

A member of the family Bethylidae belonging to the subfamily Scleroderminae.

Voronknoxus[248]

Gen. et sp. nov

Valid

Colombo & Perkovsky in Colombo, Perkovsky & Vasilenko

Eocene

Rovno amber

 Ukraine

A Scleroderminae Bethylid wasp.
The type species is V. zerovae.

Diaprioidea

Name Novelty Status Authors Age Type locality Location Notes Images

Basalys villumi[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A member of the family Diapriidae belonging to the subfamily Diapriinae.

Belyta knudhoejgaardi[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A species of Belyta.

Cinetus breviscapus[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A member of the family Diapriidae belonging to the subfamily Belytinae.

Cinetus elongatus[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A member of the family Diapriidae belonging to the subfamily Belytinae.

Doliopria baltica[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A member of the family Diapriidae belonging to the subfamily Diapriinae.

Pantoclis globosa[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A member of the family Diapriidae belonging to the subfamily Belytinae.

Pantolyta augustinusii[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A member of the family Diapriidae belonging to the subfamily Belytinae.

Pantolyta chemyrevae[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A member of the family Diapriidae belonging to the subfamily Belytinae.

Pantolyta similis[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A member of the family Diapriidae belonging to the subfamily Belytinae.

Spathiopteryx soosi[240]

Sp. nov

In press

Szabó, Brazidec & Perrichot in Szabó et al.

Late Cretaceous (Santonian)

Ajka Coal Formation

 Hungary

A member of the family Spathiopterygidae.

Spilomicrus succinalis[249]

Sp. nov

Valid

Brazidec & Vilhelmsen

Eocene

Blue Earth Formation
(Baltic amber)

Europe (Baltic Sea region)

A species of Spilomicrus.

Evanioidea

Name Novelty Status Authors Age Type locality Location Notes Images

Nevania deviata[250]

Sp. nov

Li et al.

Middle Jurassic

Haifanggou Formation

 China

A member of the family Praeaulacidae.

Praeaulacon grossus[250]

Sp. nov

Li et al.

Middle Jurassic

Haifanggou Formation

 China

A member of the family Praeaulacidae.

Pristaulacus jarzembowskii[251]

Sp. nov

Valid

Jouault & Nel

Paleocene

 France

A species of Pristaulacus.

Formicoidea

Name Novelty Status Authors Age Type locality Location Notes Images

Camponotus parvus[252]

Sp. nov

Valid

Perfilieva

Oligocene

 Russia

A carpenter ant.

Desyopone[253]

Gen. et sp. nov

Valid

Boudinot & Perrichot in Boudinot et al.

Miocene

Ethiopian amber

 Ethiopia

An ant belonging to the subfamily Ponerinae and the tribe Ponerini. The type species is D. hereon.

Emplastus amguensis[252]

Sp. nov

Valid

Perfilieva

Oligocene

 Russia

An ant, a species of Emplastus.

Emplastus obliquus[252]

Sp. nov

Valid

Perfilieva

Oligocene

 Russia

An ant, a species of Emplastus.

Emplastus similis[252]

Sp. nov

Valid

Perfilieva

Oligocene

 Russia

An ant, a species of Emplastus.

Gerontoformica sternorhabda[254]

Sp. nov

Valid

Boudinot et al.

Cretaceous

Burmese amber

 Myanmar

An ant, a member of the subfamily Sphecomyrminae.

Lasius vulgaris[252]

Sp. nov

Valid

Perfilieva

Oligocene

 Russia

An ant, a species of Lasius.

Manica andrannae[255]

Sp. nov

Zharkov & Dubovikoff in Zharkov, Dubovikoff & Abakumov

Eocene (Priabonian)

Prussian Formation
(Baltic amber)

 Russia
( Kaliningrad Oblast)

A species of Manica.

Odontoponera pseudotransversa[256]

Sp. nov

Perrichot

Miocene

Zhangpu amber

 China

A species of Odontoponera.

Paraphaenogaster ussuriensis[252]

Sp. nov

Valid

Perfilieva

Oligocene

 Russia

An ant belonging to the subfamily Myrmicinae.

Ravavy goldmani[257]

Sp. nov

Boudinot & Perrichot in Perrichot et al.

Miocene

Ethiopian amber

 Ethiopia

An ant, a species of Ravavy.

Schismiscapus[258]

Gen. et sp. nov

In press

Varela-Hernández et al.

Miocene

Mexican amber

 Mexico

An ant belonging to the subfamily Dolichoderinae. The type species is S. exstinctum.

Sussudio[259]

Gen. et comb. nov

Valid

Boudinot, Borowiec & Prebus

Eocene

Baltic amber

Europe (Baltic Sea region)

A Formicinae ant of uncertain affinities.
A new genus for "Pseudolasius" boreus Wheeler (1915).

Technomyrmex kemaensis[252]

Sp. nov

Valid

Perfilieva

Oligocene

 Russia

An ant, a species of Technomyrmex.

Technomyrmex svojtkai[257]

Sp. nov

Perrichot & Engel in Perrichot et al.

Miocene

Ethiopian amber

 Ethiopia

An ant, a species of Technomyrmex.

Ichneumonoidea

Name Novelty Status Authors Age Type locality Location Notes Images

Acerataspis? revelata[260]

Comb nov

(Brues)

Valid

Eocene
Chadronian

Florissant Formation

 USA
 Colorado

A metopiine Darwin wasp.
Moved from Pimpla revelata (1910)

Armadilleon[260]

Gen. et 2 comb. nov

Valid

Spasojevic, Broad & Klopfstein

Eocene
Chadronian

Florissant Formation

 United States
( Colorado)

A phygadeuontine Darwin wasp.
The type species is "Pimpla" morticina Brues (1910)
Additionally includes "Polysphincta" petrorum Brues (1910).

Crusopimpla collina[261]

Sp. nov

Valid

Klopfstein

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae belonging to the subfamily Pimplinae.

Crusopimpla? elongata[261]

Sp. nov

Valid

Klopfstein

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae belonging to the subfamily Pimplinae.

Crusopimpla minuta[261]

Sp. nov

Valid

Klopfstein

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae belonging to the subfamily Pimplinae.

Crusopimpla rettigi[261]

Sp. nov

Valid

Klopfstein

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae belonging to the subfamily Pimplinae.

Crusopimpla violina[261]

Sp. nov

Valid

Klopfstein

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae belonging to the subfamily Pimplinae.

Crusopimpla weltii[262]

Sp. nov

Valid

Viertler, Spasojevic & Klopfstein in Viertler et al.

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae belonging to the subfamily Pimplinae.

Dimophora? longicornis[260]

Comb nov

(Theobald)

Valid

Oligocene
Chattian

Aix-en-Provence Formation

 France
 Provence-Alpes-Côte d'Azur

A cremastine Darwin wasp.
Moved from Nemeritis longicornis (1937)

Dimophora? wickhami[260]

Comb nov

(Cockerell)

Valid

Eocene
Chadronian

Florissant Formation

 USA
 Colorado

A cremastine Darwin wasp.
Moved from Theronia wickhami (1919)

Ebriosa[262]

Gen. et sp. nov

Valid

Viertler, Spasojevic & Klopfstein in Viertler et al.

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae, possibly belonging to the subfamily Ctenopelmatinae. The type species is E. flava.

Entypoma? duergari[262]

Sp. nov

Valid

Viertler, Spasojevic & Klopfstein in Viertler et al.

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae belonging to the subfamily Orthocentrinae, possibly a species of Entypoma.

Hallocinetus? arvernus[260]

Comb nov

(Piton)

Valid

Paleocene
Thanetian

Menat Formation

 France
 Puy-de-Dôme

A possible Acaenitine Darwin wasp.
Moved from Phaenolobus arvernus (1940)

Hypsicera? solidata[260]

Comb nov

(Brues)

Valid

Eocene
Chadronian

Florissant Formation

 USA
 Colorado

A metopiine Darwin wasp.
Moved from Camerotops solidatus (1910)

Lathrolestes? zlatorog[262]

Sp. nov

Valid

Viertler, Spasojevic & Klopfstein in Viertler et al.

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae, possibly belonging to the subfamily Ctenopelmatinae and to the genus Lathrolestes.

Lithoserix antiquus[260]

Comb nov

(Saussure)

Valid

Oligocene
Chattian

Aix-en-Provence Formation

 France
 Provence-Alpes-Côte d'Azur

A cremastine Darwin wasp.
Moved from Pimpla antiqua (1852)

Lycorina? indura[260]

Comb nov

(Theobald)

Valid

Oligocene
Rupelian

Kleinkembs Formation

 Germany
 Baden[263]

A possible lycorinine Darwin wasp.
Moved from Pimpla indura (1937)
Listed as Kleinkembs, Pays de Bade, Haut-Rhin, France by Spasojevic, Broad & Klopfstein 2022[260]

Madma[264]

Gen. et sp. nov

Valid

Viertler et al.

Eocene

Oise amber

 France

A member of the family Ichneumonidae belonging to the subfamily Phygadeuontinae. The type species is M. oisella.

Monoblastus? senilis[260]

Comb nov

(Brues)

Valid

Eocene
Chadronian

Florissant Formation

 USA
 Colorado

A tryphonine Darwin wasp.
Moved from Pimpla senilis (1910)

Orthocentrus? mortuaria[260]

Comb nov

(Brues)

Valid

Eocene
Chadronian

Florissant Formation

 USA
 Colorado

An orthocentrine Darwin wasp.
Moved from Polysphincta mortuaria (1910)

Palaeorionis[265]

Gen. et sp. nov

Valid

Belokobylskij & Hovorka

Miocene

Dominican amber

 Dominican Republic

A member of the family Braconidae belonging to the subfamily Euphorinae. The type species is P. longicaudis.

Pappous[264]

Gen. et sp. nov

Valid

Viertler et al.

Eocene

Oise amber

 France

A member of the family Ichneumonidae, possibly belonging to the subfamily Tryphoninae. The type species is P. trichomatius.

Protoctonus[266]

Gen. et sp. nov

Valid

Davidian in Davidian, Manukyan & Belokobylskij

Eocene

Prussian Formation
(Baltic amber)

 Russia
( Kaliningrad Oblast)

A member of the family Braconidae belonging to the subfamily Aphidiinae. The type species is P. groehni.

Theronia? furensis[261]

Sp. nov

Valid

Klopfstein

Eocene (Ypresian)

Fur Formation

 Denmark

Possibly a species of Theronia.

Theronia? nigriscutum[261]

Sp. nov

Valid

Klopfstein

Eocene (Ypresian)

Fur Formation

 Denmark

Possibly a species of Theronia.

Triclistus bibori[262]

Sp. nov

Valid

Viertler, Spasojevic & Klopfstein in Viertler et al.

Eocene (Ypresian)

Fur Formation

 Denmark

A member of the family Ichneumonidae belonging to the subfamily Metopiinae, a species of Triclistus.

Utrillabracon[267]

Gen. et sp. nov

Valid

Álvarez-Parra & Engel in Álvarez-Parra et al.

Early Cretaceous (Albian)

Escucha Formation

 Spain

A member of the family Braconidae belonging to the subfamily Protorhyssalinae. The type species is U. electropteron.

Xanthopimpla ciboisae[261]

Sp. nov

Valid

Klopfstein

Eocene (Ypresian)

Fur Formation

 Denmark

A species of Xanthopimpla.

Xanthopimpla crescendae[261]

Sp. nov

Valid

Klopfstein

Eocene (Ypresian)

Fur Formation

 Denmark

A species of Xanthopimpla.

Xorides sejugatus[260]

Comb nov

(Brues)

Valid

Eocene
Chadronian

Florissant Formation

 USA
 Colorado

An orthocentrine Darwin wasp.
Moved from Xylonomus sejugatus (1910)

Zagryphus tilloyi[260]

Comb nov

(Theobald)

Valid

Oligocene
Chattian

Aix-en-Provence Formation

 France
 Provence-Alpes-Côte d'Azur

A tryphonine Darwin wasp.
Moved from Promethes tilloyi (1937)

Mymarommatoidea

Name Novelty Status Authors Age Type locality Location Notes Images

Cretaceomma[268]

Gen. et sp. nov

Valid

Rasnitsyn et al.

Early Cretaceous

Lebanese amber

 Lebanon

A member of the family Gallorommatidae. The type species is C. libanensis.

Proctotrupoidea

Name Novelty Status Authors Age Type locality Location Notes Images

Astarteserphus[269]

Gen. et sp. nov

Valid

Engel, Herhold & Barden

Early Cretaceous

Lebanese amber

 Lebanon

A member of the family Proctotrupidae. Genus includes new species A. grimaldii.

Cresogmus[270]

Gen. et sp. nov

Valid

Rasnitsyn & Kolyada in Rasnitsyn et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Proctotrupidae. Genus includes new species C. grimaldii.

Paekthohelorus[229]

Gen. et sp. nov

Ri et al.

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

 North Korea

A member of the family Heloridae. Genus includes new species P. sinuijuensis.

Serphitoidea

Name Novelty Status Authors Age Type locality Location Notes Images

Leptoserphites[268]

Gen. et 2 sp. nov

Valid

Rasnitsyn et al.

Early Cretaceous

Lebanese amber

 Lebanon

A member of the family Serphitidae. Genus includes new species L. pabloi and L. iriae.

Microserphites libanensis[268]

Sp. nov

Valid

Rasnitsyn et al.

Early Cretaceous

Lebanese amber

 Lebanon

A member of the family Serphitidae.

Stephanoidea

Name Novelty Status Authors Age Type locality Location Notes Images

Myanmarina diversa[271]

Sp. nov

In press

Zheng et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Myanmarinidae.

Proapocritus habitus[250]

Sp. nov

Li et al.

Middle Jurassic

Haifanggou Formation

 China

A member of the family Ephialtitidae.

Tiphioidea

Name Novelty Status Authors Age Type locality Location Notes Images

Burmatiphia[272]

Gen. et sp. nov

In press

Zheng et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Tiphiidae. Genus includes new species B. mandibulata.

Other Apocrita

Name Novelty Status Authors Age Type locality Location Notes Images

Prionaspidion[273]

Gen. et 2 sp. nov

Valid

Lepeco & Melo in Lepeco, Barbosa & Melo

Cretaceous

Burmese amber

 Myanmar

A member of Aculeata belonging to the family Trifionychidae. The type species is P. brevidens; genus also includes P. nanus.

Trifionyx[273]

Fam. et gen. et sp. nov

Valid

Lepeco & Melo in Lepeco, Barbosa & Melo

Cretaceous

Burmese amber

 Myanmar

A member of Aculeata; the type genus of the new family Trifionychidae (possibly belonging to the superfamily Bethylonymoidea). The type species is T. pilosus.

Trifionyximus[273]

Gen. et sp. nov

Valid

Lepeco & Melo in Lepeco, Barbosa & Melo

Cretaceous

Burmese amber

 Myanmar

A member of Aculeata belonging to the family Trifionychidae. The type species is T. cracens.

Apocritan research

  • A bethylid specimen belonging to the genus Eupsenella (currently confined to Australia and New Zealand and with the fossil record previously only known from the Old World) is described from the Eocene Green River Formation by Brazidec & Perrichot (2022).[274]
  • Richter et al. (2022) describe almost completely preserved internal head structures of a specimen of Gerontoformica gracilis from the Cretaceous amber from Myanmar.[275]
  • Zhuang et al. (2022) describe a female specimen of Zigrasimecia with exceptionally preserved soft tissues from the Cretaceous amber from Myanmar.[276]

Clade Neuropterida

Megaloptera

Megalopteran research

Neuroptera

Name Novelty Status Authors Age Type locality Location Notes Images

Acanthochrysa[278]

Gen. et sp. nov

In press

Liu et al.

Cretaceous

Burmese amber

 Myanmar

A member of Chrysopoidea. Genus includes new species A. langae.

Aggregataberotha[279]

Gen. et sp. nov

Wang, Huang & Wang in Wang et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Berothidae. Genus includes new species A. punctata.

Araripenymphpes burmanus[280]

Sp. nov

Valid

Lu, Xu & Liu

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of Myrmeleontoidea belonging to the family Cratosmylidae.

Arbusella platyptera[281]

Sp. nov

Ma & Wang in Ma et al.

Middle Jurassic (Aalenian/Bajocian boundary)

Jiulongshan Formation

 China

A member of the family Osmylidae belonging to the subfamily Kempyninae.

Archaeochrysa cockerelli[282]

Sp. nov

Valid

Makarkin, Antell & Archibald

Eocene

Florissant Formation

 United States
( Colorado)

A member of the family Chrysopidae belonging to the subfamily Nothochrysinae.

Archaeoconis[283]

Gen. et sp. nov

Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Coniopterygidae belonging to the subfamily Cretaconiopteryginae. Genus includes new species A. zhangzhiqiae.

Archaeomegalomus[284]

Gen. et sp. nov

Valid

Nakamine, Yamamoto & Takahashi

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of the family Hemerobiidae. The type species is A. burmiticus.

Burmotachinymphes pengi[285]

Sp. nov

Valid

Chen, Deng & Yang in Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Mesochrysopidae.

Cimbrochrysa americana[282]

Sp. nov

Valid

Makarkin, Antell & Archibald

Eocene

Florissant Formation

 United States
( Colorado)

A member of the family Chrysopidae belonging to the subfamily Nothochrysinae.

Cimbrochrysa major[282]

Sp. nov

Valid

Makarkin, Antell & Archibald

Eocene

Florissant Formation

 United States
( Colorado)

A member of the family Chrysopidae belonging to the subfamily Nothochrysinae.

Coronidilar[286]

Gen. et sp. nov

Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Dilaridae. Genus includes new species C. yanae.

Cretaconiopteryx zhengpengyueae[283]

Sp. nov

Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Coniopterygidae belonging to the subfamily Cretaconiopteryginae.

Cretapsychops skywalkeri[287]

Sp. nov

Valid

Khramov & Chemakos

Late Jurassic

Karabastau Formation

 Kazakhstan

A member of the family Psychopsidae.

Cretodilar multinervius[286]

Sp. nov

Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Dilaridae.

Cretoneuronema[288]

Gen. et sp. nov

Valid

Liu, Chen & Zhuo

Cretaceous

Burmese amber

 Myanmar

A member of the family Hemerobiidae. The type species is C. jarzembowskii.

Doratomantispa arcimaculata[289]

Sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Mantispidae.

Doratomantispa gaoyuhei[289]

Sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Mantispidae.

Doratomantispa pouilloni[290]

Sp. nov

Valid

Jouault & Nel in Jouault, Pouillon & Nel

Cretaceous (Albian-Cenomanian)

Burmese amber

 Myanmar

A member of the family Mantispidae.

Doratomantispa yumeiyingae[289]

Sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Mantispidae.

Doratomantispa zhangwenjuni[289]

Sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Mantispidae.

Doratomantispa zhangzhiqiae[289]

Sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Mantispidae.

Doratomantispa zhuozhengmingi[289]

Sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Mantispidae.

Electrobabinskaia neli[291]

Sp. nov

Valid

Jouault

Cretaceous

Burmese amber

 Myanmar

A member of the family Babinskaiidae.

Heteroconis zhangpuensis[292]

Sp. nov

In press

Chen, Li & Liu

Miocene

Zhangpu amber

 China

A species of Heteroconis.

Jersiberotha musivum[293]

Sp. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Berothidae.

Kuafupolydentes[294]

Gen. et sp. nov

In press

Luo, Liu & Jarzembowski

Cretaceous

Burmese amber

 Myanmar

An early member of Myrmeleontiformia. The type species is K. hui.

Lebanosmylus[295]

Gen. et sp. nov

Valid

Azar & Nel

Late Cretaceous (Cenomanian)

 Lebanon

A member of the family Saucrosmylidae. The type species is L. leae.

Lithochrysa meyeri[282]

Sp. nov

Valid

Makarkin, Antell & Archibald

Eocene

Florissant Formation

 United States
( Colorado)

A member of the family Chrysopidae belonging to the subfamily Nothochrysinae.

Mantispa? damzenogedanica[296]

Sp. nov

Valid

Baranov et al.

Eocene (BartonianPriabonian)

Baltic amber

 Russia
( Kaliningrad Oblast)

Possibly a species of Mantispa.

Mesypochrysa coadnata[297]

Sp. nov

In press

Chen et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Chrysopidae belonging to the subfamily Limaiinae.

Microsisyra[298]

Gen. et sp. nov

Makarkin

Cretaceous

Burmese amber

 Myanmar

A member of the family Sisyridae. Genus includes new species M. carsteni.

Olindanymphes? headsi[299]

Sp. nov

Valid

Makarkin

Early Cretaceous (Aptian)

Crato Formation

 Brazil

A member of the family Nymphidae.

Paradoxoconis[300]

Gen. et 2 sp. nov

Valid

Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Coniopterygidae. The type species is P. longipalpa; genus also includes P. szirakii.

Paradoxoleon[280]

Gen. et sp. nov

Valid

Lu, Xu & Liu

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Babinskaiidae. The type species is P. chenruii.

Paradoxomantispa mahaiyingae[289]

Sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Mantispidae.

Sinuijuala[301]

Gen. et sp. nov

In press

So & Won

Early Cretaceous (Barremian-Aptian)

Sinuiju Formation

 North Korea

A member of the family Ithonidae. The type species is S. paekthoensis.

Sinuijumantispa[302]

Gen. et sp. nov

In press

So & Won

Early Cretaceous (Barremian-Aptian)

Sinuiju Formation

 North Korea

A member of the family Mantispidae. The type species is S. ryonsangiensis.

Xiaoberotha dilatapoda[279]

Sp. nov

Wang, Huang & Wang in Wang et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Berothidae.

Xiaoleon[303]

Gen. et sp. nov

In press

Lu & Liu

Cretaceous

Burmese amber

 Myanmar

An antlion. Genus includes new species X. amoena.

Neuropteran research

  • A study on the fossil record of aphidlions and aphidlion-like lacewing larvae, including reports of new specimens from the Cretaceous Burmese amber and Eocene Baltic amber and a study on the morphological diversity of the heads of the fossil larvae, is published by Haug et al. (2022).[304]
  • Haug et al. (2022) describe an aphidlion-like larva from the Cretaceous Burmese amber, preserved in close proximity to an empty egg case, and interpreted by the authors as likely representing the first newly hatched aphidlion-like larva reported from the Burmese amber.[305]
  • A hemerobiid aphidlion preserved with possible Cf. Germaraphis prey, representing the first possible case of a cooccurrence of predator and prey for lacewing larvae reported to date, is described from a piece of the Eocene Baltic amber by Haug et al. (2022).[306]
  • An overview of the diversity and fossil record of the larvae of the families Ithonidae, Sisyridae and Coniopterygidae is published by Haug et al. (2022).[307]
  • Two probable berothid larvae with extremely enlarged trunks are described from a single piece of the Cretaceous Kachin amber from Myanmar by Haug & Haug (2022), representing the oldest case of the extreme inflation of the trunk in insects reported to date.[308]

Raphidioptera

Name Novelty Status Authors Age Type locality Location Notes Images

Siboptera lata[309]

Sp. nov

Zhang et al.

Early Cretaceous (Aptian)

Baiwan Formation

 China

A member of the family Mesoraphidiidae.

Raphidiopteran research

  • A study on the morphological diversity of snakefly larvae over the last 100 million years is published by Haug et al. (2022), who also describe new specimens of larval snakeflies preserved in Eocene and Cretaceous ambers.[310]
  • Makarkin et al. (2022) describe snakefly larvae from the Eocene Sakhalinian amber (the first confirmed representative of Raphidioptera from the Cenozoic of Asia) and Rovno amber (representing the first European Cenozoic immature snakefly found outside of Russo-Scandia).[311]

Clade †Palaeodictyopteroidea

†Megasecoptera

Name Novelty Status Authors Age Type locality Location Notes Images

Corydaloides leonensis[82]

Sp. nov

Valid

Santos et al.

Late Carboniferous

 Spain

A member of the family Corydaloididae. Announced in 2022; the final article version was published in 2023.

Mischoptera bergidensis[82]

Sp. nov

Valid

Santos et al.

Late Carboniferous

 Spain

A member of the family Mischopteridae. Announced in 2022; the final article version was published in 2023.

Piesbergptera[312]

Fam. et gen. et sp. nov

In press

Rosová et al.

Carboniferous (Moscovian)

 Germany

A megasecopteran
The type genus of the new family Piesbergpteridae.
Genus includes new species P. niesseae.

†Palaeodictyoptera

Name Novelty Status Authors Age Type locality Location Notes Images

Osnabrugapteron[312]

Gen. et sp. nov

In press

Rosová et al.

Carboniferous (Moscovian)

 Germany

A palaeodictyopteran belonging to the family Archaemegaptilidae. Genus includes new species O. rasnitsyni.

Saxonyptilus[312]

Gen. et sp. nov

In press

Rosová et al.

Carboniferous (Moscovian)

 Germany

A palaeodictyopteran belonging to the family Spilapteridae. Genus includes new species S. sowiaki.

Spilaptera peckae[312]

Sp. nov

In press

Rosová et al.

Carboniferous (Moscovian)

 Germany

A palaeodictyopteran belonging to the family Spilapteridae.

Palaeodictyopteran research

  • A study on the abdominal lateral outgrowths (flaps) of Paleozoic palaeodictyopteran larvae is published by Prokop et al. (2022), who report that these flaps show comparable structure to thoracic wings, and most likely represent wing serial homologues.[313]

Clade Palaeoptera

Ephemeroptera

Name Novelty Status Authors Age Type locality Location Notes Images

Balticobaetisca bispinata[314]

Sp. nov

Valid

Staniczek, Storari & Godunko

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A mayfly belonging to the family Baetiscidae.

Calliarcys antiquus[315]

Sp. nov

Valid

Godunko, Alba-Tercedor & Staniczek in Godunko et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Leptophlebiidae.

Cloeon tzeltal[316]

Sp. nov

Valid

Varela-Hernández, Riquelme & Montiel-Chávez

Miocene

Mexican amber

 Mexico

A species of Cloeon.

Jurassephemera[317]

Gen. et sp. nov

In press

Zhang et al.

Early Jurassic

Shiti Formation

 China

A mayfly belonging to the family Sharephemeridae. The type species is J. zhangi.

Kachinophlebia[318]

Gen. et sp. nov

In press

Chen & Zheng

Cretaceous

Burmese amber

 Myanmar

A member of the family Leptophlebiidae. Genus includes new species K. zhouchangfai.

Odonatoptera

Name Novelty Status Authors Age Type locality Location Notes Images

Aeshna caseneuvensis[319]

Sp. nov

Nel et al.

Oligocene

 France

A dragongly, a species of Aeshna.

Afrozygopteron[320]

Gen. et sp. nov

In press

Nel, Garrouste & Prevec

Permian (Guadalupian)

 South Africa

A protozygopteran belonging to the family Luiseiidae. The type species is A. inexpectatus.

Allenbya[321]

Gen. et sp. nov

Valid

Archibald & Cannings

Eocene (Ypresian)

Allenby Formation

 Canada
( British Columbia)

A possible Dysagrionidae.
The type species is A. holmesae.
Not to be confused with the Princeton Chert Allenbya (plant) of the same formation.

Allenbya holmesae

Araripegomphus ismari[322]

Sp. nov

Nel & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A member of the family Araripegomphidae.

Cratoaeschnidium[323]

Gen. et sp. nov

In press

Nel, Jouault & Ribeiro

Early Cretaceous

Crato Formation

 Brazil

A dragonfly belonging to the family Aeschnidiidae. The type species is C. martinsnetoi.

Danowhetaksa[324]

Gen. et 2 sp. nov

Valid

Simonsen et al.

Eocene (Ypresian)

Ølst Formation

 Denmark

A whetwhetaksid odonate.
Genus includes new species D. birgitteae and D. rusti.

Gansuaeschnidia[325]

Gen. et sp. nov

Valid

Zhang et al.

Early Cretaceous

Chijinpu Formation

 China

A dragonfly belonging to the family Aeschnidiidae. The type species is G. hongi.

Guangxicalopteryx[326]

Gen. et sp. nov

Valid

Huang et al.

Oligocene

Ningming Formation

 China

A damselfly belonging to the family Calopterygidae. The type species is G. huashanensis.

Kennedya ferganensis[327]

Sp. nov

Valid

Felker

Triassic

Madygen Formation

 Kyrgyzstan

A Kennedyidae Archizygopteran.

Kennedya madygensis[327]

Sp. nov

Valid

Felker

Triassic

Madygen Formation

 Kyrgyzstan

A Kennedyidae Archizygopteran

Kishenehna[328]

Gen. et sp. nov

Valid

Archibald, Cannings & Greenwalt

Eocene (Lutetian)

Kishenehn Formation

 United States
( Montana)

A Gomphaeschninae aeshnid dragonfly.
The type species is K. prima.

Koreaphlebia[329]

Gen. et sp. nov

Valid

Jouault, Nam & Nel

Triassic

Amisan Formation

 South Korea

A member of Triadophlebiomorpha belonging to the family Zygophlebiidae. Genus includes new species K. iussradiae.

Koreatriassothemis[330]

Gen. et sp. nov

Valid

Nel, Nam & Jouault

Late Triassic

Amisan Formation

 South Korea

A member of Parazygoptera belonging to the superfamily Triassolestoidea. The type species is K. elongatus.

Labandeiraia grimaldii[331]

Sp. nov

Valid

Nel

Eocene

Oise amber

 France

A member of the family Epallagidae.

Macrogomphus menatensis[332]

Sp. nov

Valid

Nel & Jouault

Paleocene

 France

A dragonfly, a species of Macrogomphus.

Menatagrion[332]

Gen. et sp. nov

Valid

Nel & Jouault

Paleocene

 France

A member of the family Dysagrionidae. The type species is M. hervetae.

Menatlestes[332]

Fam. et gen. et sp. nov

Valid

Nel & Jouault

Paleocene

 France

A damselfly belonging to the group Lestinoidea and to the new family Menatlestidae. The type species is M. palaeocenicus.

Oligaeschna sinica[333]

Sp. nov

Valid

Huang et al.

Eocene (Bartonian)

Niubao Formation

 China

An aeshnid dragonfly.

Parapetala yixianensis[334]

Sp. nov

In press

Zheng et al.

Early Cretaceous

Yixian Formation

 China

A dragonfly belonging to the group Aeshnoptera.

Pseudoliupanshania[334]

Gen. et sp. nov

In press

Zheng et al.

Early Cretaceous

Yixian Formation

 China

A dragonfly belonging to the group Aeshnoptera. Genus includes new species P. magnanicellula.

Rudiaeschna jarzembowskii[335]

Sp. nov

Valid

Huang, Jouault & Nel

Early Cretaceous

Yixian Formation

 China

A member of the family Rudiaeschnidae.

Sinomesuropetala[336]

Gen. et sp. nov

In press

Boudet, Nel & Huang

Late Jurassic (Oxfordian)

Haifanggou Formation

 China

A dragonfly belonging to the family Mesuropetalidae. The type species is S. daohugensis.

Turanophlebia liaoningensis[337]

Sp. nov

Fang & Zheng

Early Cretaceous

Yixian Formation

 China

A member of the family Tarsophlebiidae.

Viridiflumineagrion[338]

Gen. et sp. nov

Valid

Nel

Early Eocene

Green River Formation

 United States

A damselfy. Genus includes new species V. aasei.

Odonatopteran research

  • A study on the diversity of color patterns among Early Jurassic heterophlebiids is published by Jouault et al. (2022), who identify five different patterns of coloration in the studied fossil material, and interpret the appearance of patterns of colored spots and bands on the studied heterophlebiid wings as likely caused by the increase of the predation pressure by pterosaurs.[339]

Clade †Paoliidea

†Paoliida

Name Novelty Status Authors Age Type locality Location Notes Images

Glaphyrophlebia glinka[340]

Sp. nov

Valid

Aristov, Rasnitsyn & Naugolnykh

Permian

 Russia
( Perm Krai)

Glaphyrophlebia kamaensis[341]

Sp. nov

Valid

Aristov & Rasnitsyn

Permian

 Russia

A Blattinopsidae paoliid.

Glaphyrophlebia popasnaya[342]

Sp. nov

Valid

Aristov & Rasnitsyn

Carboniferous (Kasimovian)

Upper Isaeva Formation

 Ukraine

A blattinopsid.

Glaphyrophlebia rara[341]

Sp. nov

Valid

Aristov & Rasnitsyn

Permian

 Russia

A member of the family Blattinopsidae.

Glaphyrophlebia semipenna[340]

Sp. nov

Valid

Aristov, Rasnitsyn & Naugolnykh

Permian

 Russia
( Perm Krai)

Glaphyrophlebia vaga[341]

Sp. nov

Valid

Aristov & Rasnitsyn

Permian

 Russia

A member of the family Blattinopsidae.

Glaphyrophlebia victoiriensis[343]

Sp. nov

Valid

Nel, Garrouste & Jouault in Nel et al.

Carboniferous (Gzhelian)

 France

A member of the family Blattinopsidae.

Simplexpaolia[82]

Gen. et sp. nov

Valid

Santos et al.

Late Carboniferous

 Spain

A member of the family Paoliidae. Genus includes new species S. prokopi. Announced in 2022; the final article version was published in 2023.

Stephanopsis testai[210]

Sp. nov

Valid

Dvořák, Krzemiński & Prokop

Carboniferous (Moscovian)

Mazon Creek fossil beds

 United States
( Illinois)

A blattinopsid.

Clade Paraneoptera

Hemipterans

Auchenorrhyncha

Name Novelty Status Authors Age Type locality Location Notes Images

Archaeomaguviopsis[344]

Gen. et sp. nov

Valid

Fu & Huang

Middle Triassic

Yanchang Formation

 China

A maguviopseid Cicadomorpha.
The type species is A. magicus.

Beloscyta[345]

Gen. et sp. nov

Valid

Shcherbakov

Permian-Triassic boundary

 Mongolia

A scytinopterid Cicadomorph.
The type species is B. edi.

Burbungoides[346]

Gen. et sp. nov

Moulds, Frese & McCurry

Miocene

McGraths Flat Lagerstätte

 Australia

A member of the family Cicadidae. Genus includes new species B. gulgongensis.

Cretodorus multifoveatus[347]

Sp. nov

Liu et al.

Cretaceous

Burmese amber

 Myanmar

A mimarachnid planthopper.

Cycloscytina incompleta[348]

Sp. nov

Valid

Chen et al.

Jurassic

Daohugou Beds

 China

A hylicellid cicadomorph.

Eocicadellium[349]

Gen. et sp. nov

Valid

Fu & Huang

Triassic

Yanchang Formation

 China

A member of the family Archijassidae belonging to the subfamily Karajassinae. The type species is E. grimaldii.

Eumorphoptila[350]

Gen. et sp. nov

Valid

Shcherbakov

Middle Jurassic (Bathonian)

Itat Formation

 Russia
( Krasnoyarsk Krai)

A dysmorphoptilid Prosboloidea Cicadomorph.
The type species is E. pritykinae.

Gedanochila[351]

Gen. et sp. nov

Valid

Brysz, Bourgoin & Szwedo

Eocene (Lutetian)

Baltic amber

Gdańsk Bay area

An Achilidae planthopper.
The type species is G. museisucini.

Ilimpeika[352]

Gen. et sp. nov

Valid

Shcherbakov

Late Permian

Degali Formation

 Russia
( Krasnoyarsk Krai)

A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Ignotalidae. The type species is I. humerosa.

Ingensala[353]

Gen. et sp. nov

Valid

Luo, Song & Szwedo in Luo et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

An inoderbid planthopper.
The type species is I. xiai.

Jaculistilus xixuanae[354]

Sp. nov

In press

Jiang, Chen & Szwedo

Cretaceous

Burmese amber

 Myanmar

A member of the family Mimarachnidae.

Karoontina[355]

Gen. et comb. nov

Valid

Shcherbakov

Late Triassic

 South Africa

A member of the family Palaeontinidae. The type species is "Fletcheriana" magna Riek (1976).

Laiyangella[356]

Gen. et sp. nov

Zhao et al.

Early Cretaceous

Laiyang Formation

 China

A planthopper. Genus includes new species L. zhangi.

Laopsaltria[346]

Gen. et sp. nov

Moulds, Frese & McCurry

Miocene

McGraths Flat Lagerstätte

 Australia

A member of the family Cicadidae. Genus includes new species L. ferruginosa.

Longucercopis[357]

Gen. et sp. nov

Valid

Zhang, Chen & Zhang

Early Jurassic (Pliensbachian-Toarcian)

Sangonghe Formation

 China

A procercopid froghopper.
The type species is L. xinjiangensis.

Maculixius[358]

Gen. et sp. nov

Valid

Bourgoin & Wang in Wang, Liang & Bourgoin

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

An acrotiarine cixiid planthopper.
The type species is M. jiewenae.

Nangamostethos[359]

Gen. et sp. nov

Valid

Xu & Szwedo in Xu et al.

Late Eocene

Niubao Formation

 China

A member of the family Cercopidae belonging to the subfamily Cercopinae and the tribe Cosmoscartini. The type species is N. tibetense.

Papiliontina[355]

Gen. et 3 sp. nov

Valid

Shcherbakov

Triassic

Madygen Formation

 Kyrgyzstan

A member of the family Palaeontinidae. The type species is P. dracomima; genus also includes P. machaon and P. spectans.

Sinogranulus jinsuoguanensis[360]

Sp. nov

Valid

Fu & Huang in Fu, Gao & Huang

Middle Triassic

Yanchang Formation

 China

A granulid scytinopteroid cicadomorph.

Sinoscarterella[35]

Gen. et sp. nov

Valid

Nel, Fu & Huang in Wang et al.

Early Jurassic (Pliensbachian-Toarcian)

Sangonghe Formation

 China

A dysmorphoptilid Prosboloidea cicadomorph.
The type species is S. incompleta.

Stonymetopus[361]

Gen. et sp. nov

Valid

Poinar, Brown & Bourgoin

Cretaceous

Burmese amber

 Myanmar

A fulgoridiid planthopper. The type species is S. megus.

Strivicia[362]

Gen. et sp. nov

Valid

Shcherbakov

Triassic (LadinianCarnian)

Madygen Formation

 Kyrgyzstan

An ipsviciid cicadomorph. The type species is S. davidi.

Tenebricosus[363]

Gen. et sp. nov

In press

He et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Mimarachnidae. The type species is T. coriaceus.

Tennentsia koreana[364]

Sp. nov

Valid

Nel, Nam & Jouault

Late Triassic

Amisan Formation

 South Korea

A dysmorphoptilid cicadomorph.

Tithopsaltria[346]

Gen. et sp. nov

Moulds, Frese & McCurry

Miocene

McGraths Flat Lagerstätte

 Australia

A member of the family Cicadidae. Genus includes new species T. titan.

Unturella[350]

Gen. et sp. nov

Valid

Shcherbakov

Late Permian (Changhsingian), possibly also Early Triassic

Nidym Formation or lower Kochechumo Formation

 Russia
( Krasnoyarsk Krai)

A dysmorphoptilid Prosboloidea cicadomorph.
The type species is U. truncata.

Wangala[365]

Gen. et sp. nov

Chen et al.

Cretaceous

Burmese amber

 Myanmar

A froghopper belonging to the family Sinoalidae. Genus includes new species W. gigantea.

Xiaochibangus[354]

Gen. et comb. nov

In press

Jiang, Chen & Szwedo

Cretaceous

Burmese amber

 Myanmar

A member of the family Mimarachnidae. Genus includes "Dachibangus" formosus Fu et al. (2019) and "Dachibangus" hui Zhang, Yao & Pang in Zhang et al. (2021).

Yobuenahuaboshka[366]

Gen. et sp. nov

Valid

Szwedo et al.

Miocene (Burdigalian)

Dominican amber

 Dominican Republic

A planthopper belonging to the family Nogodinidae.
The type species is Y. rectangialatus.

Zhangpumazus[367]

Gen. et sp. nov

In press

Luo et al.

Miocene

Zhangpu amber

 China

A caliscelid planthopper.
The type species is Z. sheni.

Coleorrhyncha

Name Novelty Status Authors Age Type locality Location Notes Images

Permoridium[368]

Fam. et gen. et sp. nov

Valid

Burckhardt et al.

Early Permian

Saar–Nahe Basin

 Germany

A Coleorrhynchan.
The type genus of the new family Permoridiidae
The type species is P. fresenaci.

Heteroptera

Name Novelty Status Authors Age Type locality Location Notes Images

Acanthamnestus[369]

Gen. et 2 sp. nov

Valid

Du, Yao & Engel in Du et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

An amnestine cydnid bug.
The type species is A. atridorsus and also includes A. ovoideus.

Arcochterus[370]

Gen. et sp. nov

Zhang, Ren & Yao in Zhang et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Ochteridae.
The type species is A. fuscus.

Caulisoculus monlyae[371]

Sp. nov

In press

Zhuo & Chen in Zhuo et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Yuripopovinidae.

Cretocephalus[372]

Gen. et sp. nov

In press

Luo & Xie

Cretaceous

Burmese amber

 Myanmar

A member of the family Aenictopecheidae. The type species is C. stysi.

Laevigemma[369]

Gen. et sp. nov

Valid

Du, Yao & Engel in Du et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

An amnestine cydnid heteropteran.
The type species is L. lisorum.

Leptosaldinea zhengmingi[373]

Sp. nov

In press

Chen & Zhuo in Chen et al.

Cretaceous

Burmese amber

 Myanmar

A possible schizopterid dipsocoromorph heteropteran.

Megaoptocoris[374]

Gen. et 2 sp. nov

In press

Zhou et al.

Cretaceous

Burmese amber

 Myanmar

A yuripopovinid heteropteran.
Genus includes M. punctatus & M. similis.

Pachytylaradus[375]

Gen. et sp. nov

Valid

Heiss

Cretaceous

Burmese amber

 Myanmar

An aradid flat bug.
The type species is P. cretaceous.

Palaeotanyrhina[376]

Fam., gen. et sp. nov

Valid

Poinar, Brown & Kóbor

Cretaceous

Burmese amber

 Myanmar

A reduvioid, cimicomorph heteropteran
The type genus of the new family Palaeotanyrhinidae.
The type species is P. exophthalma.

Parvochterus[370]

Gen. et 2 sp. nov

Zhang, Ren & Yao in Zhang et al.

Cretaceous

Burmese amber

 Myanmar

A velvety shore bug.
The type species is P. reticulatus & also includes P. lanceolarus.

Quinalveus[369]

Gen. et sp. nov

Valid

Du, Yao & Engel in Du et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

An Amnestine burrowing bug.
The type species is Q. hui.

Simplicivenius[377]

Gen. et 2 sp. nov

Zhang, Liu & Yao in Zhang et al.

Cretaceous (Barremian-Aptian)

Yixian Formation

 China

An assassin bug.
The type species is S. tuberculosus& includes S. rectidorsius.

Varicapitatus[378]

Gen. et sp. nov

Valid

Dai, Du & Yao in Dai et al.

Early Cretaceous

Yixian Formation

 China

A member of Pentatomomorpha belonging to the family Pachymeridiidae. The type species is V. sinuolatus.

Heteropteran research
  • Fu et al. (2022) describe adult females of the water boatman Karataviella popovi from the Middle–Late Jurassic Daohugou biota (China) bearing clutches of eggs on their left mesotibia, representing the earliest direct evidence of brood care among insects reported to date.[379]
  • Redescription of Eocenocydnus lisi is published by Lis (2022), who assigns this species to the tribe Cydnini within the family Cydnidae.[380]

Sternorrhyncha

Name Novelty Status Authors Age Type locality Location Notes Images

Burmapsyllidium grimaldii[381]

Sp. nov

Valid

Hakim, Azar & Huang

Cretaceous

Burmese amber

 Myanmar

A member of Protopsyllidioidea belonging to the family Paraprotopsyllidiidae.

Burmodicus monlyae[382]

Sp. nov

In press

Chen & Zhuo in Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A whitefly belonging to the subfamily Aleurodicinae.

Medocellodes[383]

Gen. et sp. nov

Valid

Drohojowska & Szwedo in Drohojowska et al.

Eocene

Baltic amber

Europe (Gdańsk Bay region)

A whitefly. The type species is M. blackmani.

Megalophthallidion[384]

Gen. et sp. nov

Valid

Drohojowska & Szwedo in Drohojowska, Zmarzły & Szwedo

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of Protopsyllidioidea belonging to the family Postopsyllidiidae. The type species is M. burmapateron.

Talbragaropsyllidium[385]

Gen. et sp. nov

In press

Li et al.

Late Jurassic

 Australia

A protopsyllidiid Sternorrhyncha.
The type species is T. averyi.

Triassopsyllidiida[386]

Gen. et sp. nov

Valid

Huang et al.

Middle Triassic (Ladinian)

Yanchang Formation

 China

A possible permopsyllidiid Sternorrhyncha.
The type species is T. pectinata.

Sternorrhyncha research

Psocodea

Name Novelty Status Authors Age Type locality Location Notes Images

Belaphotroctes grimaldii[389]

Sp. nov

Valid

Engel & Wang

Miocene (Langhian)

Zhangpu amber

 China

A member of the family Liposcelididae.

Brachyantennum[390]

Gen. et sp. nov

Valid

Liang & Liu in Zhang, Liang & Liu

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of Trogiomorpha of uncertain affinities. The type species is B. spinosum.

Burmempheria curvatavena[391]

Sp. nov

Valid

Li, Yoshizawa & Yao in Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of Psocodea belonging to the family Empheriidae.

Latempheria[391]

Gen. et sp. nov

Valid

Li, Yoshizawa & Yao in Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of Psocodea belonging to the family Empheriidae. The type species is L. kachinensis.

Libanoglaris hespericus[392]

Sp. nov

Valid

Álvarez-Parra et al.

Early Cretaceous (Albian)

Escucha Formation

 Spain

A member of Psocodea belonging to the family Archaeatropidae.

Longiantennum[393]

Gen. et sp. nov

In press

Liang et al.

Cretaceous

Burmese amber

 Myanmar

A member of Psocodea belonging to the family Archaeatropidae. The type species is L. fashengi.

Parathylacella[394]

Gen. et sp. nov

In press

Álvarez-Parra & Nel

Eocene

Oise amber

 France

A Lepidopsocidae Psocodean. Genus includes new species P. oisensis.

Thysanoptera

Name Novelty Status Authors Age Type locality Location Notes Images

Adstrictubothrips[395]

Gen. et sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A thrips belonging to the family Rohrthripidae. Genus includes new species A. mirapterus.

Alavathrips[396]

Gen. et sp. nov

In press

Peñalver, Nel & Nel

Early Cretaceous (Albian)

Álava amber

 Spain

A thrips belonging to the family Phlaeothripidae or Rohrthripidae.[395] Genus includes new species A. moralesi.

Gemineurothrips[395]

Gen. et 2 sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A thrips belonging to the family Rohrthripidae. Genus includes new species G. microcephalus and G. peculiaris.

Paralleloalathrips[395]

Gen. et sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A thrips belonging to the family Rohrthripidae. Genus includes new species P. bivenatus.

Rohrthrips brachyvenis[395]

Sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A thrips belonging to the family Rohrthripidae.

Rohrthrips multihamuli[395]

Sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A thrips belonging to the family Rohrthripidae.

Rohrthrips pandemicus[395]

Sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A thrips belonging to the family Rohrthripidae.

Rohrthrips rhamphorhynchus[395]

Sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A thrips belonging to the family Rohrthripidae.

Rohrthrips setiger[395]

Sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A thrips belonging to the family Rohrthripidae.

Sesquithrips[395]

Gen. et 2 sp. nov

Valid

Ulitzka

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A thrips belonging to the family Rohrthripidae. Genus includes new species S. markpankowskii and S. rostratus.

Clade Perlidea

Dermapterans

Name Novelty Status Authors Age Type locality Country Notes Images

Stonychopygia laticoncava[397]

Sp. nov

In press

Wang et al.

Cretaceous

Burmese amber

 Myanmar

A pygidicranid earwig.

Tricholabidura[398]

Gen. et sp. nov

In press

Peng & Engel in Peng et al.

Cretaceous

Burmese amber

 Myanmar

A labidurid earwig.
The type species is T. elongata.

Embioptera

Name Novelty Status Authors Age Type locality Location Notes Images

Gnethoda putshkovi[399]

Sp. nov

In press

Anisyutkin & Perkovsky

Cretaceous

Burmese amber

 Myanmar

A member of Embiodea.

Parasorellembia[399]

Gen. et sp. nov

In press

Anisyutkin & Perkovsky

Cretaceous

Burmese amber

 Myanmar

A member of Embiodea. Genus includes new species P. groehni.

Phasmatodea

Name Novelty Status Authors Age Type locality Location Notes Images

Adjacivena grossa[400]

Sp. nov

Yang, Engel, Ren & Gao in Yang et al.

Middle Jurassic

Jiulongshan Formation

 China

A member of the family Susumaniidae belonging to the subfamily Aclistophasmatinae.

Ambrotophasma[400]

Gen. et sp. nov

Yang, Engel, Ren & Gao in Yang et al.

Middle Jurassic

Jiulongshan Formation

 China

A member of the family Pterophasmatidae. The type species is A. vetulum.

Araripephasma[401]

Gen. et comb. nov

Valid

Ghirotto et al.

Early Cretaceous (Aptian)

Crato Formation

 Brazil

A stick insect belonging to the group Euphasmatodea.
The type species is "Eoproscopia" reliquum (2019).

Elasmophasma longitubus[400]

Sp. nov

Yang, Engel, Ren & Gao in Yang et al.

Cretaceous

Burmese amber

 Myanmar

A member of Euphasmatodea belonging to the group Neophasmatodea.

Rhabdophasma[400]

Gen. et sp. nov

Yang, Engel, Ren & Gao in Yang et al.

Cretaceous

Burmese amber

 Myanmar

A member of Euphasmatodea belonging to the group Neophasmatodea. The type species is R. arboreum.

Tanaophasma[400]

Gen. et sp. nov

Yang, Engel, Ren & Gao in Yang et al.

Cretaceous

Burmese amber

 Myanmar

A member of Euphasmatodea belonging to the group Neophasmatodea. The type species is T. applanatum.

Plecopterans

Name Novelty Status Authors Age Type locality Location Notes Images

Amphinemura himberii[402]

Sp. nov

In press

Xu, Nel & Millet

Miocene

Zhangpu amber

 China

A species of Amphinemura.

Baltileuctra dewalti[403]

Sp. nov

Valid

Chen in Chen & Liu

Eocene

Baltic amber

 Lithuania

A member of the family Leuctridae.

Brachyptera dewalti[404]

Sp. nov

In press

Chen

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Brachyptera.

Burmaperla[405]

Gen. et sp. nov

In press

Jouault & Nel in Jouault et al.

Cretaceous

Burmese amber

 Myanmar

A member of the family Perlidae. Genus includes new species B. pouilloni.

Kachinoperla[406]

Fam. et gen. et sp. nov

Valid

Chen

Cretaceous (Albian to Cenomanian)

Burmese amber

 Myanmar

A member of Arctoperlaria belonging to the new family Kachinoperlidae. The type species is K. zwicki.

Largusoperla zhouchangfai[407]

Sp. nov

In press

Chen

Cretaceous

Burmese amber

 Myanmar

A member of the family Perlidae.

Ovaloperla[408]

Gen. et sp. nov

In press

Chen & Xu

Cretaceous

Burmese amber

 Myanmar

A member of the family Petroperlidae. The type species is O. staniczeki.

Perspicuusoperla[409]

Fam. et gen. et sp. nov

Valid

Chen

Cretaceous (Albian-Cenomanian)

Burmese amber

 Myanmar

A member of Arctoperlaria, the type genus of the new family Perspicuusoperlidae. The type species is P. lata.

Plecopteran research

  • A study on the diversity dynamics of plecopterans throughout their evolutionary history, based on data from the fossil record, is published by Jouault et al. (2022)[410]

Clade †Reculida

Name Novelty Status Authors Age Type locality Location Notes Images

Chauliodites tongchuanensis[411]

Sp. nov

Zhang, Jarzembowski & Wang

Middle Triassic (Ladinian)

Tongchuan Formation

 China

A Chaulioditidae Reculida polyneopteran.

Colubrosopterum[412]

Gen. et sp. nov

Valid

Cawood & Nel in Cawood et al.

Permian (Wordian)

Waterford Formation

 South Africa

A member of the family Liomopteridae. The type species is C. karooensis.

Geinitziella[413]

Gen. et sp. nov

Valid

Aristov

Triassic (Ladinian–Carnian)

Madygen Formation

 Kyrgyzstan

A Geinitziidae Reculida Polyneoptera.
Genus includes new species G. rasnitsyni.

Liomopterum connexus[412]

Sp. nov

Valid

Cawood & Nel in Cawood et al.

Permian (Wordian)

Waterford Formation

 South Africa

A member of the family Liomopteridae.

Liomopterum daenerys[412]

Sp. nov

Valid

Cawood & Nel in Cawood et al.

Permian (Wordian)

Waterford Formation

 South Africa

A member of the family Liomopteridae.

Permoshurabia sukhonica[413]

Sp. nov

Valid

Aristov

Late Permian

 Russia

A member of Polyneoptera belonging to the group Reculida and the family Geinitziidae.

Shurabia izyumica[413]

Sp. nov

Valid

Aristov

Late Triassic (Norian)

 Ukraine

A member of Polyneoptera belonging to the group Reculida and the family Geinitziidae.

Shurabia minutissima[413]

Sp. nov

Valid

Aristov

Triassic (Ladinian–Carnian)

Madygen Formation

 Kyrgyzstan

A member of Polyneoptera belonging to the group Reculida and the family Geinitziidae.

Shurabia taewani[414]

Sp. nov

Jouault, Nam & Nel

Late Triassic

Amisan Formation

 South Korea

A member of Polyneoptera belonging to the group Reculida and the family Geinitziidae.

Other insects

Name Novelty Status Authors Age Type locality Location Notes Images

Baharellinus orenburgensis[415]

Sp. nov

Valid

Aristov

Early Triassic

 Russia
( Orenburg Oblast)

A member of Polyneoptera belonging to the group Eoblattida and the family Blattogryllidae.

Lodevoisadia[416]

Gen. et sp. nov

Valid

Nel et al.

Permian (probably Kungurian)

Salagou Formation

 France

A "grylloblattodean" polyneopteran (i.e. a polyneopteran of uncertain affinities, possibly a member of the stem group of Grylloblattida). The type species is L. coheni.

?Partisaniferus edjarzembowskii[417]

Sp. nov

Valid

Haug & Haug

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of Holometabola belonging to the group Neuropteriformia.

Permosialis bayda[418]

Sp. nov

Valid

Aristov & Rasnitsyn

Permian

Uskat Formation

 Russia
( Kemerovo Oblast)

A member of Palaeomanteida (Miomoptera) belonging to the family Permosialidae.

Permosialis belebei[418]

Sp. nov

Valid

Aristov & Rasnitsyn

Permian

Belebei Formation

 Russia
( Kirov Oblast)

A member of Palaeomanteida (Miomoptera) belonging to the family Permosialidae.

General research

  • A study on the diversity dynamics of insects during the Permian and Triassic is published by Jouault et al. (2022), who find a pattern with three extinctions at the Roadian/Wordian, Permian/Triassic and Ladinian/Carnian boundaries, assess the effect of diversity changes between three or four guilds (herbivores, predators, detritivores/fungivores and generalists), and attempt to determine the factors influencing insect diversification dynamics throughout the Permian and Triassic.[419]
  • Conspecific aggregations of insects (dermapterans, neuropterans, orthopterans) and springtails are described from the Cretaceous Burmese amber by Hörnig et al. (2022), who attempt to determine whether these fossils could represent evidence of social behaviour.[420]
  • A flat wasp female preserved grasping and possibly stinging a beetle is described from the Cretaceous Burmese amber by Kiesmüller et al. (2022).[421]

References

  1. ^ a b Zhang, W.; Shih, C.; Engel, M. S.; Ren, D. (2022). "Cretaceous lophocoronids with short proboscis and retractable female genitalia provide the earliest evidence for their feeding and oviposition habits". Cladistics. 38 (6): 684–701. doi:10.1111/cla.12511. PMID 35758057. S2CID 250071929.
  2. ^ Sarto i Monteys, V.; Hausmann, A.; Solórzano-Kraemer, M. M.; Hammel, J. U.; Baixeras, J.; Delclòs, X.; Peñalver, E. (2022). "A new fossil inchworm moth discovered in Miocene Dominican amber (Lepidoptera: Geometridae)". Journal of South American Earth Sciences. 120. 104055. Bibcode:2022JSAES.12004055S. doi:10.1016/j.jsames.2022.104055.
  3. ^ Gauweiler, J.; Haug, C.; Müller, P.; Haug, J. T. (2022). "Lepidopteran caterpillars in the Cretaceous: were they a good food source for early birds?". Palaeodiversity. 15 (1): 45–59. doi:10.18476/pale.v15.a3. S2CID 247268740.
  4. ^ Haug, J. T.; Haug, C.; Wang, Y.; Baranov, V. A. (2022). "The fossil record of lepidopteran caterpillars in Dominican and Mexican amber". Lethaia. 55 (3): 1–14. doi:10.18261/let.55.3.7. S2CID 253416933.
  5. ^ a b Wang, J.; Zhang, W.; Engel, M. S.; Sheng, X.; Shih, C.; Ren, D. (2022). "Early evolution of wing scales prior to the rise of moths and butterflies". Current Biology. 32 (17): 3808–3814.e2. doi:10.1016/j.cub.2022.06.086. PMID 35998638. S2CID 251727701.
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